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Formicomyrmida

Insecta - Hymenoptera - Formicidae

Taxonomy
Formicomyrmida was named by Oberski et al. (2025). It is extant.

It was assigned to Formicia by Oberski et al. (2025).

Synonymy list
YearName and author
2025Formicomyrmida Oberski et al.

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaGravenhorst 1843
—Deuteropoda
—Mandibulata
—PancrustaceaZrzavý and Štys 1997
—AltocrustaceaRegier et al. 2010
superclassAllotriocarida
—Athalassocarida
—LabiocaridaSchwentner et al. 2017
superclassHexapodaLatreille 1825
—Cercofilata
classInsecta
—Dicondylia
RankNameAuthor
—Paranotalia
subclassPterygota()
—NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
—Holometabola
—Hymenopterida
orderHymenoptera
—Unicalcarida
suborderApocritaGerstaecker 1867
—Aculeata()
superfamilyFormicoidea
familyFormicidae()
—Antennoclypeata
—Euformicia
—Doryloformicia
—Formicia
—Formicomyrmida
—Formicomyrmida

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. Formicomyrmida Oberski et al. 2025
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Unr. Formicomorpha Oberski et al. 2025
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Subfm. †Formiciinae Lutz 1986
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G. †Formicium Westwood 1854
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†Formicium berryi Carpenter 1929
†Formicium brodiei Westwood 1854
Invalid names: Eoponera Carpenter 1929 [synonym]
G. †Titanomyrma Archibald et al. 2011
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†Titanomyrma gigantea Lutz 1986
†Titanomyrma lubei Archibald et al. 2011
†Titanomyrma simillima Lutz 1986
Subfm. Formicinae Latreille 1802
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G. †Antiquiformica Wu et al. 2024
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†Antiquiformica alata Wu et al. 2024
G. †Attopsis Heer 1849
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†Attopsis longipennis Heer 1849
†Attopsis longipes Heer 1867
Tr. Camponotini Forel 1878
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G. Calomyrmex Mayr 1861
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G. Camponotus Mayr 1861
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Camponotus festinatus Buckley 1866
Camponotus herculeanus Linnaeus 1758
Camponotus luteus Smith 1858
Camponotus mitis Smith 1892
†Camponotus parvus Perfilieva 2022
G. †Chimaeromyrma Dlussky 1988
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G. Colobopsis Mayr 1861
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Colobopsis vitrea Smith 1860
G. †Eocamponotus Boudinot 2024
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†Eocamponotus mengei Mayr 1868
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Invalid names: Camponotus igneus Mayr 1868 [synonym]
G. Polyrhachis Smith 1857
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†Polyrhachis annosa Wappler et al. 2009
†Polyrhachis celebensis Viehmeyer 1913
†Polyrhachis cincta Viehmeyer 1913
†Polyrhachis circumdata Viehmeyer 1913
Polyrhachis dahlii Forel 1901
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Invalid names: Polyrhachis restituta Viehmeyer 1913 [synonym]
†Polyrhachis excitata Viehmeyer 1913
†Polyrhachis inclusa Viehmeyer 1912
Polyrhachis levior Roger 1863
Polyrhachis rastellata Latreille 1802
Polyrhachis sexspinosa Latreille 1802
†Polyrhachis subfossa Viehmeyer 1913
Invalid names: Polyrhachis continua revocata Viehmeyer 1913 [invalid subgroup], Polyrhachis restituta conclusa Viehmeyer 1913 [synonym]
Invalid names: Pseudocamponotus Carpenter 1930 [nomen dubium], Pseudocamponotus elkoanus Carpenter 1930 [nomen dubium]
G. †Camponotites Steinbach 1967
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†Camponotites ambon Zhang 1989
†Camponotites cockerelli Donisthorpe 1920
†Camponotites compactus Förster 1891
†Camponotites crozei Riou 1999
†Camponotites fuscipennis Carpenter 1930
†Camponotites gracilis Zhang 1989
†Camponotites lignitus Germar 1837
†Camponotites longiventris Theobald 1937
†Camponotites longus Zhang 1989
†Camponotites microcephalus Carpenter 1930
†Camponotites novotnyi Samsinak 1967
†Camponotites obesus Piton 1935
†Camponotites penninervis Theobald 1937
†Camponotites petrifactus Carpenter 1930
†Camponotites pictus Zhang et al. 1994
†Camponotites plenus Zhang 1989
†Camponotites silvestris Steinbach 1967
†Camponotites steinbachi Dlussky et al. 2011
†Camponotites theobaldi Özdikmen 2010
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Invalid names: Camponotus saussurei Theobald 1937 [replaced]
†Camponotites tokunagai Naora 1933
†Camponotites ullrichi Bachmayer 1960
†Camponotites vehemens Förster 1891
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Invalid names: Camponotus miserabilis Förster 1891 [synonym]
†Camponotites vetus Scudder 1877
Invalid names: Camponotites Dlussky 1981 [synonym], Formica pinguicula radobojana Heer 1849 [synonym], Rabidia Hong 1984 [synonym]
G. †Drymomyrmex Wheeler 1915
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†Drymomyrmex claripennis Wheeler 1915
†Drymomyrmex fuscipennis Wheeler 1915
†Drymomyrmex rasnitsyni Radchenko 2021
Tr. Formicini Latreille 1802
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G. Cataglyphis Förster 1850
G. †Cataglyphoides Dlussky 2008
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G. †Conoformica Dlussky 2008
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G. Formica Linnaeus 1758
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†Formica alsatica Theobald 1937
†Formica annosa LaPolla and Greenwalt 2015
†Formica arcana Scudder 1877
†Formica auxillacensis Piton 1935
†Formica bauckhorni Meunier 1915
†Formica biamoensis Dlussky et al. 2015
†Formica buphthalma Novák 1878
†Formica cantalica Piton 1935
†Formica ceps Zhang 1989
†Formica cockerelli Carpenter 1930
†Formica demersa Heer 1849
†Formica eoptera Cockerell 1923
†Formica flavifemoralis Zhang et al. 1994
†Formica flori Mayr 1868
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Invalid names: Formica (Serviformica) antiqua Dlussky 1967 [synonym], Formica (Serviformica) baltica Dlussky 1967 [synonym], Formica (Serviformica) parvula Dlussky 1967 [synonym], Formica egecomerta Özdikmen 2010 [synonym]
Formica fusca Linnaeus 1758
†Formica globiventris Heer 1849
†Formica grandis Carpenter 1930
†Formica gravida Heer 1849
†Formica gustawi Dlussky 2002
†Formica heteroptera Cockerell 1920
†Formica horrida Wheeler 1915
†Formica immersa Heer 1849
†Formica kutscheri Dlussky 2008
†Formica latinodosa Theobald 1937
†Formica lavateri Heer 1849
†Formica linquensis Zhang 1989
†Formica longicollis Heer 1849
†Formica macrocephala Heer 1849
†Formica macrophthalma Heer 1849
†Formica maculipennis Théobald 1935
†Formica martynovi Popov 1933
†Formica obscura Heer 1849
†Formica orbata Heer 1849
†Formica ovala Zhang 1989
†Formica palaeopolonica Dlussky 2008
†Formica paleosibirica Dlussky et al. 2015
†Formica parexsecta Dlussky and Putyatina 2014
†Formica phaethusa Wheeler 1915
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Invalid names: Formica clymene Wheeler 1915 [synonym]
†Formica pitoni Théobald 1935
†Formica primitiva Heer 1849
†Formica primordialis Heer 1849
†Formica procera Heer 1849
†Formica pulchella Heer 1849
†Formica radchenkoi Dlussky 2008
†Formica ribbeckei Radchenko and Perkovsky 2021
†Formica robusta Carpenter 1930
†Formica sepulta Theobald 1937
†Formica serresi Theobald 1937
†Formica seuberti Heer 1849
†Formica strangulata Wheeler 1915
†Formica surinamensis Berendt 1830
