Basic info Taxonomic history Classification Included Taxa
Morphology Ecology and taphonomy External Literature Search Age range and collections

Perissodactyla (odd-toed ungulate)

Osteichthyes - Perissodactyla

Taxonomy
Perissodactyla was named by Owen (1848) [Owen does not give a rank in his original description of Perissodactyla. Owen does not give a rank in his original description of Perissodactyla. Owen does not give a rank in his original description of Perissodactyla.]. It is extant.

It was reranked as the suborder Perissodactyla by Gadow (1898), Hay (1902), Lambe (1908), Gregory (1910); it was synonymized subjectively with Mesodactyli by Jaekel (1911); it was reranked as the order Perissodactyla by Leidy (1873), Flower (1883), Ferrusquia-Villafranca (1969), McKenna (1975), Thurmond and Jones (1981), Schoch (1984), Kumar and Sahni (1985), Hooker (1986), MacFadden (1986), Carroll (1988), Prothero and Schoch (1989), Nowak (1991), Madsen et al. (2001), Wilson and Reeder (2005), Prothero (2005), Symeonidis et al. (2006), Hulbert and Whitmore (2006), MacFadden (2009), Asher and Helgen (2010), Cooper et al. (2014), Ferrusquía-Villafranca et al. (2014), Bai et al. (2017), Rose et al. (2020).

It was assigned to Diplarthra by Cope (1889), Cope (1891); to Ungulata by Owen (1848), Flower (1883), Ameghino (1894); to Ungulata by Lydekker (1876), Flower and Lydekker (1891), Gadow (1898), Hay (1902), Lambe (1908); to Mesaxonia by Gregory (1910); to Phenacodonta by McKenna (1975); to Eutheria by Hooker (1986); to Altungulata by Prothero and Schoch (1989); to Eutheria by Carroll (1988), Nowak (1991); to Zooamata by Waddell et al. (1999); to Laurasiatheria by Madsen et al. (2001); to Euungulata by Waddell et al. (2001), Asher and Helgen (2010); to Placentalia by Cooper et al. (2014); to Mammalia by Leidy (1873), Ferrusquia-Villafranca (1969), Thurmond and Jones (1981), Schoch (1984), Kumar and Sahni (1985), MacFadden (1986), Wilson and Reeder (2005), Prothero (2005), Symeonidis et al. (2006), Hulbert and Whitmore (2006), MacFadden (2009), Ferrusquía-Villafranca et al. (2014), Bai et al. (2017); and to Perissodactylamorpha by Rose et al. (2020).

Synonyms
Synonymy list
YearName and author
1848Perissodactyla Owen p. 131
1873Perissodactyla Leidy p. 321
1876Perissodactyla Lydekker
1883Perissodactyla Flower p. 185
1889Perissodactyla Cope p. 876
1891Perissodactyla Cope p. 8
1891Perissodactyla Flower and Lydekker p. 90
1894Perissodactyla Ameghino p. 13
1897Hippopotamus iravaticus Noetling
1898Perissodactyla Gadow p. 46
1902Perissodactyla Hay p. 607
1908Perissodactyla Lambe
1910Perissodactyla Gregory p. 466
1969Perissodactyla Ferrusquia-Villafranca
1975Perissodactyla McKenna p. 43
1981Perissodactyla Thurmond and Jones p. 182
1984Perissodactyla Schoch p. 3
1985Perissodactyla Kumar and Sahni p. 153
1986Perissodactyla Hooker p. 342
1986Perissodactyla MacFadden p. 467
1988Perissodactyla Carroll
1989Perissodactyla Prothero and Schoch p. 530
1991Perissodactyla Nowak
1997Selenida McKenna and Bell
1999Perissodactyla Waddell et al. p. 3
2001Perissodactyla Madsen et al. p. 610
2001Perissodactyla Waddell et al. p. 148
2005Perissodactyla Prothero
2005Perissodactyla Wilson and Reeder
2006Perissodactyla Hulbert and Whitmore p. 13
2006Perissodactyla Symeonidis et al.
2009Perissodactyla MacFadden
2010Perissodactyla Asher and Helgen p. 3 figs. Figure 1
2014Perissodactyla Cooper et al. p. 3 fig. 3
2014Perissodactyla Ferrusquía-Villafranca et al. p. 204
2017Perissodactyla Bai et al.
2020Perissodactyla Rose et al. p. 101 fig. 81

Is something missing? Join the Paleobiology Database and enter the data

RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
classOsteichthyes
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
—Tetrapoda
—Reptiliomorpha
—Anthracosauria
subclassAmphibiosauriaKuhn 1967
—Cotylosauria()
—Amniota
subclassSynapsida
—Therapsida()
RankNameAuthor
infraorderCynodontia()
—Mammaliamorpha
—Mammaliaformes
classMammalia
—Cladotheria
—Zatheria
subclassTribosphenida()
subclassTheria
—Eutheria()
—Placentalia
—Boreoeutheria
—Laurasiatheria
—Scrotifera
—Euungulata
—Panperissodactyla
superorderPerissodactylamorpha
orderPerissodactyla()
orderPerissodactyla()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Or. Perissodactyla Owen 1848 [odd-toed ungulate]
show all | hide all
Fm. †Anthracobunidae Wells and Gingerich 1983
+
G. †Anthracobune Pilgrim 1940
+
†Anthracobune aijiensis Kumar 1991
†Anthracobune daviesi Pilgrim 1940
†Anthracobune pinfoldi Pilgrim 1940
+
Invalid names: Pilgrimella pilgrimi Dehm and Oettingen-Spielberg 1958 [synonym]
†Anthracobune wardi Pilgrim 1940
G. †Ishatherium Sahni and Kumar 1980
+
†Ishatherium subathuensis Sahni and Kumar 1980
G. †Jozaria Wells and Gingerich 1983
+
†Jozaria palustris Wells and Gingerich 1983
G. †Lammidhania Gingerich 1977
G. †Obergfellia Cooper et al. 2014
+
†Obergfellia occidentalis Cooper et al. 2014
G. †Pilgrimella Dehm and Oettingen-Spielberg 1958
Fm. †Brontotheriidae Marsh 1873 [brontothere]
+
G. †Balochititanops Missiaen et al. 2011
+
†Balochititanops haqi Missiaen et al. 2011
Subfm. †Brontotheriinae Marsh 1873
+
G. †Acrotitan Ye 1983
+
Tr. †Brontotheriini Marsh 1873
+
Subtr. †Brontotheriina Marsh 1873
+
Unr. †Brontotheriita Marsh 1873
G. †Diplacodon Marsh 1875
+
Invalid names: Eotitanotherium Peterson 1914 [synonym], Pseudodiplacodon Mader 2000 [synonym]
G. †Epimanteoceras Granger and Gregory 1943
G. †Pachytitan Granger and Gregory 1943
G. †Protitan Granger and Gregory 1943
G. †Rhinotitan Granger and Gregory 1943
G. †Microtitan Granger and Gregory 1943
+
†Microtitan mongoliensis Osborn 1925
Invalid names: Microtitan elongatus Qi 1987 [nomen dubium]
G. †Nanotitanops Qi and Beard 1998
+
†Nanotitanops shanghuangensis Qi and Beard 1998
+
Invalid names: Nanotitan shanghuangensis Qi and Beard 1996 [replaced]
Invalid names: Nanotitan Qi and Beard 1996 [synonym]
Subtr. †Telmatheriina Osborn 1914
+
G. †Qufutitan Wang and Wang 1997
G. †Wickia Mihlbachler 2008
Invalid names: Metatelmatheriinae Granger and Gregory 1943 [synonym]
G. †Bunobrontops Holroyd and Ciochon 2000
+
†Bunobrontops savagei Holroyd and Ciochon 2000
G. †Desmatotitan Granger and Gregory 1943
+
†Desmatotitan tukhumensis Granger and Gregory 1943
Tr. †Dolichorhinini Riggs 1912
+
G. †Mesatirhinus Osborn 1908
+
+
Invalid names: Mesatirhinus petersoni Osborn 1908 [synonym]
Invalid names: Mesatirhinus junius Leidy 1872 [nomen dubium]
G. †Sphenocoelus Osborn 1895
+
†Sphenocoelus hyognathus Osborn 1889
+
Invalid names: Dolichorhinus longiceps Douglass 1909 [synonym], Sphenocoelus cornutum Osborn 1895 [synonym]
†Sphenocoelus intermedius Osborn 1908
+
Invalid names: Dolichorhinus fluminalis Riggs 1912 [synonym], Dolichorhinus heterodon Douglass 1909 [synonym], Mesatirhinus superior Riggs 1912 [synonym]
†Sphenocoelus uintensis Osborn 1895
+
Invalid names: Sphenocoelus blairi Cook 1926 [synonym], Sphenocoelus bridgeri Cook 1926 [synonym], Sphenocoelus harundivoras Cook 1926 [synonym]
Invalid names: Dolichorhinoides Granger and Gregory 1943 [synonym], Dolichorhinus Hatcher 1895 [synonym], Tanyorhinus Cook 1926 [synonym]
Invalid names: Rhadinorhininae Osborn 1929 [synonym]
Unr. †Embolotheriita Osborn 1929
+
G. †Aktautitan Mihlbachler et al. 2004
+
†Aktautitan hippopotamopus Mihlbachler et al. 2004
G. †Brachydiastematherium Bockh and Maty 1876
+
†Brachydiastematherium transylvanicum Bockh and Maty 1876
G. †Embolotherium Osborn 1929
+
†Embolotherium andrewsi Osborn 1929
†Embolotherium grangeri Osborn 1929
+
Invalid names: Embolotherium insigne Yanovskaya 1980 [synonym], Embolotherium loucksii Osborn 1929 [synonym], Titanodectes ingens Granger and Gregory 1943 [synonym], Titanodectes minor Granger and Gregory 1943 [synonym]
†Embolotherium ultimum Granger and Gregory 1943
G. †Gnathotitan Granger and Gregory 1943
+
†Gnathotitan berkeyi Osborn 1925
G. †Maobrontops Averianov et al. 2018
+
†Maobrontops paganus Averianov et al. 2018
G. †Metatitan Granger and Gregory 1943
+
†Metatitan khaitshinus Yanovskaya 1954
+
Invalid names: Metatitan reshetovi Yanovskaya 1980 [synonym]
†Metatitan primus Granger and Gregory 1943
†Metatitan relictus Granger and Gregory 1943
+
Invalid names: Protitan khaitshinus Yanovskaya 1980 [synonym]
G. †Nasamplus Mihlbachler 2008
+
†Nasamplus progressus Granger and Gregory 1943
G. †Pollyosbornia Mihlbachler 2008
+
†Pollyosbornia altidens Osborn 1908
G. †Protembolotherium Yanovskaya 1954
+
†Protembolotherium efremovi Yanovskaya 1954
G. †Pygmaetitan Miao 1982
+
G. †Metarhinus Osborn 1908
+
†Metarhinus abbotti Riggs 1912
†Metarhinus diploconus Osborn 1895
+
Invalid names: Rhadinorhinus abbotti Riggs 1912 [synonym]
†Metarhinus fluviatilis Osborn 1908
+
Invalid names: Metarhinus riparius Riggs 1912 [synonym]
†Metarhinus pater Stock 1937
Invalid names: Heterotitanops Peterson 1914 [synonym], Metarhinus cristatus Riggs 1912 [nomen dubium], Metarhinus earlei Osborn 1908 [nomen dubium], Rhadinorhinus Riggs 1912 [synonym], Telmatherium accola Cook 1926 [nomen dubium], Telmatherium advocata Cook 1926 [nomen dubium]
G. †Protitanotherium Osborn 1908
+
+
Invalid names: Protitanotherium superbum Osborn 1908 [synonym], Sthenodectes australis Wilson 1977 [synonym]
Invalid names: Protitanotherium koreanicum Takai 1939 [nomen dubium]
G. †Telmatherium Marsh 1872
+
†Telmatherium validum Marsh 1872
†Telmatherium validus Marsh 1872
+
Invalid names: Telmatherium cultridens Osborn et al. 1878 [synonym], Telmatherium manteoceras Osborn in Matthew 1899 [synonym], Telmatherium washakiensis Osborn 1908 [synonym]
Invalid names: Leurocephalus Osborn et al. 1878 [synonym], Manteoceras Hatcher 1895 [synonym], Telmatotherium Marsh 1872 [synonym]
G. †Danjiangia Wang 1995
+
†Danjiangia lambdodon Bai et al. 2018
†Danjiangia pingi Wang 1995
Subfm. †Embolotheriinae Osborn 1929
+
G. †Titanodectes Granger and Gregory 1943
Subfm. †Eotitanopinae Osborn 1913
+
G. †Eotitanops Osborn 1907
+
†Eotitanops borealis Cope 1880
+
Invalid names: Eotitanops brownianus Cope 1881 [synonym], Eotitanops major Osborn 1913 [synonym], Eotitanops princeps Osborn 1913 [synonym]
†Eotitanops dayi Dehm and Oettingen-Spielberg 1958
†Eotitanops gregoryi Osborn 1913
+
Invalid names: Eotitanops minimus Osborn 1919 [synonym]
†Eotitanops pakistanensis Missiaen et al. 2011
G. †Lambdotherium Cope 1880
+
+
Invalid names: Lambdotherium magnum Osborn 1919 [synonym], Lambdotherium primaevum Loomis 1907 [synonym], Lambdotherium priscum Osborn 1913 [synonym], Lambdotherium progressum Osborn 1913 [synonym]
G. †Metatelmatherium Granger and Gregory 1938
+
†Metatelmatherium cristatum Granger and Gregory 1938
†Metatelmatherium ultimum Osborn 1908
+
Invalid names: Manteoceras uintensis Douglass 1909 [synonym]
Invalid names: Metatelmatherium lahirii Pilgrim 1925 [nomen dubium], Metatelmatherium parvum Granger and Gregory 1943 [nomen dubium]
G. †Mulkrajanops Kumar and Sahni 1985
+
†Mulkrajanops moghliensis Kumar and Sahni 1985
†Mulkrajanops mogliensis Kumar and Sahni 1985
G. †Pakotitanops West 1980
+
Subfm. †Palaeosyopinae Steinmann and Döderlein 1890
+
G. †Palaeosyops Leidy 1870
+
+
Invalid names: Eometarhinus huerfanensis Osborn 1919 [synonym]
†Palaeosyops junius Leidy 1873
†Palaeosyops manteoceras Osborn in Matthew 1899
†Palaeosyops paludosus Leidy 1870
+
Invalid names: Canis marshii Hay 1899 [synonym], Limnohyops matthewi Osborn 1908 [synonym], Palaeosyops humilis Leidy 1872 [synonym], Palaeosyops laevidens Cope 1873 [synonym], Palaeosyops longirostris Earle 1892 [synonym], Palaeosyops major Leidy 1871 [synonym], Palaeosyops minor Marsh 1871 [synonym], Palaeosyops minor Earle 1891 [synonym], Palaeosyops priscus Osborn 1908 [synonym]
†Palaeosyops robustus Marsh 1872
+
Invalid names: Palaeosyops copei Osborn 1908 [synonym], Palaeosyops diaconus Cope 1873 [synonym], Palaeosyops grangeri Osborn 1908 [synonym], Palaeosyops laticeps Leidy 1872 [synonym], Palaeosyops leidyi Osborn 1908 [synonym]
Invalid names: Eometarhinus Osborn 1919 [synonym], Limnohyops Marsh 1890 [synonym], Limnohyus Marsh 1872 [synonym], Palaeosyops ultimus Matthew 1897 [nomen nudum]
G. †Sivatitanops Pilgrim 1925
+
†Sivatitanops cotteri Pilgrim 1925
Invalid names: Sivatitanops birmanicum Pilgrim and Cotter 1916 [nomen dubium]
G. †Sthenodectes Gregory 1912
+
†Sthenodectes incisivum Douglass 1909
+
Invalid names: Sthenodectes priscus Peterson 1934 [synonym]
G. †Xylotitan Mihlbachler and Samuels 2016
+
†Xylotitan cenosus Mihlbachler and Samuels 2016
Invalid names: Brontopinae Osborn 1914 [invalid subgroup], Epimanteoceratinae Granger and Gregory 1943 [empty], Menodontidae Cope 1881 [invalid subgroup], Titanotheriidae [synonym]
Fm. †Cambaytheriidae Bajpai et al. 2005
+
G. †Kalitherium Bajpai et al. 2006
G. †Cambaytherium Bajpai et al. 2005
+
†Cambaytherium gracilis Smith et al. 2016
†Cambaytherium marinus Bajpai et al. 2006
†Cambaytherium thewissi Bajpai et al. 2005
+
Invalid names: Cambaytherium bidens Bajpai et al. 2005 [synonym], Cambaytherium minor Bajpai et al. 2005 [synonym], Indobune vastanensis Rose et al. 2006 [synonym]
Invalid names: Indobune Rose et al. 2006 [synonym]
Or. †Desmostylia Reinhart 1953 [desmostylian]
+
G. †Behemotops Domning et al. 1986
+
†Behemotops katsuiei Inuzuka 2000
†Behemotops proteus Domning et al. 1986
Or. †Desmostyloidea Abel 1933
+
Fm. †Desmostylidae Osborn 1905
+
G. †Ashoroa Inuzuka 2000
+
†Ashoroa laticosta Inuzuka 2000
G. †Cornwallius Hay 1923
+
†Cornwallius sookensis Cornwall 1922
G. †Desmostylus Marsh 1888
+
†Desmostylus hesperus Marsh 1888
+
Invalid names: Desmostylella typica Nagao 1937 [synonym], Desmostylus californicus Hay 1923 [synonym], Desmostylus cymatias Hannibal 1922 [synonym], Desmostylus minor Nagao 1937 [synonym], Desmostylus mirabilis Nagao 1935 [synonym], Desmostylus watasei Hay 1915 [synonym]
†Desmostylus japonicus Tokunaga and Iwasaki 1914
Invalid names: Desmostylella Nagao 1937 [synonym]
G. †Ounalashkastylus Chiba et al. 2016
+
†Ounalashkastylus tomidai Chiba et al. 2016
G. †Vanderhoofius Reinhart 1959
+
†Vanderhoofius coalingensis Reinhart 1959
Fm. †Paleoparadoxiidae Reinhart 1959
+
G. †Archaeoparadoxia Barnes 2013
+
G. †Neoparadoxia Barnes 2013
+
†Neoparadoxia cecilialina Barnes 2013
†Neoparadoxia repenningi Domning and Barnes 2007
G. †Paleoparadoxia Reinhart 1959
+
†Paleoparadoxia media Inuzuka 2005
†Paleoparadoxia tabatai Tokunaga 1939
Invalid names: Behemotopsinae Inuzuka 1987 [empty], Paleoparadoxiinae Reinhart 1959 [empty]
G. †Jamilcotatus Aranda-Manteca and Barnes 1998
+
†Jamilcotatus boreios Aranda-Manteca and Barnes 1998
†Jamilcotatus kumiaica Aranda-Manteca and Barnes 1998
G. †Kronokotherium Pronina 1957
+
G. †Seuku Beatty and Cockburn 2015
+
†Seuku emlongi Domning et al. 1986
Invalid names: Cornwalliidae Shikama 1966 [empty], Desmostyliformes Hay 1923 [synonym]