†Formica tripartita Theobald 1937
†Formica ungeri Heer 1849
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Invalid names: Formica acuminata Heer 1849 [synonym], Formica aemula Heer 1867 [synonym], Formica kollari Heer 1867 [synonym], Formica oblita Heer 1867 [synonym], Formica redtenbacheri Heer 1849 [synonym], Ponera elongatula Heer 1849 [synonym]
†Formica zherikhini Dlussky 2008
Invalid names: Serviformica Forel 1913 [synonym]
G. †Glaphyromyrmex Wheeler 1915
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G. †Protoformica Dlussky 1967
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Tr. Gesomyrmecini Forel 1912
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G. Gesomyrmex Mayr 1868
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†Gesomyrmex bremii Heer 1849
†Gesomyrmex breviceps Dlussky et al. 2009
†Gesomyrmex flavescens Dlussky et al. 2009
†Gesomyrmex gallicus Aria et al. 2023
†Gesomyrmex germanicus Dlussky et al. 2009
†Gesomyrmex hoernesi Mayr 1868
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Invalid names: Dimorphomyrmex mayri Wheeler 1915 [synonym], Dimorphomyrmex theryi Emery 1905 [synonym], Gesomyrmex annectens Wheeler 1915 [synonym], Gesomyrmex curiosus Dlussky et al. 2009 [synonym], Gesomyrmex incertus Dlussky et al. 2015 [synonym]
†Gesomyrmex macrops Dlussky et al. 2015
†Gesomyrmex magnus Dlussky et al. 2015
†Gesomyrmex pulcher Dlussky et al. 2009
Invalid names: Dimorphomyrmex André 1892 [synonym]
G. †Prodimorphomyrmex Wheeler 1915
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G. †Sicilomyrmex Wheeler 1915
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†Sicilomyrmex corniger Emery 1891
Invalid names: Sicelomyrmicini Wheeler 1929 [synonym]
G. †Heeridris Dlussky and Putyatina 2014
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†Heeridris croaticus Dlussky and Putyatina 2014
G. †Imhoffia Heer 1849
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†Imhoffia nigra Heer 1849
†Imhoffia pallida Heer 1864
G. †Kyromyrma Grimaldi and Agosti 2000
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†Kyromyrma neffi Grimaldi and Agosti 2000
Tr. Lasiini Ashmead 1905
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G. Euprenolepis Emery 1906
G. Lasius Fabricius 1805
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†Lasius anthracinus Heer 1867
†Lasius chambonensis Théobald 1935
†Lasius crispus Théobald 1935
†Lasius epicentrus Theobald 1937
†Lasius globularis Heer 1849
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Invalid names: Formica capito Heer 1856 [synonym]
†Lasius glom LaPolla and Greenwalt 2015
†Lasius inflatus Zhang 1989
†Lasius longaevus Heer 1849
†Lasius longipennis Heer 1849
†Lasius mordicus Zhang 1989
†Lasius nemorivagus Wheeler 1915
†Lasius oblongus Assmann 1870
†Lasius occultatus Heer 1849
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Invalid names: Formica minutula Heer 1849 [synonym], Formica obliterata Heer 1849 [synonym], Formica pumila Heer 1849 [synonym]
†Lasius ophthalmicus Heer 1849
†Lasius peritulus Cockerell 1927
†Lasius punctulatus Mayr 1868
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Invalid names: Lasius edentatus Mayr 1868 [synonym]
†Lasius tertiarius Zalessky 1949
†Lasius truncatus Zhang 1989
†Lasius validus Zhang 1989
†Lasius vetulus Dlussky 1981
†Lasius vulgaris Perfilieva 2022
Invalid names: Formica macrocephala radobojana Heer 1849 [synonym], Tylolasius Zhang 1989 [synonym]
G. Metalasius Boudinot et al. 2022
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†Metalasius pumilus Mayr 1868
G. Nylanderia Emery 1906
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†Nylanderia pygmaea Mayr 1868
†Nylanderia vetula LaPolla and Dlussky 2010
G. Paratrechina Motschoulsky 1863
G. Prenolepis Mayr 1861
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†Prenolepis henschei Mayr 1868
G. Pseudolasius Emery 1887
G. †Sussudio Boudinot et al. 2022
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†Sussudio boreus Wheeler 1915
G. Zatania LaPolla et al. 2012
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†Zatania electra LaPolla et al. 2012
Invalid names: Prenolepidini Forel 1912 [synonym]
G. †Leucotaphus Donisthorpe 1920
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†Leucotaphus donisthorpei Dlussky and Perfilieva 2014
†Leucotaphus gurnetensis Cockerell 1915
G. Myrmecorhynchus André 1896
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†Myrmecorhynchus novaeseelandiae Kaulfuss and Dlussky 2016
Tr. Oecophyllini Emery 1895
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G. †Eoecophylla Archibald et al. 2024
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†Eoecophylla quilchenensis Archibald et al. 2024
G. Oecophylla Smith 1860 [weaver ant]
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†Oecophylla atavina Cockerell 1915
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Invalid names: Oecophylla perdita Cockerell 1915 [synonym]
†Oecophylla bartoniana Cockerell 1920
†Oecophylla brischkei Mayr 1868
†Oecophylla crassinoda Wheeler 1922
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Invalid names: Oecophylla brevinodis Wheeler 1915 [replaced]
†Oecophylla eckfeldiana Dlussky et al. 2008
†Oecophylla kraussei Dlussky and Rasnitsyn 1999
†Oecophylla leakeyi Wilson and Taylor 1964
†Oecophylla longiceps Dlussky et al. 2008
†Oecophylla macroptera Dlussky 1981
†Oecophylla megarche Cockerell 1915
†Oecophylla obesa Heer 1849
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Invalid names: Attopsis anthracina Heer 1849 [synonym], Attopsis nigra Heer 1849 [synonym], Formica pinguis Heer 1849 [synonym]
†Oecophylla praeclara Förster 1891
†Oecophylla sicula Emery 1891
Oecophylla smaragdina Fabricius 1775
†Oecophylla superba Theobald 1937
†Oecophylla taurica Perfilieva et al. 2017
Tr. Plagiolepidini Forel 1886
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G. Acropyga Roger 1862
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†Acropyga glaesaria LaPolla 2005
G. Lepisiota Santschi 1926
G. Plagiolepis Mayr 1861
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†Plagiolepis kuenowi Mayr 1868
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Invalid names: Plagiolepis balticus Dlussky 1997 [synonym], Rhopalomyrmex pygmaeus Mayr 1868 [synonym]
†Plagiolepis labilis Emery 1891
†Plagiolepis minutissima Dlussky 2002
†Plagiolepis paradoxa Dlussky 2010
†Plagiolepis solitaria Mayr 1868
†Plagiolepis wheeleri Dlussky 2010
Invalid names: Rhopalomyrmex Mayr 1868 [synonym]
G. †Protrechina Wilson 1985
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†Protrechina carpenteri Wilson 1985
Invalid names: Andrena tertiaria Meunier 1920 [nomen dubium], Camponotinae Forel 1878 [synonym], Curtipalpulus Hong 2002 [nomen nudum], Eoleptocerites Hong 2002 [nomen nudum], Formica fragilis Heer 1867 [nomen dubium], Fushuniformica Hong 2002 [nomen nudum], Huaxiaformica Hong 2002 [nomen nudum], Leptogasteritus Hong 2002 [nomen nudum], Liaoformica Hong 2002 [nomen nudum], Longiformica Hong 2002 [nomen nudum], Magnogasterites Hong 2002 [nomen nudum], Orbicapitia Hong 2002 [nomen nudum], Ovalicapito Hong 2002 [nomen nudum], Ovaligastrula Hong 2002 [nomen nudum], Palaeosminthurus Pierce and Gibron 1962 [nomen dubium], Palaeosminthurus juliae Pierce and Gibron 1962 [nomen dubium], Sinoformica Hong 2002 [nomen nudum], Sinotenuicapito Hong 2002 [nomen nudum], Wilsonia Hong 2002 [nomen nudum]
Unr. Myrmicomorpha Oberski et al. 2025
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Subfm. Ectatomminae Emery 1895