Superfm. Equoidea Hay 1902
+
G. †Cymbalophus Hooker 1980
+
†Cymbalophus cuniculus Owen 1842
†Cymbalophus hookeri Godinot et al. 1987
Fm. Equidae Gray 1821 [horse]
+
Subfm. †Anchitheriinae Leidy 1869 [anchitheriine horse]
+
G. †Anchitherium von Meyer 1844
+
Subg. †Anchitherium (Hypohippus) Leidy 1858
Subg. †Anchitherium (Parahippus) Leidy 1858
†Anchitherium clarencei Simpson 1932
†Anchitherium hypohippoides Matsumoto 1924
†Anchitherium zitteli Schlosser 1903
G. †Archaeohippus Gidley 1906
+
+
Invalid names: Archaeohippus nanus Simpson 1932 [synonym], Parahippus minutalis Hay 1924 [synonym]
†Archaeohippus mannulus O'Sullivan 2003
†Archaeohippus minimus Douglass 1899
†Archaeohippus mourningi Merriam 1913
†Archaeohippus penultimus Matthew 1924
†Archaeohippus ultimus Cope 1886
G. †Desmatippus Scott 1893
+
†Desmatippus avus Marsh 1874
+
Invalid names: Anchippus brevidens Marsh 1874 [synonym]
†Desmatippus crenidens Scott 1893
†Desmatippus integer Matthew 1924
†Desmatippus nebrascensis Peterson 1907
†Desmatippus texanus Leidy 1868
†Desmatippus tyleri Loomis 1908
Invalid names: Anchippus Leidy 1868 [synonym]
G. †Hypohippus Leidy 1858
+
†Hypohippus affinis Leidy 1858
†Hypohippus chico MacFadden 1982
†Hypohippus equinus Scott 1893
†Hypohippus nevadensis Merriam 1913
†Hypohippus osborni Gidley 1907
†Hypohippus pertinax Matthew 1918
Invalid names: Hypohippus (Drymohippus) Merriam 1913 [nomen dubium]
G. †Kalobatippus Osborn 1915
+
†Kalobatippus agatensis Osborn 1918
†Kalobatippus australis Leidy 1873
†Kalobatippus avus Schlaikjer 1935
+
Invalid names: Kalobatippus gracilis Marsh 1892 [synonym]
G. †Megahippus McGrew 1938
+
†Megahippus matthewi Barbour 1914
†Megahippus mckennai Tedford and Alf 1962
G. †Mesohippus Marsh 1875
+
†Mesohippus bairdi Leidy 1850
+
Invalid names: Mesohippus hypostylus Osborn 1904 [synonym], Mesohippus proteulophus Osborn 1904 [synonym], Mesohippus viejensis Clark and Beerbower 1967 [synonym]
†Mesohippus barbouri Schlaikjer 1931
†Mesohippus celer Marsh 1874
†Mesohippus exoletus Cope 1874
+
Invalid names: Mesohippus trigonostylus Osborn 1918 [synonym]
†Mesohippus latidens Douglass 1903
†Mesohippus propinquus Lambe 1905
†Mesohippus texanus McGrew 1971
†Mesohippus westoni Cope 1889
+
Invalid names: Mesohippus montanensis Osborn 1904 [synonym], Mesohippus praecocidens Lambe 1905 [synonym]
Invalid names: Mesohippus cuneatus Cope 1873 [nomen dubium]
G. †Miohippus Marsh 1874
+
†Miohippus anceps Marsh 1874
†Miohippus annectens Marsh 1874
+
Invalid names: Miohippus crassicuspis Osborn 1904 [synonym]
+
Invalid names: Mesohippus planidens Lambe 1905 [synonym]
†Miohippus condoni Leidy 1870
†Miohippus equiceps Cope 1878
+
Invalid names: Anchitherium brachylophum Cope 1878 [synonym], Mesohippus acutidens Sinclair 1905 [synonym]
†Miohippus equinanus Osborn 1918
†Miohippus gemmarosae Osborn 1918
†Miohippus gidleyi Osborn 1904
+
Invalid names: Mesohippus grallipes Sinclair 1925 [synonym], Miohippus validus Osborn 1904 [synonym]
†Miohippus grandis Clark and Beerbower 1967
†Miohippus intermedius Osborn and Wortman 1895
†Miohippus obliquidens Osborn 1904
+
Invalid names: Mesohippus brachystylus Osborn 1904 [synonym], Mesohippus eulophus Osborn 1904 [synonym], Mesohippus meteulophus Osborn 1904 [synonym], Pediohippus antiquus Schlaikjer 1935 [synonym]
†Miohippus primus Osborn 1918
†Miohippus quartus Osborn 1918
†Miohippus taxus Douglass 1908
Invalid names: Altippus Douglass 1908 [synonym], Anchitherium longicristis Cope 1878 [nomen dubium], Pediohippus Schlaikjer 1935 [synonym]
G. †Parahippus Leidy 1858
+
†Parahippus agrestis Leidy 1873
†Parahippus atavus Osborn 1918
†Parahippus cognatus Leidy 1858
†Parahippus coloradensis Gidley 1907
†Parahippus leonensis Sellards 1916
+
Invalid names: Merychippus socius Hay 1924 [synonym], Parahippus barbouri White 1942 [synonym], Parahippus vellicans Hay 1924 [synonym]
†Parahippus maxsoni Jahns 1940
†Parahippus pawniensis Gidley 1907
†Parahippus pristinus Osborn 1918
†Parahippus wyomingensis Schlaikjer 1935
G. †Sinohippus Zhai 1962
+
†Sinohippus sampelayoi Villalta and Crusafont 1945
Invalid names: Anchitherium australe Leidy 1873 [nomen dubium]
G. †Cremohipparion Zhanxiang et al. 1987
G. †Duchesnehippus Peterson 1932
+
†Duchesnehippus intermedius Peterson 1932
G. †Eohippus Marsh 1876
+
†Eohippus angustidens Cope 1875
+
Invalid names: Eohippus validus Marsh 1876 [synonym], Hyracotherium cuspidatum Cope 1875 [synonym], Hyracotherium seekinsi Morris 1968 [synonym]
†Eohippus borealis Granger 1908
†Eohippus cristatus Wortman 1896
†Eohippus cristonensis Cope 1877
†Eohippus resartus Granger 1908
G. †Epihippus Marsh 1878
+
Subg. †Epihippus (Duchesnehippus) Peterson 1932
Subg. †Epihippus (Epihippus) Stirton 1940
†Epihippus gracilis Marsh 1871
+
Invalid names: Epihippus parvus Granger 1908 [synonym]
†Epihippus uintensis Marsh 1875
Subfm. Equinae Steinmann and Döderlein 1890
+
G. †Acritohippus Kelly 1995
+
†Acritohippus isonesus Cope 1889
+
Invalid names: Merychippus praecocidens Russell 1933 [synonym]
†Acritohippus quinni Kelly 1995
†Acritohippus stylodontus Merriam 1919
†Acritohippus tertius Osborn 1918
Tr. Equini Quinn 1955 [horse]
+
G. †Allohippus Kretzoi 1938
G. †Boreohippidion Dos Santos Avilla et al. 2015
G. †Calippus Matthew and Stirton 1930
G. †Dinohippus Quinn 1955
G. Equus Linnaeus 1758 [horse]
+
Invalid names: Allozebra Trumler 1961 [synonym], Asinus Gray 1822 [synonym], Dolichohippus Heller 1912 [synonym], Haringtonhippus Heintzman et al. 2017 [synonym], Hippotigris Hamilton-Smith 1841 [synonym], Kolpohippus van Hoepen 1930 [synonym], Kraterohippus van Hoepen 1930 [synonym], Neohippus Abel 1913 [synonym], Onager Brisson 1762 [synonym], Sterrohippus van Hopen 1930 [synonym], Tomolabis Cope 1892 [synonym]
G. †Heteropliohippus Kelly 1995
G. †Hippidion Owen 1869
+
Invalid names: Onohippidium Moreno 1891 [synonym], Parahipparion Ameghino 1904 [synonym]
G. †Parapliohippus Kelly 1995
G. †Plesippus Matthew 1924
Subtr. Pliohippina Prado and Alberdi 1996
G. †Pliohippus Marsh 1874
G. †Protohippus Leidy 1858
+
Invalid names: Eoequus Quinn 1955 [synonym]
Invalid names: Equus proversus Merriam 1916 [nomen dubium], Protohippina Quinn 1955 [empty]
Tr. †Hipparionini Quinn 1955 [hipparionine horse]
+
G. †Cormohipparion Skinner and MacFadden 1977
G. †Eurygnathohippus Van Hoepen 1930
+
Invalid names: Notohipparion Haughton 1932 [synonym]
G. †Hipparion de Christol 1832
+
Invalid names: Hemihipparion Wehrli 1941 [synonym], Hipparitherium Christol 1847 [synonym], Stylohipparion van Hoepen 1932 [synonym]
G. †Hippotherium Kaup 1832
G. †Nannippus Matthew 1926
G. †Neohipparion Gidley 1903
+
Invalid names: Hesperohipparion Dalquest 1981 [synonym]
G. †Pseudhipparion Ameghino 1904
+
Invalid names: Griphippus Quinn 1955 [synonym]
Invalid names: Hipparion anthonyi Merriam 1916 [nomen dubium], Hippotherium sinclairi Wortman 1882 [nomen dubium]
G. †Merychippus Leidy 1856
+
Subg. †Merychippus (Merychippus) Stirton 1940
Subg. †Merychippus (Protohippus) Leidy 1858
†Merychippus californicus Merriam 1915
†Merychippus coloradense Osborn 1918
+
Invalid names: Hipparion sanfondensis Frick 1933 [synonym]
†Merychippus eohipparion Osborn 1918
†Merychippus eoplacidus Osborn 1918
†Merychippus gunteri Simpson 1930
†Merychippus insignis Leidy 1856
+
Invalid names: Merychippus paniensis Cope 1874 [synonym]
†Merychippus labrosus Cope 1874
†Merychippus missouriensis Douglass 1908
†Merychippus patruus Osborn 1918
†Merychippus primus Osborn 1918
†Merychippus proparvulus Osborn 1918
†Merychippus quintus Osborn 1918
†Merychippus relictus Cope 1889
†Merychippus republicanus Osborn 1918
†Merychippus sejunctus Cope 1874
†Merychippus seversus Cope 1878
†Merychippus stevensi Dougherty 1940
Invalid names: Merychippus isonesus secundus Osborn 1918 [nomen vanum], Merychippus quartus Osborn 1918 [nomen vanum]
G. †Scaphohippus Pagnac 2006
+
†Scaphohippus sumani Merriam 1915
+
Invalid names: Merychippus stylodus Merriam 1915 [synonym]
Invalid names: Calippini Quinn 1955 [empty], Hippodon Leidy 1854 [invalid subgroup], Hippotheriini Bonaparte 1850 [empty], Protohippini Quinn 1955 [empty], Stylonus Cope 1878 [nomen dubium]
G. †Erihippus Bai et al. 2018
+
†Erihippus tingae Bai et al. 2018
G. †Gobihippus Dashzeveg 1979
G. †Hipphaplus Ameghino 1882
+
†Hipphaplous entrerianus Ameghino 1885
G. †Hyperhippidium Sefve 1910
G. †Hypohippidium Fernandez de Alvarez 1957
+
†Hypohippidium humahuaquense Fernandez de Alvarez 1957
Subfm. †Hyracotheriinae Cope 1881
+
G. †Haplohippus McGrew 1953
+
†Haplohippus texanus McGrew 1953
G. †Lophiopus Ellenberger 1980
G. †Minippus Froehlich 2002
+
†Minippus index Cope 1873
+
Invalid names: Hyracotherium sulcatus Gingerich 1989 [synonym], Orotherium loevii Cope 1877 [synonym]
†Minippus jicarillai Froehlich 2002
G. †Orohippus Marsh 1872
+
†Orohippus agilis Marsh 1873
†Orohippus major Marsh 1874
†Orohippus progressus Granger 1908
†Orohippus proteros Korth and Evander 1982
†Orohippus pumilus Marsh 1871
+
Invalid names: Helotherium procyoninum Cope 1871 [synonym], Hyracotherium osbornianum Cope 1883 [synonym], Lophiotherium ballardi Marsh 1871 [synonym], Oligotomus cinctus Cope 1873 [synonym], Orohippus atavus Granger 1908 [synonym], Orohippus typicus Granger 1908 [synonym]
†Orohippus sylvaticus Leidy 1870
+
Invalid names: Orotherium uintanum Marsh 1872 [synonym]
Invalid names: Helohippus Marsh 1892 [synonym], Helotherium Cope 1872 [synonym], Oligotomus Cope 1873 [synonym], Orohippus (Aminippus) Granger 1908 [invalid subgroup]
G. †Plesiohipparion Qiu et al. 1987
G. †Pliolophus Owen 1858
+
†Pliolophus barnesi Hooker 2010
†Pliolophus quesnoyensis Bronnert et al. 2017
†Pliolophus vulpiceps Owen 1858
G. †Protorohippus Wortman 1896
+
†Protorohippus montanum Wortman 1896
G. †Sifrhippus Froehlich 2002
+
†Sifrhippus grangeri Gingerich 1989
†Sifrhippus sandrae Gingerich 1989
Invalid names: Arenahippus Froehlich 2002 [synonym]
G. †Sivalhippus Lydekker 1877
+
†Sivalhippus theobaldi Lydekker 1877
G. †Xenicohippus Bown and Kihm 1981
+
†Xenicohippus grangeri Bown and Kihm 1981
†Xenicohippus osborni Bown and Kihm 1981
Invalid names: Protohippinae Gidley 1907 [empty]
Fm. †Pachynolophidae Pavlow 1888
+
Subfm. †Plagiolophinae Remy 1976
+
G. †Metaplagiolophus Checa Soler 1993
+
†Metaplagiolophus atoae Checa Soler 1993
Fm. †Palaeotheriidae Bonaparte 1850
+
G. †Cantabrotherium Casanova-Cladellas and Santafe-Llopis 1987
+
†Cantabrotherium truyolsi Casanovas and Santafe 1987
G. †Eurohippus Franzen 2006
+
†Eurohippus messlense Haupt 1925
†Eurohippus parvulus Laurillard 1849
G. †Franzenium Casanovas and Santafe 1989
+
†Franzenium durense Cuesta 1993
†Franzenium tetradactylum Casanovas and Santafe 1989
G. †Hyracotherium Owen 1841
+
†Hyracotherium levei Hooker 1994
†Hyracotherium remyi Checa-Soler 1997
+
Invalid names: Hyrachyus singularis Cope 1875 [synonym]
G. †Iberolophus Badiola and Cuesta 2008
+
†Iberolophus arabensis Badiola and Cuesta 2008
†Iberolophus jimenezi Badiola and Cuesta 2008
G. †Leptolophus Remy 1965
+
†Leptolophus cuestai Perales-Gogenola et al. 2021
†Leptolophus franzeni Perales-Gogenola et al. 2021
†Leptolophus magnus Remy 1998
†Leptolophus nouleti Remy 1998
†Leptolophus stehlini Remy 1965
G. †Orolophus Remy 2017
+
†Orolophus maldani Remy 2017
Invalid names: Orotherium Marsh 1872 [replaced]
Subfm. †Pachynolophinae Paulow 1877
+
Tr. †Anchilophini Remy 2012
+
G. †Anchilophus Gervais 1852
G. †Lophiohippus Bai 2017
G. †Bepitherium Checa and Colombo 2004
+
†Bepitherium jordifusalbae Checa and Colombo 2004
G. †Lophiotherium Marsh 1871
+
†Lophiotherium cervulum Gervais 1849
†Lophiotherium robiacense Deperet 1917
G. †Metanchilophus Remy 2012
+
†Metanchilophus depereti Stehlin 1905
†Metanchilophus dumasi Gervais 1849
†Metanchilophus gaudini Pictet and Humbert 1869
G. †Pachynolophus Pomel 1847
+
†Pachynolophus boixedatensis Crusafont Pairo and Remy 1970
†Pachynolophus cayluxi Filhol 1888
†Pachynolophus duvali Pomel 1847
†Pachynolophus eulaliensis Danilo et al. 2013
†Pachynolophus gaytei Remy 2015
†Pachynolophus hookeri Godinot et al. 1987
†Pachynolophus livinierensis Savage et al. 1965
†Pachynolophus molipontiensis Checa-Soler 1997
†Pachynolophus ruscassierensis Remy et al. 2019
†Pachynolophus zambranensis Badiola et al. 2005
G. †Propalaeotherium Gervais
+
†Propalaeotherium gaudryi Lemoine 1878
+
Invalid names: Lophiotherium magnum Matthes 1977 [synonym]
†Propalaeotherium helveticum Savage et al. 1965
†Propalaeotherium sinense Zdansky 1930
†Propalaeotherium sudrei Remy et al. 2016
†Propalaeotherium voigti Matthes 1977
G. †Qianohippus Miao 1982
+
†Qianohippus magicus Miao 1982
Subfm. †Palaeotheriinae Bonaparte 1850
+
G. †Palaeotherium Cuvier 1804
+
†Palaeotherium cartieri Stehlin 1904
†Palaeotherium castrense Noulet 1863
†Palaeotherium crassum Cuvier 1805
†Palaeotherium crusafonti Cassanovas-Cladellas 1975
†Palaeotherium curtum Cuvier 1812
†Palaeotherium duvali Pomel 1853
†Palaeotherium eocaenum Gervais 1875
†Palaeotherium giganteum Cuesta 1993
†Palaeotherium isselanum Cuvier 1824
†Palaeotherium lautricense Stehlin 1904
†Palaeotherium magnum Cuvier 1804
†Palaeotherium maximum Leidy 1852
†Palaeotherium medium Cuvier 1804
†Palaeotherium minimum Cuvier 1836
†Palaeotherium muehlbergi Stehlin 1904
†Palaeotherium pomeli Franzen 1968
†Palaeotherium renevieri Stehlin 1904
†Palaeotherium robustum Franzen 1968
†Palaeotherium ruetimeyeri Franzen 1968
†Palaeotherium siderolithicum Pictet and Humbert 1869
†Palaeotherium tapiroides Cuvier 1812
G. †Plagiolophus Pomel 1847
+
Subg. †Plagiolophus (Plagiolophus) Pomel 1847
†Plagiolophus cartailhaci Stehlin 1904
†Plagiolophus codiciensis Gaudry 1865
†Plagiolophus curtisi Hooker 1986
†Plagiolophus markgrafi Lőrenthey 1907
†Plagiolophus ministri Brunet and Jehenne 1989
†Plagiolophus minor Cuvier 1804
†Plagiolophus minus Cuvier 1804
G. †Paraplagiolophus Depéret 1917
G. †Propachynolophus Lemoine 1891
+
G. †Pseudopalaeotherium Franzen 1972
G. †Ghazijhippus Missiaen and Gingerich 2014
+
†Ghazijhippus talibhasani Missiaen and Gingerich 2014
G. †Hallensia Franzen and Haubold 1986
+
†Hallensia louisi Hooker 1994
†Hallensia matthesi Franzen and Haubold 1986
Subor. †Lophodontomorpha Hooker and Dashzeveg 2004
+
Infraor. †Euperissodactyla Hooker and Dashzeveg 2004
+
Unr. †Hippomorpha
+
Fm. †Indolophidae Schoch 1984
+
G. †Indolophus Pilgrim 1925
Invalid names: Pachynolophoidea Pavlow 1888 [empty]
G. †Orientolophus Ting 1993
+
G. †Mesolambdolophus Holbrook and Lapergola 2011
+
†Mesolambdolophus setoni Holbrook and Lapergola 2011
G. †Nakusia Ginsburg et al. 1999
+
†Nakusia shahrigensis Ginsburg et al. 1999
G. †Paschatherium Russell 1963
+
†Paschatherium dolloi Teilhard de Chardin 1927
†Paschatherium levei Hooker and Russell 2012
†Paschatherium marianae Estravís and Russell 1992
†Paschatherium plaziati Marandat 1989