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Tr. Ectatommini Emery 1895
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G. Ectatomma Smith 1858
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†Ectatomma gracile Emery 1891
G. Gnamptogenys Roger 1863
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†Gnamptogenys casca Lattke 2002
†Gnamptogenys europaea Mayr 1868
†Gnamptogenys levinates Baroni Urbani 1980
†Gnamptogenys pristina Baroni Urbani 1980
†Gnamptogenys rohdendorfi Dlussky 2009
G. Poneracantha Emery 1897
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†Poneracantha brunoi Lattke 2002
G. Rhytidoponera Mayr 1862
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Rhytidoponera araneoides Le Guillou 1842
†Rhytidoponera gibsoni Kaulfuss and Dlussky 2016
†Rhytidoponera kirghizorum Dlussky 1981
†Rhytidoponera waipiata Kaulfuss and Dlussky 2016
G. †Electroponera Wheeler 1915
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†Electroponera dubia Wheeler 1915
G. †Pseudectatomma Dlussky and Wedmann 2012
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†Pseudectatomma eocenica Dlussky and Wedmann 2012
†Pseudectatomma striatula Dlussky and Wedmann 2012
Subfm. Myrmicinae Lepeletier de Saint-Fargeau 1835 [ant]
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G. †Afromyrma Dlussky et al. 2004
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†Afromyrma petrosa Dlussky et al. 2004
G. †Agastomyrma Dlussky et al. 2015
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†Agastomyrma laticeps Dlussky et al. 2015
Tr. Attini Smith 1858
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G. Acanthognathus Mayr 1887
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†Acanthognathus poinari Baroni Urbani 1994
Subtr. Attina Smith 1858
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G. Acromyrmex Mayr 1865
G. Apterostigma Mayr 1865
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†Apterostigma eowilsoni Schultz 2007
G. Atta Fabricius 1804
G. Cyphomyrmex Mayr 1862
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†Cyphomyrmex maya de Andrade 2003
†Cyphomyrmex taino de Andrade 2003
G. Mycetomoellerius Solomon et al. 2019
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†Mycetomoellerius primaevus Baroni Urbani 1980
G. Trachymyrmex Forel 1893
G. Basiceros Schulz 1906
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†Basiceros enana Fiorentino et al. 2025
G. Cephalotes Latreille 1802
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†Cephalotes alveolatus Vierbergen and Scheven 1995
†Cephalotes bloosi Baroni Urbani and de Andrade 1999
†Cephalotes brevispineus de Andrade 1999
†Cephalotes caribicus de Andrade 1999
†Cephalotes dieteri Baroni Urbani 1999
†Cephalotes hispaniolicus de Andrade 1999
†Cephalotes integerrimus Vierbergen and Scheven 1995
†Cephalotes jansei Vierbergen and Scheven 1995
†Cephalotes maya de Andrade 1999
†Cephalotes obscurus Vierbergen and Scheven 1995
†Cephalotes olmecus de Andrade 1999
†Cephalotes poinari Baroni Urbani 1999
†Cephalotes resinae de Andrade 1999
†Cephalotes serratus Vierbergen and Scheven 1995
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Invalid names: Exocryptocerus elevatus Vierbergen and Scheven 1995 [synonym], Exocryptocerus truncatus Vierbergen and Scheven 1995 [synonym]
†Cephalotes squamosus Vierbergen and Scheven 1995
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Invalid names: Zacryptocerus pseudoaztecus Vierbergen and Scheven 1995 [synonym]
†Cephalotes sucinus de Andrade 1999
†Cephalotes taino de Andrade 1999
†Cephalotes ventriosus de Andrade 1999
Invalid names: Exocryptocerus Vierbergen and Scheven 1995 [synonym], Zacryptocerus Ashmead 1905 [synonym]
G. Pheidole Westwood 1840
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†Pheidole anticua Casadei-Ferreira et al. 2019
†Pheidole chaan Varela-Hernández and Flores-Zapoteco 2024
†Pheidole cordata Holl 1829
Pheidole pallidula Nylander 1849
†Pheidole pauchil Varela-Hernández and Riquelme 2021
Pheidole plagiaria Smith 1860
†Pheidole primigenia Baroni Urbani 1995
†Pheidole rasnitsyni Dubovikoff 2011
†Pheidole tertiaria Carpenter 1930
†Pheidole tethepa Wilson 1985
G. Strumigenys Smith 1860
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†Strumigenys electrina de Andrade 1994
†Strumigenys pilosula de Andrade 2007
†Strumigenys poinari Baroni Urbani and de Andrade 2007
†Strumigenys schleeorum Baroni Urbani 1994
Invalid names: Smithistruma Brown 1948 [synonym]
Invalid names: Cephalotini Smith 1949 [synonym], Dacetini Forel 1892 [synonym], Pheidolini Emery 1877 [synonym]
G. †Biamomyrma Dlussky et al. 2015
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†Biamomyrma lata Dlussky et al. 2015
†Biamomyrma rugosa Dlussky et al. 2015
†Biamomyrma zherikhini Dlussky et al. 2015
G. †Bilobomyrma Radchenko and Dlussky 2013
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†Bilobomyrma baltica Radchenko and Dlussky 2013
†Bilobomyrma ukrainica Radchenko and Dlussky 2013
G. †Boltonidris Radchenko and Dlussky 2012
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†Boltonidris mirabilis Radchenko and Dlussky 2012
G. †Cephalomyrmex Carpenter 1930
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†Cephalomyrmex rotundatus Carpenter 1930
G. †Cephites Heer 1849
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†Cephites fragilis Heer 1849
†Cephites oeningensis Heer 1849
Tr. Crematogastrini Forel 1893
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G. Cardiocondyla Emery 1869
G. Carebara Westwood 1840
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†Carebara antiqua Mayr 1868
†Carebara bohemica Novák 1878
†Carebara groehni Radchenko and Dlussky 2019
†Carebara kutscheri Radchenko et al. 2019
†Carebara nitida Dlussky 2002
†Carebara rugiceps Heer 1849
†Carebara schossnicensis Assmann 1870
†Carebara sophiae Emery 1891
†Carebara thorali Théobald 1937
†Carebara ucrainica Dlussky 2002
Invalid names: Aeromyrma Forel 1891 [synonym], Erebomyrma Wheeler 1903 [synonym], Oligomyrmex Mayr 1867 [synonym], Pheidologeton Mayr 1862 [synonym]
G. Cataulacus Smith 1853
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†Cataulacus planiceps Emery 1891
†Cataulacus plebeius de Andrade and Baroni Urbani 2004
†Cataulacus resinosus Viehmeyer 1913
†Cataulacus silvestrii Emery 1891
G. Crematogaster Lund 1831
G. †Eocenomyrma Dlussky and Radchenko 2006
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†Eocenomyrma breviscapa Radchenko and Dlussky 2016
†Eocenomyrma electrina Dlussky and Radchenko 2006
†Eocenomyrma elegantula Dlussky and Radchenko 2006
†Eocenomyrma groehni Radchenko 2020
†Eocenomyrma orthospina Dlussky and Radchenko 2006
†Eocenomyrma ukrainica Radchenko and Dlussky 2016
G. Gauromyrmex Menozzi 1933
G. †Hypopomyrmex Emery 1891
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†Hypopomyrmex bombiccii Emery 1891
G. Leptothorax Mayr 1855
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Subg. Leptothorax (Macromischa) Roger 1863
Leptothorax muscorum Nylander 1846
G. †Lonchomyrmex Mayr 1867
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†Lonchomyrmex freyeri Heer 1867
†Lonchomyrmex nigritus Assmann 1870
G. Lophomyrmex Emery 1892
G. Lordomyrma Emery 1897
G. Meranoplus Smith 1853
G. Myrmecina Curtis 1829
G. Nesomyrmex Wheeler 1910
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†Nesomyrmex caritatis de Andrade 1999
†Nesomyrmex dominicanus de Andrade 1999
G. †Oxyidris Wilson 1985
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†Oxyidris antillana Wilson 1985