†Paschatherium russelli Godinot 1978
†Paschatherium yvettae Gheerbrant et al. 1997
G. †Perissobune Missiaen and Gingerich 2014
+
†Perissobune intizarkhani Missiaen and Gingerich 2014
†Perissobune munirulhaqi Missiaen and Gingerich 2014
G. †Skopaiolophus Métais et al. 2006
+
†Skopaiolophus burmese Métais et al. 2006
Subor. †Solidungula Cuvier
+
Fm. †Anchitheridae Leidy 1869 [anchitheriine horse]
+
G. †Limnohyus Leidy 1872
Unr. Tapiromorpha Haeckel 1873
+
Unr. †Ancylopoda Cope 1889
+
Superfm. †Chalicotherioidea Gill 1872
+
Fm. †Chalicotheriidae Gill 1872 [chalicothere]
+
G. †Eomoropus Osborn 1913
G. †Limognitherium Filhol 1880
Subfm. †Macrotheriinae Holland and Peterson 1913
G. †Moropus Marsh 1877
G. †Pernatherium Gervais 1876
Subfm. †Schizotheriinae Holland and Peterson 1913
G. †Schizotherium Gervais 1876
Invalid names: Moropodinae Marsh 1877 [empty]
Fm. †Eomoropidae Matthew 1929
+
G. †Litolophus Radinsky 1964
G. †Pappomoropus Tong and Wang 2006
G. †Grangeria Zdansky 1930
+
†Grangeria canina Zdansky 1930
Fm. †Lophiodontidae Gill 1872
+
G. †Atalonodon Dal-Piaz 1929
G. †Lophiaspis Depéret 1910
G. †Lophiodochoerus Lemoine 1880
Subfm. †Lophiodontinae Cope 1881
G. †Paleomoropus Radinsky 1964
G. †Paralophiodon Dedieu 1977
+
Invalid names: Rhinocerolophiodon Fischer 1977 [synonym]
G. †Protomoropus Hooker and Dashzeveg 2004
+
†Protomoropus gabuniai Hooker and Dashzeveg 2004
Invalid names: Artionychia Osborn and Wortman 1893 [empty], Lophiodontoidea [empty], Perissonychia Osborn and Wortman 1893 [empty]
G. †Cardiolophus Gingerich 1991
+
†Cardiolophus radinskyi Gingerich 1991
+
Invalid names: Homogalax planeticus Gingerich 1980 [synonym]
Unr. †Ceratomorpha Wood 1937
+
G. †Breviodon Radinsky 1965
+
†Breviodon lumeiyiensis Huang and Qi 1982
G. †Fouchia Emry 1989
+
†Fouchia elyensis Emry 1989
G. †Karagalax Maas et al. 2001
+
†Karagalax mamikhelensis Maas et al. 2001
G. †Parabreviodon Reshetov 1975
Superfm. Rhinocerotoidea Gray 1821
+
Fm. †Amynodontidae Scott and Osborn 1883
+
G. †Amynodon Marsh 1877
G. †Amynodontopsis Stock 1933
G. †Andarakodon Averianov and Potapova 1996
Tr. †Metamynodontini Kretzoi 1942
G. †Orthocynodon Scott and Osborn 1882
G. †Rostriamynodon Wall and Manning 1986
G. †Sharamynodon Kretozi 1942
+
Invalid names: Lushiamynodon Chow and Xu 1965 [synonym]
Invalid names: Amynodontinae Scott and Osborn 1883 [empty], Cadurcotheriinae Kretzoi 1942 [invalid subgroup], Gigantamynodon Gromova 1954 [nomen dubium], Gigantamynodon cessator Gromova 1954 [nomen dubium], Gigantamynodon giganteus Xu 1961 [nomen dubium], Gigantamynodon maguanensis Qi 1992 [nomen dubium], Paramynodontinae Kretzoi 1942 [invalid subgroup], Rostriamynodontinae Wall and Manning 1986 [empty]
G. †Caenolophus Matthew and Granger 1925
+
†Caenolophus magnus Shi 1989
†Caenolophus obliquus Matthew and Granger 1925
†Caenolophus progressus Matthew and Granger 1925
†Caenolophus promissus Matthew and Granger 1925
Invalid names: Euryodon Xu et al. 1979 [synonym], Teilhardia Prothero and Schoch 1989 [synonym], Teilhardina Prothero and Schoch 1989 [synonym]
Fm. †Eggysodontidae Breuning 1923
+
G. †Allacerops Wood 1932
G. †Eggysodon Roman 1910
+
Invalid names: Engyodon Stehlin 1930 [synonym]
G. †Proeggysodon Bai and Wang 2012
Invalid names: Eggysodontinae Breuning 1923 [empty]
G. †Ergilia Gromova 1952
+
†Ergilia kazachstanica Gromov 1960
Fm. †Hyrachyidae Osborn 1892
+
G. †Hyrachyus Leidy 1871
+
Invalid names: Chasmotherium Rutimeyer [synonym], Colonoceras Marsh 1873 [synonym], Ephyrachyus Wood 1934 [synonym], Metahyrachyus Troxell 1922 [synonym]
G. †Subhyrachyus Gabunia 1999
Invalid names: Hyrachyus douglassi Wood 1934 [nomen dubium]
Fm. †Hyracodontidae Cope 1879
+
Subfm. †Alloceropinae Wood 1932
G. †Ardynia Matthew and Granger 1923
G. †Epitriplopus Wood 1927
G. †Hyracodon Leidy 1856
+
Invalid names: Mesamynodon Peterson 1931 [synonym]
G. †Ilianodon Chow 1961
G. †Pataecops Radinsky 1966
G. †Praeaceratherium Abel 1910
G. †Prohyracodon Koch 1897
+
Invalid names: Meninatherium Abel 1910 [synonym], Prothyracodon Scott and Osborn 1887 [synonym]
G. †Rhodopagus Radinsky 1965
G. †Triplopides Radinsky 1967
G. †Triplopus Cope 1880
+
Invalid names: Eotrigonias Wood 1927 [synonym]
G. †Ulania Qi 1990
G. †Yimengia Wang 1988
Invalid names: Allaceropinae Wood 1932 [empty], Hyracodontinae Steinmann and Doderlein 1890 [empty], Triplopodinae Cope 1881 [empty]
G. †Meschotherium Gabunia 1964
+
G. †Pakitherium Malkani et al. 2013
+
†Pakitherium shagalai Malkani et al. 2013
Fm. †Paraceratheriidae Osborn 1923
+
Subfm. †Forstercooperiinae Wood 1963
Subfm. †Paraceratheriinae Osborn 1923
+
Invalid names: Indricotheriinae Borissiak 1923 [synonym]
Invalid names: Forstercooperia ergiliinensis Gabunia and Dashzeveg 1974 [nomen dubium], Forstercooperia jigniensis Sahni and Khare 1973 [nomen dubium]
G. †Punjabitherium Khan 1971
Fm. Rhinocerotidae Gray 1821 [rhinoceros]
+
Subfm. †Aceratheriinae Dollo 1885
Tr. †Aceratheriini Dollo 1885
G. †Amphicaenopus Wood 1927
G. †Atelodus Pomel 1853
Subfm. †Diceratheriinae Dollo 1885
Subtr. †Elasmotheriina Bonaparte 1850
G. †Gulfoceras Albright 1999
Subfm. †Iranotheriinae Kretzoi 1942
G. †Miodiceros Giaourtsakis 2022
G. †Penetrigonias Tanner and Martin 1976
Subfm. Rhinocerotinae Gray 1821
G. †Subhyracodon Wood 1927
+
Invalid names: Anchisodon Cope 1879 [synonym], Caenopus Cope 1880 [synonym], Leptaceratherium Osborn 1898 [synonym]
G. †Teletaceras Hanson 1989
G. †Trigonias Lucas 1900
+
Invalid names: Procaenopus Figgins 1934 [synonym]
G. †Uintaceras Holbrook and Lucas 1997
G. †Woodoceras Prothero 2005
G. †Zalabis
Invalid names: Anchisodon tubifer Cope 1879 [nomen dubium], Dicerorhininae Simpson 1945 [empty], Forstercooperia crudus Gabounia 1977 [nomen dubium], Gigantamynodon akespensis Bayshashov 1993 [nomen dubium], Rhinoceros crassus Leidy 1858 [nomen dubium], Rhinoceros oregonensis Marsh 1873 [nomen dubium], Rhinoceros pacificus Leidy 1872 [nomen dubium], Teleoceratinae [empty]
G. †Siamolophus Ducrocq et al. 2006
+
†Siamolophus krabiense Ducrocq et al. 2006
G. †Symphysorrhachis Belyaeva 1954
+
G. †Toxotherium Wood 1961
+
†Toxotherium hunteri Wood 1961
+
Invalid names: Schizotheriodes jackwilsoni Schiebout 1977 [synonym], Toxotherium woodi Skinner and Gooris 1966 [synonym]
G. †Veragromovia Gabunia 1961
+
Invalid names: Huananodon You 1977 [nomen dubium]
G. †Schizotheroides Hough 1955
+
†Schizotheroides parvus Hough 1955
G. †Selenaletes Radinsky 1966
+
†Selenaletes scopaeus Radinsky 1966
Superfm. Tapiroidea Gill 1872
+
G. †Cambaylophus Kapur and Bajpai 2015
+
†Cambaylophus vastanensis Kapur and Bajpai 2015
Fm. †Deperetellidae Radinsky 1965
+
G. †Bahinolophus Tsubamoto 2005
G. †Deperetella Matthew and Granger 1925
+
Invalid names: Cristidentinus Zdansky 1930 [synonym], Diplolophodon Zdansky 1930 [synonym]
G. †Irdinolophus Dashzeveg and Hooker 1997
G. †Euryletes
G. †Haagella Heissig 1978
G. †Helaletes Marsh 1872
+
†Helaletes intermedius Osborn et al. 1878
+
Invalid names: Desmatotherium guyotii Scott 1883 [synonym]
†Helaletes medius Qi 1987
†Helaletes mongoliensis Osborn 1923
†Helaletes nanus Marsh 1871
+
Invalid names: Helaletes boops Marsh 1872 [synonym]
Fm. †Helaletidae Osborn 1892
+
G. †Desmatotherium Scott 1883
+
Invalid names: Chasmotheroides Wood 1934 [synonym]
G. †Jhagirilophus Thewissen et al. 2001
G. †Paracolodon Matthew and Granger 1925
G. †Vastanolophus Smith et al. 2015