G. †Parameranoplus Wheeler 1915
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†Parameranoplus primaevus Wheeler 1915
G. Podomyrma Smith 1859
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†Podomyrma mayri Emery 1891
Invalid names: Acrostigma Emery 1891 [synonym]
G. Pristomyrmex Mayr 1866
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†Pristomyrmex archaios Radchenko and Dlussky 2018
†Pristomyrmex elmesi Radchenko and Dlussky 2018
†Pristomyrmex rasnitsyni Dlussky and Radchenko 2011
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Invalid names: Pristomyrmex parumpunctatus Emery 1887 [synonym]
G. Proatta Forel 1912
G. †Proleptothorax Radchenko et al. 2018
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†Proleptothorax primitivus Radchenko et al. 2018
G. †Stigmomyrmex Mayr 1868
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†Stigmomyrmex rugulosus Radchenko and Dlussky 2017
†Stigmomyrmex venustus Mayr 1868
G. †Stiphromyrmex Wheeler 1915
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G. Temnothorax Mayr 1861
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†Temnothorax praecreolus de Andrade 1992
G. Tetramorium Mayr 1855
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†Tetramorium kulickae Radchenko and Dlussky 2015
†Tetramorium paraarmatum Radchenko and Dlussky 2015
G. †Thanacomyrmex Chény et al. 2019
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†Thanacomyrmex hoffeinsorum Chény et al. 2019
G. Vollenhovia Mayr 1865
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†Vollenhovia beyrichi Mayr 1868
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Invalid names: Propodomyrma samlandica Wheeler 1910 [synonym]
†Vollenhovia prisca André 1895
Invalid names: Propodomyrma Wheeler 1910 [synonym]
Invalid names: Cataulacini Emery 1895 [synonym], Formicoxenini Forel 1893 [synonym], Meranoplini Emery 1914 [synonym], Myrmecinini Ashmead 1905 [synonym], Tetramoriini Emery 1895 [synonym]
G. †Electromyrmex Wheeler 1910
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†Electromyrmex klebsi Wheeler 1910
†Electromyrmex wheeleri Radchenko and Dlussky 2018
G. †Enneamerus Mayr 1868
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†Enneamerus costatus Radchenko and Dlussky 2017
G. †Eocenidris Wilson 1985
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†Eocenidris crassa Wilson 1985
G. †Eomyrmex Hong 1974
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G. †Eotemnothorax Radchenko 2024
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†Eotemnothorax armatum Radchenko 2024
†Eotemnothorax balticus Radchenko 2024
†Eotemnothorax foveocephalus Radchenko 2024
†Eotemnothorax glaesarius Wheeler 1915
†Eotemnothorax groehni Radchenko 2024
†Eotemnothorax hystriculus Wheeler 1915
†Eotemnothorax khomychi Radchenko 2024
†Eotemnothorax longaevus Wheeler 1915
†Eotemnothorax odontos Radchenko 2024
†Eotemnothorax pedunculatus Radchenko 2024
†Eotemnothorax perkovskii Radchenko 2024
†Eotemnothorax placivus Wheeler 1915
†Eotemnothorax punctatus Radchenko 2024
†Eotemnothorax rav Radchenko 2024
†Eotemnothorax rectispinus Radchenko 2024
†Eotemnothorax rhytidus Radchenko 2024
G. †Fallomyrma Dlussky and Radchenko 2006
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†Fallomyrma anodonta Radchenko and Dlussky 2018
†Fallomyrma marginata Radchenko and Dlussky 2018
†Fallomyrma robusta Radchenko and Dlussky 2018
†Fallomyrma transversa Dlussky and Radchenko 2006
G. †Ilemomyrmex Wilson 1985
+
†Ilemomyrmex caecus Wilson 1985
G. †Incertogaster Boudinot 2024
+
†Incertogaster aurora LaPolla and Greenwalt 2015
†Incertogaster primitiva Radchenko and Dlussky 2019
G. †Lelejus Radchenko and Proshchalykin 2021
+
†Lelejus venustus Radchenko and Proshchalykin 2021
G. Melissotarsus Emery 1877
+
†Melissotarsus ethiopiensis Coty et al. 2016
G. †Miosolenopsis Zhang 1989
+
†Miosolenopsis fossilis Zhang 1989
G. †Myrmecites Dlussky and Rasnitsyn 2002
+
†Myrmecites dubius Dlussky et al. 2015
†Myrmecites latus Dlussky and Putyatina 2014
†Myrmecites major Dlussky et al. 2015
†Myrmecites pusillus Heer 1849
†Myrmecites rotundiceps Dlussky and Rasnitsyn 2002
†Myrmecites sibiricus Dlussky et al. 2015
Tr. Myrmicini Lepeletier de Saint-Fargeau 1835
+
G. Manica Jurine 1807
+
†Manica andrannae Zharkov and Dubovikoff 2022
G. Myrmica Latreille 1804
+
†Myrmica andryushchenkoi Zharkov and Dubovikoff 2025
†Myrmica damzeni Radchenko 2023
†Myrmica dictyosa Radchenko 2023
†Myrmica electrina Radchenko 2023
†Myrmica eocenica Radchenko et al. 2007
Myrmica incompleta Provancher 1881
†Myrmica intermedia Wheeler 1915
Myrmica lobifrons Pergande 1900
†Myrmica longispinosa Mayr 1868
†Myrmica nungesseri Jessen 2020
†Myrmica paradoxa Radchenko et al. 2007
†Myrmica rudis Mayr 1868
†Myrmica saxonica Radchenko 2023
†Myrmica schaefersi Jessen 2020
Invalid names: Myrmica aemula Heer 1849 [nomen dubium], Myrmica angusticollis Heer 1849 [nomen dubium], Myrmica macrocephala Heer 1849 [nomen dubium], Myrmica molassica Heer 1849 [nomen dubium], Myrmica nebulosa Novák 1878 [nomen dubium], Myrmica obsoleta Heer 1849 [nomen dubium], Nothomyrmica Wheeler 1915 [synonym]
G. †Plesiomyrmex Dlussky and Radchenko 2009
+
†Plesiomyrmex tubulatus Dlussky and Radchenko 2009
G. †Protomyrmica Dlussky and Radchenko 2009
+
†Protomyrmica atavia Dlussky and Radchenko 2009
Tr. Pogonomyrmecini Ward et al. 2014
+
G. Pogonomyrmex Mayr 1868
+
†Pogonomyrmex fossilis Carpenter 1930
Tr. Solenopsidini Forel 1893
+
G. Monomorium Mayr 1855
+
†Monomorium kugleri Radchenko and Perkovsky 2009
†Monomorium mayrianum Wheeler 1915
+
Invalid names: Lampromyrmex gracillimus Mayr 1868 [replaced]
†Monomorium pilipes Mayr 1868
Invalid names: Lampromyrmex Mayr 1868 [synonym]
G. Myrmicaria Saunders 1842
G. Rogeria Emery 1894
G. Solenopsis Westwood 1840
+
†Solenopsis alena Özdikmen 2010
+
Invalid names: Solenopsis major Theobald 1937 [replaced]
†Solenopsis atavinus Perfilieva et al. 2017
†Solenopsis blanda Förster 1891
+
Invalid names: Attopsis acuta Förster 1891 [synonym]
†Solenopsis foersteri Theobald 1937
†Solenopsis maxima Förster 1891
†Solenopsis moesta Förster 1891
†Solenopsis privata Förster 1891
†Solenopsis superba Förster 1891
+
Invalid names: Attopsis extensa Förster 1891 [synonym]
†Solenopsis valida Förster 1891
Invalid names: Eurytarsites Hong 2002 [nomen nudum], Monomoriini Wheeler 1915 [synonym]
G. †Solenopsites Dlussky and Rasnitsyn 2002
+
†Solenopsites abdita LaPolla and Greenwalt 2015
†Solenopsites minutus Dlussky and Rasnitsyn 2002
†Solenopsites rossi Dlussky and Perfilieva 2014
Tr. Stenammini Ashmead 1905
+
G. Aphaenogaster Mayr 1853
+
†Aphaenogaster amphioceanica de Andrade 1995
†Aphaenogaster antiqua Dlussky 2002
†Aphaenogaster archaica Meunier 1915
†Aphaenogaster dlusskyana Radchenko and Perkovsky 2016
†Aphaenogaster donisthorpei Carpenter 1930
†Aphaenogaster groehni Radchenko 2024
†Aphaenogaster khomychi Radchenko 2025
†Aphaenogaster mayri Carpenter 1930
†Aphaenogaster mersa Wheeler 1915
†Aphaenogaster oligocenica Wheeler 1915
†Aphaenogaster praerelicta de Andrade 1995
†Aphaenogaster ribbeckei Radchenko 2024
†Aphaenogaster wennebergi Zharkov and Dubovikoff 2025
Invalid names: Deromyrma Forel 1913 [synonym], Rectorhagas Lin 1982 [synonym], Sinaphaenogaster Zhang 1989 [synonym]
G. Goniomma Emery 1895
+
†Goniomma poschmanni Jessen 2020
G. Messor Forel 1890
+
†Messor sculpturatus Carpenter 1930
G. †Paraphaenogaster Dlussky 1981
+
†Paraphaenogaster avita Fujiyama 1970
†Paraphaenogaster bizeri Jessen 2020