Invalid names: Chasmotheriinae Viret 1858 [empty], Helaletinae Osborn 1892 [empty]
G. †Heptodon Cope 1882
+
†Heptodon calciculus Cope 1880
+
Invalid names: Lophiodon ventorum Cope 1880 [synonym]
†Heptodon minimus Qi 1987
†Heptodon niushanensis Chow and Li 1965
†Heptodon posticus Cope 1882
+
Invalid names: Heptodon brownorum Seton 1931 [synonym]
G. †Hesperaletes Colbert 2006
+
†Hesperaletes borineyi Colbert 2006
†Hesperaletes walshi Colbert 2006
Fm. Tapiridae Gray 1821 [tapir]
+
G. †Colodon Marsh 1890
+
Invalid names: Mesotapirus Scott and Osborn 1890 [synonym]
G. †Dilophodon Scott 1883
G. †Eotapirus Cerdeno and Ginsburg 1988
G. †Heteraletes Peterson 1931
G. †Miotapirus Schlaikjer 1937
G. †Nexuotapirus Albright 1998
G. †Palaeotapirus Filhol 1888
G. †Paratapirus Depéret and Douxami 1902
G. †Plesiocolopirus Schoch 1989
G. †Plesiotapirus Qiu et al. 1991
G. †Protapirus Filhol 1877
+
Invalid names: Tanyops Marsh 1894 [synonym]
G. †Tapiravus Marsh 1877
G. Tapirella Palmer 1903
+
Invalid names: Elasmognathus Gill 1865 [replaced]
Subfm. Tapirinae Burnett 1830
G. †Tapiriscus Kretzoi 1951
G. Tapirus Brisson 1762 [tapir]
+
Invalid names: Acrocodia Goldman 1913 [synonym], Cinchacus Gray 1873 [synonym], Pinchacus Hershkovitz 1954 [synonym], Rhinochoerus Wagler 1830 [synonym], Syspotamus Billberg 1827 [synonym], Tapir Blumenbach 1799 [synonym], Tapyra Liais 1872 [synonym]
G. †Teleolophus Matthew and Granger 1925
+
Invalid names: Pachylophus Tong and Lei 1984 [synonym]
G. †Thuliadanta Eberle 2005
+
†Thuliadanta mayri Eberle 2005
Invalid names: Heptodontidae Holbrook 1999 [empty]
Invalid names: Deperetellidea Radinsky 1965 [empty], Lophialetidea Matthew and Granger 1925 [empty], Tapiridea Burnett 1830 [empty]
G. †Homogalax Hay 1899
+
†Homogalax aureus Gingerich 1991
†Homogalax namadicus Dashzeveg 1979
†Homogalax primaevus Wortman 1896
†Homogalax protapirinus Wortman 1896
†Homogalax wutuensis Chow and Li 1965
Fm. †Isectolophidae Peterson 1919
+
G. †Chowliia Tong and Wang 2006
+
†Chowliia europea Bronnert et al. 2019
†Chowliia laoshanensis Tong and Wang 2006
G. †Gandheralophus Missiaen and Gingerich 2012
+
†Gandheralophus minor Missiaen and Gingerich 2012
†Gandheralophus robustus Missiaen and Gingerich 2012
Subfm. †Isectolophinae Peterson 1919
+
G. †Schizolophodon Peterson 1919
G. †Meridiolophus Bai et al. 2014
+
†Meridiolophus expansus Bai et al. 2014
G. †Sastrilophus Sahni and Khare 1971
+
†Sastrilophus dehmi Sahni and Khare 1971
Invalid names: Homogalaxinae Peterson 1919 [empty], Systemodontinae Osborn and Wortman 1894 [invalid subgroup]
G. †Isectolophus Scott and Osborn 1887
+
†Isectolophus annectens Scott and Osborn 1887
+
Invalid names: Homogalax uintensis Troxell 1922 [synonym], Isectolophus scotti Peterson 1919 [synonym], Schizolophodon cuspidens Peterson 1919 [synonym]
†Isectolophus latidens Osborn et al. 1878
+
Invalid names: Homogalax bridgerensis Troxell 1922 [synonym], Homogalax reliquius Qi 1987 [synonym], Isectolophus bogdulensis Reshetov 1979 [synonym]
†Isectolophus radinskyi Schoch 1983
Invalid names: Parisectolophus Peterson 1919 [synonym]
G. †Kalakotia Ranga Rao 1972
+
†Kalakotia simplicidentata Ranga Rao 1972
+
Invalid names: Chasmotherium mckennai Sahni and Khare 1973 [synonym], Schlosseria radinskyi Sahni and Khare 1973 [synonym]
Invalid names: Aulaxolophus Ranga Rao 1972 [synonym]
Fm. †Lophialetidae Matthew and Granger 1925
+
G. †Ampholophus Wang and Tong 1996
+
†Ampholophus luensis Wang and Tong 1996
G. †Eoletes Biryukov 1974
+
†Eoletes gracilis Biryukov 1974
†Eoletes tianshanicus Averianov and Godinot 2005
†Eoletes tshakpaktasensis Gabunia 1999
†Eoletes xianensis Zhang and Qi 1981
G. †Lophialetes Matthew and Granger 1925
+
†Lophialetes expeditus Matthew and Granger 1925
G. †Minchenoletes Wang et al. 2011
+
†Minchenoletes erlianensis Wang et al. 2011
G. †Schlosseria Matthew and Granger 1926
+
†Schlosseria hetaoyuanensis Tong and Lei 1984
†Schlosseria magister Matthew and Granger 1926
+
Invalid names: Lophialetes primus Cheng and Ma 1990 [synonym], Simplaletes sujiensis Qi 1980 [synonym], Simplaletes ulanshirehensis Qi 1980 [synonym]
Invalid names: Simplaletes Qi 1980 [synonym]
G. †Zhongjianoletes Ye 1983
+
Invalid names: Eoletidae Shoch 1989 [invalid subgroup]
G. †Systemodon Cope 1881
+
†Systemodon tapirinus Cope 1875
+
Invalid names: Systemodon cristatus Wortman 1896 [synonym]
Invalid names: Breviodontidae [empty], Moropomorpha [synonym], Rhodopagidae [empty]
G. †Teilhardimys Kretzoi and Kretzoi 2000
+
†Teilhardimys brisswalteri Hooker and Russell 2012
†Teilhardimys musculus Kretzoi and Kretzoi 2000
+
Invalid names: Microhyus musculus Teilhard de Chardin 1927 [replaced]
†Teilhardimys reisi Antunes et al. 1987
Invalid names: Microhyus Teilhard de Chardin 1927 [replaced]
+
Superfm. †Brontotherioidea Marsh 1873
+
Invalid names: Lambdotheriidae Cope 1889 [empty]
Invalid names: Chelodactyla Scott 1913 [empty], Equina Bonaparte 1850 [empty], Hippopotamus iravaticus Noetling 1897 [nomen dubium], Hyracotheriidae Cope 1881 [empty], Mesaxonia Marsh 1884 [empty], Rhinocerotida Owen 1845 [empty], Selenida McKenna and Bell 1997 [invalid subgroup], Stereopterna Ameghino 1889 [empty], Triplopodidae Cope 1881 [empty]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: phosphaticsubp
Maximum body mass: 1000 kgg
Minimum body mass: 10 kgg
Environment: terrestrialsubc
Locomotion: actively mobilec
Life habit: ground dwellingo
Diet: browsero
Diet 2: grazero
Reproduction: viviparoussubc
Created: 2005-08-26 13:57:26
Modified: 2005-08-26 15:57:26
Source: g = genus, o = order, subc = subclass, c = class, subp = subphylum
References: Nowak 1999, Ji et al. 2002, Hendy et al. 2009, Lillegraven 1979, Carroll 1988, Legendre 1988

Age range

Maximum range based only on fossils: base of the Late/Upper Paleocene to the top of the Holocene or 59.24000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 56.0 Ma

Collections (6610 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Late/Upper Paleocene59.24 - 56.0China (Nei Mongol) Perissodactyla indet. (133389)
Late/Upper Paleocene59.24 - 56.0France (Oise) Teilhardimys musculus (166598)
Thanetian59.24 - 56.0France (Champagne-Ardenne) Microhyus sp., Teilhardimys brisswalteri (68428) Teilhardimys brisswalteri (87943 137125) Teilhardimys brisswalteri, Paschatherium dolloi (87925)
Thanetian59.24 - 56.0Spain (Cataluña) Microhyus musculus, Paschatherium dolloi (79306) Microhyus musculus, Paschatherium dolloi, Paschatherium sp. (79305) Paschatherium dolloi (79304 79307)
Thanetian59.24 - 56.0Romania Paschatherium sp. (83307 138240 138241)
Thanetian59.24 - 56.0Spain (Catalunia) Paschatherium yvettae (182623)
Graybullian56.0 - 51.1USA (Wyoming) Hyracotherium cristatum, Hyracotherium pernix, Homogalax protapirinus (15701) Hyracotherium sp. (15875) Hyracotherium vasacciense (15878)
Lostcabinian56.0 - 51.1USA (Wyoming) Heptodon posticus, Lambdotherium popoagicum, Eotitanops borealis, Hyracotherium venticolum (15998) Hyracotherium vasacciense (16076) Hyracotherium vasacciense, Heptodon posticus, Lambdotherium popoagicum, Heptodon ventorum (15903) Hyracotherium vasacciense, Heptodon posticus, Lambdotherium popoagicum, Heptodon ventorum, Hyracotherium craspedotum (15915 15917 15930) Hyracotherium vasacciense, Heptodon posticus, Lambdotherium popoagicum, Hyracotherium craspedotum (16078) Hyracotherium vasacciense, Lambdotherium popoagicum (16068) Hyracotherium vasacciense, Lambdotherium popoagicum, Hyracotherium craspedotum (16073) Hyracotherium vasacciense, Orohippus sp., Heptodon posticus, Lambdotherium popoagicum, Hyracotherium craspedotum (15945) Hyracotherium vasacciense, Orohippus sp., Lambdotherium popoagicum (15901) Hyracotherium venticolum (16000) Lambdotherium popoagicum, Hyracotherium craspedotum, Hyracotherium venticolum (15997) Lambdotherium popoagicum, Hyracotherium venticolum (16005) Lambdotherium sp. (124856 124857) Xenicohippus craspedotum, Lambdotherium popoagicum, Eotitanops borealis, Hyracotherium venticolum (15999)