†Paraphaenogaster hooleyana Dlussky and Perfilieva 2014
+
Invalid names: Myrmica concinna Heer 1867 [synonym]
†Paraphaenogaster loosi Jessen 2020
†Paraphaenogaster maculata Theobald 1937
†Paraphaenogaster maculipes Theobald 1937
†Paraphaenogaster pannonica Bachmayer 1960
+
Invalid names: Myrmica bicolor Heer 1867 [synonym]
†Paraphaenogaster ussuriensis Perfilieva 2022
†Paraphaenogaster wuttkei Jessen 2020
G. Stenamma Westwood 1840
+
†Stenamma berendti Mayr 1868
G. Veromessor Forel 1917
+
G. †Zhangidris Bolton 2003
+
†Zhangidris atopogaster Zhang 1989
Invalid names: Heteromyrmex Zhang 1989 [replaced]
Invalid names: Aphaenogaster longaeva Scudder 1877 [nomen dubium], Brachytarsites Hong 2002 [nomen nudum], Clavipetiola Hong 2002 [nomen nudum], Fushunomyrmex Hong 2002 [nomen nudum], Orbigastrula Hong 2002 [nomen nudum], Pseudomyrmini Emery 1914 [empty], Quadrulicapito Hong 2002 [nomen nudum], Quineangulicapito Hong 2002 [nomen nudum], Sinomyrmex Hong 2002 [nomen nudum], Solenopsidiini Forel 1893 [empty], Sphaerogasterites Hong 2002 [nomen nudum], Wumyrmex Hong 2002 [nomen nudum]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Early/Lower Cenomanian to the top of the Holocene or 100.50000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 93.9 Ma

Collections (251 total)


Time interval Ma Country or state Original ID and collection number
Early/Lower Cenomanian100.5 - 93.9Myanmar Antiquiformica alata (165561)
Turonian93.9 - 89.8Botswana Afromyrma petrosa (123430)
Turonian93.9 - 89.8USA (New Jersey) Kyromyrma neffi (110895)
Thanetian59.24 - 56.0USA (Alaska) Formicinae indet. (196713)
Wasatchian56.0 - 51.1USA (Wyoming) Titanomyrma sp. (229109)
Ypresian56.0 - 48.07USA (Wyoming) Titanomyrma lubei (108779)
Ypresian56.0 - 48.07Canada (British Columbia) Formicidae indet. A and B (31677) Leptothorax sp., Technomyrmex sp. (123795) Oecophyllini indet. A (192140) Titanomyrma sp. (226231)
Ypresian56.0 - 48.07China (Liaoning) Eomyrmex guchengziensis (125469) Sinomyrmex brunnea, Fushunomyrmex liaoningensis, Orbigastrula guchengziensis, Quineangulicapito fushunensis, Brachytarsites dongzhouheensis, Quadrulicapito longa, Wumyrmex furvis, Eoleptocerites chinensis, Eoleptocerites fushunensis, Leptogasteritus capricornutes, Sinotenuicapito badis, Magnogasterites curticornutus, Ovalicapito fushunensis, Ovaligastrula xilutianensis, Orbicapitia reticulata, Longiformica dongzhouheensis, Longiformica guchengziensis, Wilsonia megagastrosa, Wilsonia liaoningensis, Fushuniformica spinata, Sinoformica longicapitata (124820) Sinomyrmex furvis, Sphaerogasterites longipetiolatus, Clavipetiola asiatica, Huaxiaformica mallepetiolata, Liaoformica longipetiolata, Eurytarsites fushunensis, Curtipalpulus eocenicus (124530)
Ypresian56.0 - 48.07France (Oise) Gesomyrmex gallicus (123850)
Ypresian56.0 - 48.07USA (Washington) Camponotites kraussei (124086)
Bridgerian51.1 - 45.9USA (Colorado) Myrmecites rotundiceps, Solenopsites minutus (138664)
Bridgerian51.1 - 45.9USA (Utah) Camponotus vetus (121147)
Bridgerian51.1 - 45.9USA (Wyoming) Myrmica sp. (121306)
Lutetian48.07 - 41.03Russian Federation Chimaeromyrma brachycephala, Aphaenogaster dlusskyana (107340)
Lutetian48.07 - 41.03China (Liaoning) Camponotus tokunagai (162504)
Lutetian48.07 - 41.03USA (Montana) Camponotites spp., Formica annosa, Lasius glom, Crematogaster aurora, Solenopsites abdita (117614)
Lutetian48.07 - 41.03USA (Mississippi) Oecophylla sp. (157347)
Lutetian48.07 - 41.03United Kingdom (England) Oecophylla bartoniana, Formica heteroptera (127644)
MP 1148.07 - 41.03Germany (Hessen) Formicium giganteum, Formicium simillimum (120036 138656) Myrmicinae indet., Camponotus sp., Formicium giganteum, Formicium simillimum, Oecophylla longiceps, Formicinae indet., Pseudectatomma striatula, Gesomyrmex pulcher, Gesomyrmex breviceps, Gesomyrmex curiosus, Ponerinae indet. 1 (120798)
MP 1348.07 - 41.03Germany (Eifel) Oecophylla eckfeldiana (135644) Oecophylla eckfeldiana, Gesomyrmex flavescens (138912) Oecophylla eckfeldiana, Gesomyrmex germanicus (135640 138911)
Uintan45.9 - 39.7USA (Colorado) Formicinae indet. (138682)
Bartonian41.03 - 37.71India (Gujarat) Formica sp., Gesomyrmex sp. (228739)
Bartonian41.03 - 37.71USA (Arkansas) Eocenidris crassa, Formicidae indet. (122391)
Bartonian41.03 - 37.71USA (Tennessee) Eoponera berryi (138891)
Late/Upper Eocene37.71 - 33.9USA (Colorado) Aphaenogaster mayri, Messor sculpturatus, Formica robusta, Lasius peritulus (122093)
Late/Upper Eocene37.71 - 33.9Czechia Oecophylla obesa (138883)
Late/Upper Eocene37.71 - 33.9USA (Texas) Formica eoptera (130390)
Priabonian37.71 - 33.9United Kingdom (Isle of Wight) Oecophylla megarche, Leucotaphus gurnetensis, Camponotus cockerelli (157669)
Priabonian37.71 - 33.9Denmark Aphaenogaster wennebergi (242225) Gesomyrmex hoernesi, Myrmica eocenica, Plagiolepis klinsmanni, Prenolepis henschei, Formica gustawi, Gnamptogenys europaea, Eocenomyrma electrina, Fallomyrma transversa, Nylanderia pygmaea, Plagiolepis kuenowi, Pristomyrmex rasnitsyni, Eotemnothorax armatum (123762)
Priabonian37.71 - 33.9France Formica sp. (113709)
Priabonian37.71 - 33.9Russian Federation Gesomyrmex hoernesi, Formica flori, Ectatomma europaeum, Nothomyrmica rudis, Plagiolepis kunowi, Lasius schiefferdeckeri, Formica constricta (123930) Thanacomyrmex hoffeinsorum (123419)
Priabonian37.71 - 33.9Russian Federation (Primorskiy) Formica sp., Lasius sp. A, Lasius sp. B, Formica biamoensis, Formica paleosibirica, Myrmecites sibiricus, Myrmecites dubius, Myrmecites major (172678) Formica sp., Lasius sp. B, Formica paleosibirica, Gesomyrmex magnus, Biamomyrma zherikhini (123794) Myrmicinae indet., Formica sp., Lasius sp. A, Lasius sp. B, Formica paleosibirica, Gesomyrmex macrops, Gesomyrmex incertus, Agastomyrma laticeps, Biamomyrma lata, Biamomyrma rugosa (172679)
Priabonian37.71 - 33.9Lithuania Camponotus mengei, Prenolepis henschei, Lasius schiefferdeckeri, Formica antiqua, Formica baltica, Formica parvula, Formica gustawi, Formica proformicoides, Gnamptogenys rohdendorfi (113470) Oecophylla crassinoda, Myrmica andryushchenkoi (232023)
Priabonian37.71 - 33.9Canada (British Columbia) Aphaenogaster longaeva, Formica arcana (121480)
Priabonian37.71 - 33.9Germany (Halle) Eotemnothorax odontos (149487) Gesomyrmex hoernesi, Aphaenogaster sommerfeldti, Plagiolepis klinsmanni, Plagiolepis squamifera, Plagiolepis solitaria, Prenolepis henschei, Lasius schiefferdeckeri, Formica phaethusa, Pseudolasius boreus, Conoformica bitterfeldiana, Formica gustawi, Formica kutscheri, Gnamptogenys europaea, Eocenomyrma rugosostriata, Fallomyrma transversa, Nylanderia pygmaea, Plagiolepis kuenowi, Stigmomyrmex rugulosus, Electromyrmex wheeleri, Myrmica saxonica, Eotemnothorax gracilis (129227) Gesomyrmex hoernesi, Formica flori, Monomorium mayrianum, Stigmomyrmex venustus, Plagiolepis solitaria, Prenolepis henschei, Lasius schiefferdeckeri, Formica gustawi, Formica kutscheri, Gnamptogenys europaea, Fallomyrma transversa, Nylanderia pygmaea, Plagiolepis kuenowi, Plagiolepis paradoxa, Plesiomyrmex tubulatus, Pristomyrmex archaios, Carebara kutscheri, Eotemnothorax gracilis, Eotemnothorax longaevus, Eotemnothorax rectispinus, Eotemnothorax pedunculatus (129228) Myrmica paradoxa, Eocenomyrma rugosostriata (123910)
Priabonian37.71 - 33.9Russian Federation (Kaliningrad) Aphaenogaster sommerfeldti, Eotemnothorax rectispinus (229453) Camponotus mengei, Camponotus constrictus (138655) Eotemnothorax petiolatus (234648) Eotemnothorax rectispinus, Eotemnothorax foveocephalus, Eotemnothorax punctatus (147612) Formica flori, Camponotus mengei, Aphaenogaster sommerfeldti, Oecophylla brischkei, Prenolepis henschei, Lasius schiefferdeckeri (138654) Formica flori, Lasius schiefferdeckeri (138658) Formica flori, Macromischa rugosostriata (123962) Formica flori, Prenolepis henschei (138660) Formica flori, Prenolepis henschei, Prenolepis pygmaea, Lasius schiefferdeckeri (138661) Gesomyrmex hoernesi (154709) Gesomyrmex hoernesi, Formica flori, Camponotus mengei, Monomorium mayrianum, Erebomyrma antiqua, Aphaenogaster sommerfeldti, Electromyrmex klebsi, Nothomyrmica rudis, Leptothorax gracilis, Parameranoplus primaevus, Enneamerus reticulatus, Plagiolepis klinsmanni, Plagiolepis kunowi, Plagiolepis squamifera, Dimorphomyrmex theryi, Prodimorphomyrmex primigenius, Oecophylla brischkei, Prenolepis henschei, Prenolepis pygmaea, Lasius schiefferdeckeri, Lasius pumilus, Lasius punctulatus, Formica phaethusa, Formica constricta, Pseudolasius boreus, Drymomyrmex claripennis, Formica gustawi, Propodomyrma samlandica (124073) Gesomyrmex hoernesi, Formica flori, Camponotus mengei, Plagiolepis klinsmanni, Dimorphomyrmex theryi, Oecophylla brischkei, Prenolepis henschei, Lasius schiefferdeckeri, Formica constricta, Macromischa beyrichi, Formica palaeopolonica (123757) Gesomyrmex hoernesi, Formica flori, Dimorphomyrmex theryi, Prenolepis henschei, Prenolepis pygmaea, Lasius schiefferdeckeri, Pseudolasius boreus (124205) Gesomyrmex hoernesi, Formica flori, Glaphyromyrmex oligocenicus, Drymomyrmex fuscipennis, Camponotus mengei, Myrmica longispinosa, Macromischa rudis, Nothomyrmica intermedia, Ectatomma europaeum, Electroponera dubia, Monomorium pilipes, Stenamma berendti, Aphaenogaster sommerfeldti, Aphaenogaster oligocenica, Aphaenogaster mersa, Leptothorax gracilis, Leptothorax glaesarius, Leptothorax longaevus, Leptothorax hystriculus, Leptothorax placivus, Stigmomyrmex venustus, Enneamerus reticulatus, Plagiolepis klinsmanni, Plagiolepis squamifera, Rhopalomyrmex pygmaeus, Dimorphomyrmex theryi, Dimorphomyrmex mayri, Gesomyrmex annectens, Oecophylla brischkei, Oecophylla brevinodis, Prenolepis henschei, Prenolepis pygmaea, Lasius schiefferdeckeri, Lasius pumilus, Lasius punctulatus, Lasius nemorivagus, Formica horrida, Formica clymene, Formica strangulata, Pseudolasius boreus, Camponotus constrictus, Macromischa beyrichi, Macromischa rugosostriata, Macromischa petiolata, Pheidologeton antiquus, Lampromyrmex gracillimus, Stigmomyrmex robustus, Plagiolepis kuenowi (123927) Gesomyrmex hoernesi, Formica flori, Macromischa rudis, Aphaenogaster sommerfeldti, Prenolepis henschei, Lasius schiefferdeckeri (138663) Gesomyrmex hoernesi, Formica flori, Myrmica eocenica, Oecophylla brischkei, Prenolepis henschei, Formica phaethusa, Formica gustawi, Formica zherikhini, Plagiolepis minutissima, Nylanderia pygmaea, Plagiolepis kuenowi, Bilobomyrma baltica, Eotemnothorax hystriculus (122591) Gesomyrmex hoernesi, Formica flori, Prenolepis henschei, Lasius schiefferdeckeri (138657) Gesomyrmex hoernesi, Formica flori, Prenolepis henschei, Lasius schiefferdeckeri, Formica gustawi, Sussudio boreus (123458) Gesomyrmex hoernesi, Myrmica intermedia, Aphaenogaster sommerfeldti, Stigmomyrmex venustus, Plagiolepis solitaria, Prenolepis henschei, Lasius schiefferdeckeri, Carebara antiqua, Cataglyphoides intermedius, Gnamptogenys europaea, Eocenomyrma rugosostriata, Eocenomyrma elegantula, Nylanderia pygmaea, Plagiolepis kuenowi, Enneamerus costatus, Electromyrmex wheeleri, Carebara groehni, Eotemnothorax gracilis, Eotemnothorax hystriculus, Eotemnothorax groehni, Eotemnothorax balticus, Eotemnothorax rhytidus, Aphaenogaster groehni, Aphaenogaster ribbeckei, Eocenomyrma orthospina (110735) Lasius sp. (124318) Lasius sp., Prenolepis henschei (125139) Macromischa prisca (138940) Manica andrannae (192258) Metalasius pumilus (123928) Myrmica longispinosa, Myrmica eocenica, Aphaenogaster sommerfeldti, Myrmica electrina, Eotemnothorax gracilis, Eotemnothorax rectispinus (123764) Myrmica longispinosa, Myrmica rudis, Myrmica intermedia, Aphaenogaster mersa, Myrmica damzeni (123964) Nylanderia pygmaea, Eotemnothorax balticus (124003) Plagiolepis klinsmanni, Dimorphomyrmex theryi, Camponotus igneus, Nylanderia pygmaea, Protomyrmica atavia (123766) Plagiolepis minutissima, Eotemnothorax balticus, Rhopalomyrmex pygmaeus (123420)
Priabonian37.71 - 33.9Ukraine Aphaenogaster sommerfeldti, Eotemnothorax gracilis, Eotemnothorax glaesarius, Eotemnothorax hystriculus, Eotemnothorax armatum, Eotemnothorax rhytidus, Eotemnothorax punctatus, Eotemnothorax khomychi, Aphaenogaster khomychi (238338) Camponotus mengei, Myrmica rudis, Fallomyrma transversa, Formica ribbeckei (81780) Drymomyrmex rasnitsyni (207958) Eocenomyrma rugosostriata, Eotemnothorax balticus (196704) Eotemnothorax rectispinus (154710 185682) Formica flori, Prenolepis henschei, Lasius schiefferdeckeri (122998) Gesomyrmex hoernesi, Monomorium kugleri, Enneamerus reticulatus, Plagiolepis klinsmanni, Plagiolepis solitaria, Prenolepis henschei, Lasius schiefferdeckeri, Pseudolasius boreus, Formica gustawi, Formica radchenkoi, Plagiolepis minutissima, Eocenomyrma orthospina, Fallomyrma transversa, Nylanderia pygmaea, Plagiolepis kuenowi, Bilobomyrma ukrainica, Boltonidris mirabilis, Eocenomyrma ukrainica, Eocenomyrma breviscapa, Fallomyrma anodonta, Fallomyrma marginata, Fallomyrma robusta, Pristomyrmex elmesi, Proleptothorax primitivus, Crematogaster primitiva, Formica ribbeckei, Myrmica electrina, Eotemnothorax gracilis, Eotemnothorax hystriculus, Eotemnothorax placivus, Eotemnothorax armatum, Eotemnothorax punctatus, Eotemnothorax pedunculatus, Eotemnothorax perkovskii (122997) Lasius schiefferdeckeri (153882 234940) Myrmicinae indet., Formica flori, Camponotus mengei, Plagiolepis klinsmanni, Prenolepis henschei, Lasius schiefferdeckeri, Pseudolasius boreus, Formica gustawi, Plagiolepis minutissima, Aphaenogaster antiqua, Oligomyrmex nitidus, Oligomyrmex ucrainicus, Gnamptogenys europaea, Fallomyrma transversa, Lelejus venustus, Eotemnothorax rav (113621) Nylanderia pygmaea (122996) Plagiolepis klinsmanni, Formica ribbeckei (170205) Prenolepis henschei (124272)
Priabonian37.71 - 33.9United Kingdom (England) Formicium brodiei, Leptothorax gurnetensis, Oecophylla atavina, Oecophylla perdita, Oecophylla megarche, Leucotaphus cockerelli, Leucotaphus donisthorpei, Solenopsites rossi (123959) Oecophylla megarche, Leucotaphus gurnetensis, Camponotus cockerelli (21976) Oecophylla megarche, Leucotaphus gurnetensis, Camponotus cockerelli, Leucotaphus donisthorpei, Paraphaenogaster hooleyana (123960)
Priabonian37.71 - 33.9Poland Gesomyrmex hoernesi, Formica flori, Camponotus mengei, Aphaenogaster sommerfeldti, Enneamerus reticulatus, Plagiolepis klinsmanni, Oecophylla brischkei, Prenolepis henschei, Prenolepis pygmaea, Lasius schiefferdeckeri, Lasius pumilus, Camponotus constrictus, Aphaenogaster berendti, Macromischa petiolata, Lampromyrmex gracillimus, Stigmomyrmex robustus, Formica surinamensis (109625) Gesomyrmex hoernesi, Formica flori, Camponotus mengei, Ectatomma europaeum, Monomorium pilipes, Stigmomyrmex venustus, Enneamerus reticulatus, Plagiolepis klinsmanni, Plagiolepis singularis, Plagiolepis squamifera, Plagiolepis solitaria, Oecophylla brischkei, Prenolepis henschei, Prenolepis pygmaea, Lasius schiefferdeckeri, Lasius pumilus, Lasius punctulatus, Lasius edentatus, Camponotus igneus, Camponotus constrictus, Pheidologeton antiquus, Lampromyrmex gracillimus (120811) Monomorium sp., Gesomyrmex hoernesi, Formica flori, Camponotus mengei, Myrmica rudis, Stiphromyrmex robustus, Enneamerus reticulatus, Plagiolepis klinsmanni, Prenolepis henschei, Lasius schiefferdeckeri, Gnamptogenys europaea, Eocenomyrma rugosostriata, Eocenomyrma orthospina, Nylanderia pygmaea, Plagiolepis kuenowi, Tetramorium paraarmatum, Tetramorium kulickae (120813) Stigmomyrmex rugulosus, Myrmica dictyosa (110737)