Lysitean56.0 - 51.1USA (Wyoming) Cardiolophus semihians (16002 16004) Hyracotherium sp. (16061) Hyracotherium vasacciense (15881)
Lysitean56.0 - 51.1USA (Colorado) Hyracotherium vasacciense, Hyracotherium craspedotum (15653) Lambdotherium popoagicum (15654)
Lysitean - Lostcabinian56.0 - 51.1USA (Wyoming) Heptodon sp., Hyracotherium sp. (15451)
Wasatchian56.0 - 51.1Mexico (Baja California) Eohippus angustidens, Hyracotherium sp. (15671)
Wasatchian56.0 - 51.1USA (Utah) Hyracotherium sp. (15716) Lambdotherium sp. (15715)
Wasatchian56.0 - 51.1USA (New Mexico) Homogalax protapirinus (15440 15460) Homogalax protapirinus, Hyracotherium tapirinum (15439) Hyracotherium angustidens (15598 15607 15611) Hyracotherium sp. (15442 15443 15446 15447 15448 15449 15453 15459 15481) Hyracotherium vasacciense, Xenicohippus sp., Hyracotherium tapirinum (15441) Xenicohippus osborni, Hipposyus sp. (15430) Xenicohippus sp., Hyracotherium vasacciense, Hyracotherium tapirinum (15663)
Wasatchian56.0 - 51.1Canada (Nunavut) Thuliadanta mayri (68720)
Wasatchian56.0 - 51.1USA (Wyoming) Arenahippus aemulor (15826 15835) Arenahippus aemulor, Homogalax protapirinus (15790) Arenahippus grangeri (15489 15497 15773 15884 16039 16040 16041 16042 16049 16050) Arenahippus grangeri, Cardiolophus radinskyi (15736 15747) Arenahippus pernix, Cardiolophus semihians, Homogalax protapirinus (118482) Arenahippus pernix, Cardiolophus semihians, Homogalax protapirinus, Hyracotherium sp. (15673) Cardiolophus radinskyi (46658 46723 46796) Eohippus index (15483) Eotitanops sp. (15911) Equidae indet., Helaletes sp., Lambdotherium popoagicum (15841) Heptodon brownorum (15910) Heptodon brownorum, Lambdotherium popoagicum, Hyracotherium boreale (15996) Heptodon calciculus (186940) Heptodon calciculus, Eotitanops sp., Lambdotherium primaevum (15565) Heptodon sp. (15642) Heptodon sp., Hyracotherium sp. (15597) Heptodon sp., Hyracotherium sp., Lambdotherium sp. (15931) Heptodon sp., Lambdotherium sp. (15958) Heptodon sp., Lambdotherium sp., Hyracotherium sp. (15842) Homogalax protapirinus (15474 46842) Homogalax protapirinus, Homogalax sp., Heptodon calciculus, Lambdotherium popoagicum, Hyracotherium sp. (15934) Homogalax protapirinus, Homogalax sp., Hyracotherium sp. (15933) Homogalax protapirinus, Hyracotherium angustidens (15468 15476 15478 15994 16055) Hyrachyus sp. (16057) Hyrachyus sp., Brontops sp. (16059) Hyracotherium aemulor (15511 15513 15514 15516 15517 15518 15520 15522 15523 15524 15531 15545 15767 15801 15804 15810 15827 15828 15829 46686 46770 46772 46775 46776 46779 46784 46791 46799 46801 46815 46828 46835 46838 46852) Hyracotherium aemulor, Cardiolophus radinskyi (46672 46693 46694 46794 46795 46844) Hyracotherium aemulor, Cardiolophus radinskyi, Homogalax protapirinus (15592 15683 46667 46673 46687 46689 46769 46792 46805 46814 46830 46831 46834 46836 46837) Hyracotherium aemulor, Cardiolophus semihians, Homogalax protapirinus (46800) Hyracotherium aemulor, Cardiolophus sp. (46656) Hyracotherium aemulor, Homogalax protapirinus (15699 15728 15741 15742 15743 15744 15745 15757 15758 15770 15798 15799 15800 15802 46641 46666 46674 46675 46688 46697 46698 46721 46724 46771 46773 46777 46778 46782 46789 46797 46798 46816 46817 46818 46819 46832 46833 46839 46840 46841) Hyracotherium aemulor, Hyracotherium grangeri (15510 15512 15532 15536 15538 15541 15544 15546) Hyracotherium aemulor, Hyracotherium grangeri, Cardiolophus radinskyi (46807) Hyracotherium aemulor, Hyracotherium grangeri, Cardiolophus radinskyi, Homogalax protapirinus (46809) Hyracotherium aemulor, Hyracotherium grangeri, Cardiolophus semihians, Heptodon calciculus, Perissodactyla indet. (15548) Hyracotherium aemulor, Hyracotherium grangeri, Perissodactyla indet. (15549) Hyracotherium aemulor, Hyracotherium grangeri, Systemodon sp. (15542) Hyracotherium angustidens (15467 15469 15471 15472 15491 15493 15504 15515 15717 15879 15889 15912 15932 15943 16016 16017 16020 16023 16028) Hyracotherium craspedotum (15457) Hyracotherium cristatum (15783) Hyracotherium cristatum, Hyracotherium pernix (15703 46701 46712 46714 46729 46734 46742 46755 46826 46827 46847) Hyracotherium cristatum, Hyracotherium pernix, Cardiolophus semihians (46764) Hyracotherium cristatum, Hyracotherium pernix, Cardiolophus semihians, Heptodon calciculus (46700 46849) Hyracotherium cristatum, Hyracotherium pernix, Cardiolophus semihians, Homogalax protapirinus (15692 15694 46683 46736 46761) Hyracotherium cristatum, Hyracotherium pernix, Heptodon calciculus (46699 46703 46711) Hyracotherium cristatum, Hyracotherium pernix, Homogalax protapirinus (15695 15697 15702 15704 15705 15706 15707 46682 46696 46704 46707 46718 46733 46740 46741 46745 46747 46748 46754 46760 46767 46825) Hyracotherium cristatum, Hyracotherium pernix, Homogalax protapirinus, Heptodon calciculus (46746) Hyracotherium cristatum, Hyracotherium pernix, Hyracotherium grangeri, Cardiolophus semihians, Homogalax protapirinus, Heptodon calciculus (46692) Hyracotherium grangeri (15458 15495 15503 15505 15507 15509 15540 15543 15547 15550 15587 15588 15589 15734 15738 15739 15740 15748 15750 15751 15752 15754 15755 15763 15764 15765 15766 15772 15774 15775 15778 15786 15791 15794 15795 15808 15809 15815 15816 15820 15825 15944 46637 46638 46639 46640 46642 46644 46647 46648 46660 46661 46662 46664 46670 46681 46684 46685 46691 46725 46726 46727 46744 46774 46781 46785 46793 46804 46813 46821 46822 46823 46824) Hyracotherium grangeri, Cardiolophus radinskyi (15586 15590 15726 15727 15731 15732 15733 15735 15737 15746 15749 15753 15777 15781 15787 15788 15792 15813 15822 15824 46636 46643 46645 46646 46650 46652 46657 46659 46668 46669 46671 46680 46690 46768 46780 46783 46787 46788 46790 46803 46806 46812) Hyracotherium grangeri, Cardiolophus radinskyi, Homogalax protapirinus (15762 15779 15796 15819 46808) Hyracotherium grangeri, Heptodon calciculus (15560) Hyracotherium grangeri, Heptodon posticus, Lambdotherium primaevum (15566) Hyracotherium grangeri, Homogalax planeticus (15594) Hyracotherium grangeri, Homogalax protapirinus (15730 15797 15818 15821 46786) Hyracotherium grangeri, Homogalax sp. (15793) Hyracotherium grangeri, Lambdotherium primaevum (15568) Hyracotherium index (16056) Hyracotherium index, Heptodon sp. (15937) Hyracotherium index, Hyracotherium aemulor, Hyracotherium grangeri, Heptodon calciculus (15551 15553 15554) Hyracotherium index, Hyracotherium grangeri, Heptodon posticus, Heptodon calciculus (15555) Hyracotherium index, Hyracotherium pernix, Hyracotherium grangeri, Heptodon calciculus (15557) Hyracotherium index, Hyracotherium pernix, Hyracotherium grangeri, Heptodon posticus, Heptodon calciculus (15556) Hyracotherium index, Hyracotherium venticolum, Orohippus proteros, Heptodon calciculus, Lambdotherium