Priabonian37.71 - 33.9Belarus Lasius schiefferdeckeri (224403)
Late/Upper Eocene - Early/Lower Oligocene37.71 - 27.3USA (Nevada) Pseudocamponotus elkoanus (138653)
Chadronian37.0 - 33.9USA (Colorado) Aphaenogaster mayri, Aphaenogaster donisthorpei, Messor sculpturatus, Pheidole tertiaria, Pogonomyrmex fossilis, Cephalomyrmex rotundatus, Formica robusta, Formica cockerelli, Formica grandis, Lasius peritulus, Camponotus fuscipennis, Camponotus microcephalus, Camponotus petrifactus (106384) Aphaenogaster mayri, Formica robusta (113749) Aphaenogaster mayri, Formica robusta, Lasius peritulus, Camponotus petrifactus (123119) Aphaenogaster mayri, Messor sculpturatus, Formica robusta, Formica cockerelli, Formica grandis, Lasius peritulus (124624) Aphaenogaster mayri, Messor sculpturatus, Formica robusta, Formica cockerelli, Lasius peritulus, Camponotus fuscipennis, Camponotus petrifactus (124002) Aphaenogaster mayri, Messor sculpturatus, Lasius peritulus (132362) Aphaenogaster mayri, Messor sculpturatus, Pogonomyrmex fossilis, Formica robusta, Formica cockerelli, Lasius peritulus, Camponotus fuscipennis, Camponotus petrifactus (127617) Aphaenogaster mayri, Tetramorium peritulum (124680)
Early/Lower Oligocene33.9 - 27.3France Attopsis nigra, Camponotus vehemens, Attopsis extensa, Attopsis moesta, Attopsis maxima, Attopsis valida, Attopsis longipennis, Attopsis longipes (121476) Attopsis superba, Attopsis valida (121474) Camponotus miserabilis, Camponotus compactus, Oecophylla praeclara, Attopsis privata, Attopsis blanda, Attopsis acuta, Attopsis longipennis (121473) Solenopsis superba, Solenopsis foersteri, Solenopsis blanda, Aphaenogaster maculipes (122408)
Rupelian33.9 - 27.3Germany Myrmicidae indet., Camponotus vehemens, Solenopsis maxima, Solenopsis valida, Solenopsis major, Solenopsis superba, Solenopsis foersteri, Solenopsis blanda, Aphaenogaster maculipes, Oecophylla superba, Formica flori, Formica tripartita, Formica alsatica, Formica sepulta, Glaphyromyrmex oligocenicus, Drymomyrmex fuscipennis, Camponotus mengei (122407)
Rupelian33.9 - 27.3Czechia Camponotus novotnyi (138762)
Chattian27.3 - 23.04France (Bouches-Du-Rhone) Erebomyrma thorali (138838) Formica capito (122876) Formica minutula (122877) Formica sp. (113712) Formica sp., Formica minutula, Formica sepulta, Aphaenogaster maculata, Camponotus longiventris, Camponotus saussurei, Lasius epicentrus, Formica latinodosa (122501)
Chattian27.3 - 23.04Russian Federation Camponotus parvus, Lasius vulgaris, Paraphaenogaster ussuriensis (125971)
Chattian27.3 - 23.04France (Provence-Alpes-Côte d'Azur) Aphaenogaster maculata (122487) Formica sepulta, Camponotus penninervis, Formica serresi (122488)
MP 2827.3 - 23.04Germany (Rheinland-Pfalz) Myrmicinae indet., Paraphaenogaster sp., Paraphaenogaster loosi, Paraphaenogaster schindleri, Paraphaenogaster wettlauferi, Paraphaenogaster wuttkei, Goniomma poschmanni, Myrmica schaefersi, Myrmica nungesseri (160097) Paraphaenogaster schindleri (123354) Paraphaenogaster sp., Paraphaenogaster loosi, Paraphaenogaster schindleri, Paraphaenogaster bizeri, Paraphaenogaster freihauti, Aphaenogaster enspelensis, Goniomma poschmanni (211302) Paraphaenogaster sp., Stenammini indet., Paraphaenogaster loosi (211301) Stenammini indet. (215791)
MP 3027.3 - 23.04Germany (Nordrhein-Westfalen) Formica sp. (127628) Formica sp., Myrmica sp., Myrmica archaica, Formica bauckhorni, Andrena tertiaria (138696)
MP 3027.3 - 23.04Germany Formica lignitum (115802)
Aquitanian23.04 - 20.45Germany Myrmicinae indet., Formicinae indet., Aphaenogaster sp., Lasius sp. (123872)
Aquitanian23.04 - 20.45New Zealand Rhytidoponera waipiata, Rhytidoponera gibsoni, Myrmecorhynchus novaeseelandiae (50035)
Early/Lower Miocene23.04 - 15.98Mexico Camponotus sp., Pheidole sp. 1, Pheidole sp. 2, Solenopsis sp., Crematogaster sp., Cephalotes sp. 1, Cephalotes sp. 2, Cephalotes ventriosus (21977)
Early/Lower Miocene23.04 - 15.98Ethiopia Melissotarsus ethiopiensis (183971)
Early/Lower Miocene23.04 - 15.98Mexico (Chiapas) Aphaenogaster praerelicta, Cyphomyrmex maya, Cephalotes poinari, Cephalotes maya, Cephalotes olmecus (125757) Camponotus sp. (153340) Pheidole chaan (217216) Pheidole pauchil (198592)
Burdigalian20.45 - 15.98Japan (Gifu) Camponotus sp., Formica sp. A, Formica sp. B, Solenopsis sp. (181074)
Burdigalian20.45 - 15.98China (Shandong) Camponotus shanwangense (128549) Formica flavifemoralis, Camponotus pictus (138944) Formica linquensis, Formica ovala, Camponotus shanwangensis, Camponotus gracilis, Camponotus ampullosus, Camponotus festinus, Camponotus microthoracus, Camponotus ambon (128552) Formica sp. 1, Rectorhagas dumetorum (130412) Miosolenopsis fossilis, Tylolasius inflatus, Formica sp. 2, Oecophylla xiejiaheensis, Aphaenogaster lapidescens, Aphaenogaster paludosa, Formica ceps, Formica ovala, Formica fusca, Lasius mordicus, Camponotus shanwangensis, Camponotus mitis, Camponotus plenus, Camponotus luteus, Camponotus ambon (128551) Rabidia xiejiaheensis, Camponotus shanwangense, Heteromyrmex atopogaster, Paraphaenogaster shanwangensis, Lasius vetulus, Formica linquensis, Lasius validus, Lasius truncatus, Camponotus curviansatus, Camponotus mitis, Camponotus longus, Camponotus microthoracus (128550)
Burdigalian20.45 - 15.98Austria Myrmicinae indet., Camponotus sp. (139942)
Burdigalian20.45 - 15.98Kenya Oecophylla leakeyi (138938)
Burdigalian - Langhian20.45 - 13.82Dominican Republic Acropyga glaesaria (124429) Acropyga sp. (124596) Acropyga sp., Acropyga glaesaria, Apterostigma electropilosum, Apterostigma eowilsoni, Gnamptogenys brunoi, Gnamptogenys casca, Nylanderia vetula, Zatania electra (122982) Basiceros enana (152045) Cephalotes integerrimus (124282 138708) Cephalotes serratus, Cephalotes jansei, Cephalotes squamosus, Acanthognathus poinari, Cephalotes brevispineus, Cephalotes bloosi, Cephalotes caribicus, Cephalotes sucinus, Leptothorax dominicanus, Strumigenys pilosula, Strumigenys poinari (122472) Cephalotes serratus, Cephalotes obscurus, Cephalotes squamosus, Leptothorax praecreolus (124281) Exocryptocerus jansei, Zacryptocerus squamosus, Zacryptocerus pseudoaztecus, Zacryptocerus obscurus, Cephalotes integerrimus, Cyphomyrmex taino, Ilemomyrmex caecus, Zatania electra (21978) Exocryptocerus serratus, Cephalotes squamosus, Pheidole tethepa, Leptothorax caritatis (138700) Exocryptocerus serratus, Exocryptocerus elevatus, Exocryptocerus truncatus, Exocryptocerus jansei, Zacryptocerus integerrimus, Zacryptocerus squamosus, Zacryptocerus pseudoaztecus, Zacryptocerus alveolatus (124846) Gnamptogenys sp., Nesomyrmex sp. (222615) Leptothorax caritatis (138937) Oxyidris antillana (138698) Pheidole anticua (203120) Pheidole sp., Cephalotes serratus, Cephalotes alveolatus, Cephalotes obscurus, Cephalotes integerrimus, Cephalotes squamosus, Strumigenys schleeorum, Strumigenys electrina, Aphaenogaster amphioceanica, Cephalotes hispaniolicus, Cephalotes caribicus, Cephalotes dieteri, Gnamptogenys pristina, Gnamptogenys levinates, Pheidole primigenia, Trachymyrmex primaevus (122474)
Hemingfordian18.5 - 16.3USA (California) Palaeosminthurus juliae (138761) Palaeosminthurus sp. (106431)
MN 416.9 - 15.98Czechia Pheidologeton bohemicus, Myrmica nebulosa, Formica buphthalma (117612)
Langhian15.98 - 13.82Ukraine Oecophylla taurica (188087)
Langhian15.98 - 13.82Japan Aphaenogaster avita (103101)
Langhian15.98 - 13.82USA (Idaho) Camponotus sp., Lasius sp., Messor sp. (97616)
Langhian15.98 - 13.82Malaysia (Sarawak) Cataulacus plebeius (138840)
Langhian15.98 - 13.82China (Fujian) Camponotus sp., Aphaenogaster sp., Gnamptogenys sp., Monomorium sp., Pheidole sp., Prenolepis sp., Vollenhovia sp., Cataulacus sp., Crematogaster sp., Polyrhachis sp., Tetramorium sp., Carebara sp., Nylanderia sp., Strumigenys sp., Colobopsis sp., Paratrechina sp., Euprenolepis sp., Cardiocondyla sp., Gauromyrmex sp., Lophomyrmex sp., Lordomyrma sp., Meranoplus sp., Myrmicaria sp., Proatta sp. (220273)
Middle Miocene15.98 - 11.63Russian Federation Camponotites sp., Camponotites macropterus (125988) Lasius vetulus (125990) Oecophylla macroptera (125996) Paraphaenogaster microphthalmus (138756)
Middle Miocene - Late/Upper Miocene15.98 - 5.333Kyrgyzstan Camponotus sp. (138758) Camponotus sp., Rhytidoponera kirghizorum (138754)
Badenian13.82 - 12.8Austria (Steiermark) Myrmica obsoleta, Myrmica aemula (115306)
Sarmatian12.8 - 11.63Croatia Camponotus sp., Oecophylla obesa, Camponotus induratus, Formica ungeri, Lasius globularis, Lasius longaevus, Lasius occultatus, Lasius longipennis, Formica fragilis, Lasius ophthalmicus, Heeridris croaticus, Lasius anthracinus, Paraphaenogaster jurinei, Myrmecites pusillus, Myrmecites latus (117247) Formica lignitum, Formica heraclea (117250) Formica obesa, Formica indurata, Formica obscura, Formica ophthalmica, Formica ungeri, Formica redtenbacheri, Formica globularis, Formica longaeva, Formica occultata, Formica longipennis, Formica minutula, Formica pumila, Formica acuminata, Formica obliterata, Ponera elongatula, Attopsis longipennis, Attopsis anthracina, Myrmica tertiaria, Myrmica bremii, Myrmica jurinei, Myrmica pusilla, Attopsis nigra, Attopsis longipes, Formica fragilis, Formica freyeri, Formica oblita, Formica capito, Formica kollari, Formica aemula, Poneropsis anthracina, Myrmica bicolor, Myrmica concinna, Formica pinguicula, Formica macrocephala (117246) Formica occultata (122874) Lasius occultatus (156759) Lasius ophthalmicus (113693) Lasius ophthalmicus, Solenopsis longaevus (156758) Oecophylla obesa, Camponotus induratus, Formica ungeri, Lasius globularis, Lasius longaevus, Lasius occultatus, Lasius longipennis, Formica fragilis, Lasius ophthalmicus, Formica parexsecta, Lasius anthracinus, Paraphaenogaster tertiaria (113698)
Sarmatian12.8 - 11.63Georgia Lasius martynovi (138759)
Sarmatian12.8 - 11.63Romania Formica sp., Cataulacus sp. (156760)
Sarmatian12.8 - 11.63Germany (Baden-Württemberg) Formica globiventris (117339) Formica lignitum, Formica heraclea, Formica orbata, Imhoffia pallida (117252) Formica primitiva, Formica demersa, Myrmica tertiaria (122866) Formica primordialis (117338) Formica primordialis, Formica immersa, Formica lavateri, Formica seuberti, Formica primitiva, Myrmica macrocephala, Myrmica rugiceps, Myrmica molassica (113695) Formica procera, Formica lignitum, Formica gravida, Formica heraclea, Formica demersa, Formica pulchella, Imhoffia nigra, Cephites fragilis, Formica pinguicula (113689) Formica procera, Formica lignitum, Formica gravida, Formica macrophthalma, Formica demersa, Formica orbata, Imhoffia nigra, Myrmica angusticollis, Formica pinguicula (115304) Formica procera, Formica lignitum, Formica longicollis, Formica immersa, Formica macrophthalma, Formica seuberti, Formica orbata, Formica pulchella, Myrmica macrocephala, Cephites oeningensis (115305) Formica pulchella, Myrmica tertiaria (122867) Myrmica rugiceps (122868)
Sarmatian12.8 - 11.63Ukraine Lasius tertiarius (138935) Paraphaenogaster microphthalmus, Solenopsis atavinus (152283)
Tortonian11.63 - 7.246France Camponotus obesus, Formica cantalica (138763) Lasius crispus (138766)
Late/Upper Miocene11.63 - 5.333Poland Lasius oblongus, Lonchomyrmex nigritus, Myrmica ruficeps (138936)
Late/Upper Miocene11.63 - 5.333Italy Hypopomyrmex bombiccii, Gesomyrmex corniger, Podomyrma mayri, Ectatomma gracile, Cataulacus silvestrii, Cataulacus planiceps, Aeromyrma sophiae, Crematogaster praecursor, Plagiolepis labilis, Oecophylla sicula (123432)
Turolian8.725 - 5.09France (Ardèche) Camponotus crozei, Oecophylla grandimandibula (156164) Formica sp., Solenopsis sp., Formica auxillacensis, Lasius crispus, Lasius chambonensis, Formica maculipennis, Pheidole pallidula (172761)
Messinian7.246 - 5.333Greece (Crete) Polyrhachis annosus (133428)
Messinian7.246 - 5.333France Formica maculipennis (138772) Formica sp., Formica auxillacensis (138771)
Messinian7.246 - 5.333USA (Alaska) Camponotus sp., Leptothorax sp. (123352) Myrmicinae indet., Aphaenogaster sp., Messor sp. (103102)
Messinian7.246 - 5.333Italy (Tuscany) Camponotinae indet. (113751)
Messinian7.246 - 5.333Austria Aphaenogaster pannonicus, Camponotus ullrichi (127417)
Blancan4.7 - 1.4USA (Texas) Myrmicinae indet., Formicinae indet. (154909)
Piacenzian3.6 - 2.58Germany Camponotus sp. (164952)
Piacenzian3.6 - 2.58Germany (Niedersachsen) Camponotites steinbachi (155691) Myrmicinae indet., Camponotus sp., Camponotites silvestris, Camponotini indet., Formicinae indet. (109196)
Gelasian2.58 - 1.8France Erebomyrma antiqua, Formica pitoni (138769) Lasius sp., Lasius chambonensis (113996)
Gelasian2.58 - 1.8Greenland Camponotus herculeanus (160159 160163) Camponotus spp., Formica sp., Myrmica sp., Formicinae indet., Camponotus herculeanus (160109) Camponotus spp., Myrmica sp., Camponotus herculeanus (160144) Formicinae indet., Camponotus herculeanus (160145) Leptothorax sp. (160153)
Calabrian1.8 - 0.774Portugal Lasius sp. (201818)
Chibanian0.774 - 0.129Japan Camponotus sp. (194482) Camponotus sp., Lasius sp. (211426)
Rancholabrean0.21 - 0.014USA (California) Veromessor andrei (152007 161529 161535)
Late/Upper Pleistocene0.129 - 0.0117USA (Wisconsin) Formica sp., Myrmica sp. (169406)
Late/Upper Pleistocene0.129 - 0.0117USA (Colorado) Formica neorufibarbis (169333 169347) Formica neorufibarbis, Myrmica lobifrons (169336) Myrmica lobifrons (169350 169353)
Late/Upper Pleistocene0.129 - 0.0117Canada (British Columbia) Myrmica sp. 1, Leptothorax muscorum, Myrmica incompleta (167673)
Holocene0.0117 - 0.0Indonesia Rhytidoponera araneoides, Pristomyrmex parumpunctatus, Cataulacus resinosus, Oecophylla smaragdina, Calomyrmex levissimus, Camponotus vitreus, Polyrhachis excitata, Polyrhachis restituta, Polyrhachis cincta, Polyrhachis continua, Polyrhachis circumdata, Polyrhachis inclusa, Polyrhachis paromalus, Polyrhachis tibialis, Polyrhachis subfossa, Polyrhachis erosispina, Pheidole plagiaria, Polyrhachis levior, Polyrhachis celebensis (182105)
Holocene0.0117 - 0.0Tanzania Aphaenogaster sp. (152160) Camponotus sp., Crematogaster sp., Pheidole cordata, Lepisiota sp. (234540) Formica cordata (147705) Pheidole rasnitsyni (124004)
Holocene0.0117 - 0.0Colombia (Boyacá) Myrmicinae indet., Camponotus sp., Gnamptogenys sp., Pheidole sp., Solenopsis sp., Crematogaster sp., Zacryptocerus sp., Cephalotes sp., Paratrechina sp., Acromyrmex sp., Rogeria sp. (215881)
Holocene0.0117 - 0.0Dominican Republic Cephalotes resinae, Cephalotes taino (138880)
Holocene0.0117 - 0.0Colombia (Santander) Camponotus sp., Gnamptogenys sp., Pheidole sp., Solenopsis sp., Crematogaster sp., Zacryptocerus sp., Strumigenys sp., Paratrechina sp., Atta sp., Myrmecina sp., Smithistruma sp. (213295)