popoagicum, Eotitanops borealis (15454) Hyracotherium montanum (15581) Hyracotherium pernix (15696 15803 15811 15812 46695 46702 46709 46710 46713 46719 46720 46728 46848 46851) Hyracotherium pernix, Cardiolophus semihians, Homogalax protapirinus (46735 46763) Hyracotherium pernix, Cardiolophus semihians, Homogalax protapirinus, Heptodon calciculus (46810) Hyracotherium pernix, Cardiolophus semihians, Homogalax protapirinus, Hyracotherium cristatum (15687) Hyracotherium pernix, Heptodon calciculus (46706 46715) Hyracotherium pernix, Heptodon posticus, Heptodon calciculus, Heptodon sp. (15564) Hyracotherium pernix, Homogalax protapirinus (15698 15768 15769 15782 15785 15805 15806 15814 46716 46717 46730 46731 46732 46738 46739 46743 46749 46753 46756 46757 46758 46759 46762 46765 46766 46820 46843 46845 46846) Hyracotherium pernix, Homogalax protapirinus, Heptodon calciculus (46811) Hyracotherium pernix, Hyracotherium grangeri, Heptodon calciculus (15558) Hyracotherium pernix, Hyracotherium grangeri, Heptodon posticus, Lambdotherium primaevum (15567) Hyracotherium pernix, Lambdotherium popoagicum (46705) Hyracotherium pernix, Paleomoropus jepseni, Homogalax protapirinus, Hyracotherium cristatum (15807) Hyracotherium sandrae (15760 15771 15780) Hyracotherium sandrae, Hyracotherium grangeri (15759 15761) Hyracotherium sp. (15462 15572 15596 15657 15688 15720 15721 15722 15723 15724 15846 15891 15892 15893 15894 15898 15900 15914 15936 16008 16060 16391 46737 46751 46752) Hyracotherium sp., Helaletidae indet. (15577) Hyracotherium sp., Heptodon calciculus, Helaletidae indet. (15552) Hyracotherium sp., Heptodon sp. (15689) Hyracotherium sp., Heptodon sp., Helaletidae indet. (15559) Hyracotherium sp., Heptodon ventorum (15963) Hyracotherium sp., Homogalax protapirinus (15573 15593 15682 46829) Hyracotherium sp., Homogalax sp. (15729 15756) Hyracotherium sp., Hyracotherium pernix (15455) Hyracotherium sp., Lambdotherium popoagicum (27268) Hyracotherium sp., Xenicohippus grangeri, Helaletidae indet. (15575 15578) Hyracotherium sp., Xenicohippus grangeri, Homogalax sp., Heptodon calciculus (15576) Hyracotherium vasacciense (16063) Hyracotherium vasacciense, Heptodon calciculus, Lambdotherium popoagicum, Hyracotherium craspedotum (15852) Hyracotherium vasacciense, Heptodon posticus, Heptodon calciculus, Lambdotherium popoagicum, Eotitanops borealis, Hyracotherium craspedotum (15486) Hyracotherium vasacciense, Heptodon ventorum (15661) Hyracotherium vasacciense, Homogalax sp., Lambdotherium popoagicum, Heptodon ventorum, Hyracotherium craspedotum (15479) Hyracotherium vasacciense, Hyrachyus modestus, Heptodon posticus, Heptodon calciculus, Lambdotherium popoagicum, Hyracotherium craspedotum (15839) Hyracotherium vasacciense, Hyrachyus modestus, Lambdotherium popoagicum, Heptodon ventorum, Hyracotherium craspedotum (15710) Hyracotherium vasacciense, Hyracotherium craspedotum, Heptodon calciculus, Lambdotherium popoagicum (15935) Hyracotherium vasacciense, Hyracotherium index, Heptodon calciculus, Hyracotherium craspedotum (15964) Hyracotherium vasacciense, Lambdotherium popoagicum (15665 15711 15714 15848 15957 16067) Hyracotherium vasacciense, Lambdotherium sp. (16075) Hyracotherium vasacciense, Orohippus proteros, Homogalax sp., Heptodon sp., Lambdotherium popoagicum, Eotitanops borealis (15902) Hyracotherium venticolum (15927 15928 15946) Hyracotherium venticolum, Heptodon brownorum, Palaeosyops borealis (15904) Hyracotherium venticolum, Lambdotherium popoagicum (15908) Lambdotherium popoagicum (15664 15666) Lambdotherium popoagicum, Eotitanops sp., Hyracotherium venticolum, Heptodon brownorum (15906) Lambdotherium popoagicum, Heptodon brownorum (15905) Lambdotherium popoagicum, Hyracotherium sp. (16074) Lambdotherium popoagicum, Hyracotherium vasacciense (15456) Lambdotherium sp. (15569 15853 16009) Lambdotherium sp., Hyracotherium sp. (15843 16062 16392) Lophiotherium vasacciensis, Perissodactyla indet. (131859) Orohippus sp., Heptodon posticus, Helaletes nanus, Lambdotherium popoagicum, Heptodon ventorum, Hyracotherium craspedotum (15942) Palaeosyops borealis (15909) Perissodactyla indet. (15886) Protorohippus venticolum (186942 186943 186945 186947 186948) Protorohippus venticolum, Equidae indet. (186932 186933 186935 186936 186939 186944 186946 186952) Protorohippus venticolum, Equidae indet., Heptodon calciculus (186937) Protorohippus venticolum, Heptodon calciculus (186951) Protorohippus venticolum, Heptodon posticus, Lambdotherium popoagicum, Hyracotherium vasacciense, Heptodon ventorum, Hyracotherium craspedotum (15662) Xenicohippus craspedotum (186950) Xenicohippus craspedotum, Equidae indet. (186934) Xenicohippus craspedotum, Equidae indet., Lambdotherium popoagicum (186931)
Wasatchian56.0 - 51.1Canada (British Columbia) Heptodon sp. (161203)
Wasatchian56.0 - 51.1Canada (Northwest Territories) Equidae indet., Lambdotherium sp., Hyrachyus sp. (15583)
Wasatchian56.0 - 51.1USA (Colorado) Arenahippus grangeri, Hyracotherium angustidens (15450 15480) Eotitanops brownianus, Eotitanops borealis (15651) Eotitanops minimus (15882) Heptodon calciculus (15302) Heptodon sp. (15270 15407) Heptodon sp., Lambdotherium popoagicum, Hyracotherium vasacciense (15649) Homogalax protapirinus (15275) Homogalax sp. (15425) Hyracotherium angustidens (15865) Hyracotherium angustidens, Homogalax primaevus (15485 15725 15845 15887) Hyracotherium craspedotum, Heptodon calciculus, Heptodon sp. (15293) Hyracotherium craspedotum, Hyracotherium vasacciense (15652) Hyracotherium grangeri (15429) Hyracotherium sp. (15260 15261 15262 15263 15264 15266 15267 15268 15269 15271 15278 15282 15283 15285 15286 15289 15292 15294 15296 15297 15298 15300 15301 15304 15307 15310 15311 15313 15314 15315 15317 15319 15320 15324 15326 15329 15332 15340 15343 15349 15354 15355 15357 15360 15363 15365 15366 15367 15368 15371 15374 15375 15376 15377 15379 15381 15382 15383 15384 15385 15387 15388 15389 15391 15392 15396 15397 15401 15403 15405 15416 15420 15422 15428 15484 15646 15836 15854 15855 15859 15860 15861 15862 15870 15871 15873 15874) Hyracotherium sp., Heptodon calciculus (15325 15393 15395 15426) Hyracotherium sp., Heptodon calciculus, Lambdotherium popoagicum (15338) Hyracotherium sp., Heptodon posticus (15410) Hyracotherium sp., Heptodon posticus, Lambdotherium popoagicum (15413) Hyracotherium sp., Heptodon sp. (15220 15221 15272 15350 15372) Hyracotherium sp., Homogalax primaevus (15864 15868 15869) Hyracotherium sp., Homogalax protapirinus (15273) Hyracotherium sp., Homogalax protapirinus, Heptodon sp. (15277) Hyracotherium sp., Homogalax protapirinus, Homogalax sp. (15274) Hyracotherium sp., Lambdotherium popoagicum (15331 15335 15409 15414 15418 15421) Hyracotherium sp., Xenicohippus grangeri (15290 15295 15370) Hyracotherium sp., Xenicohippus grangeri, Heptodon sp. (15225) Hyracotherium sp., Xenicohippus osborni (15399 15404) Hyracotherium sp., Xenicohippus osborni, Xenicohippus sp. (15402) Hyracotherium sp., Xenicohippus sp. (15309) Hyracotherium vasacciense, Lambdotherium priscum, Lambdotherium magnum, Lambdotherium popoagicum, Hyracotherium craspedotum (15650) Lambdotherium popoagicum (15328 15330 15337 15342 15411 15415 15419) Xenicohippus grangeri (15362 15406) Xenicohippus osborni, Eohippus angustidens (15655)
Wasatchian56.0 - 51.1USA (Mississippi) Hyracotherium sp. (15719)