Parana, Pueblo Brugo to Diamante, Ituzaingo Fm. (Miocene of Argentina)

Also known as “Conglomerado osífero” (= “Mesopotamiense”, sensu Frenguelli, 1920)

Where: Entre Rios, Argentina (31.5° S, 60.2° W: paleocoordinates 31.6° S, 58.1° W)

• coordinate estimated from map

• local area-level geographic resolution

When: Huayquerian mammal zone, Ituzaingo Formation, Huayquerian (9.0 - 6.8 Ma)

• Schmidt 2015: fossils from the base of the Ituzaingo Fm.

Environment/lithology: marine; sandstone and mudstone

Size classes: macrofossils, mesofossils

Preservation: original phosphate

Primary reference: A. L. Cione, M. M. Azpelicueta, M. Bond, A. A. Carlini, J. R. Casciotta, M. A. Cozzuol, M. Fuente, Z. Gasparini, F. J. Goin, J. Noriega, G. J. Scillato-Yane, L. Soibelzon, E. P. Tonni, D. Verzi, and M. G. Vucetich. 2000. Miocene vertebrates from Entre Rios province, eastern Argentina. Serie Correlacion Geologica 14:191-237 [M. Uhen/M. Uhen]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 55600: authorized by Mark Uhen, entered by Mark Uhen on 01.11.2005, edited by Philip Mannion, Juan Carrillo and Miranta Kouvari

Creative Commons license: CC BY (attribution)

Taxonomic list

unclassified
  -
Actinopteri
 Characiformes - Characidae
 Characiformes - Cynodontidae
Reptilia
 Testudines - Panchelidae
Phrynops cf. geoffroanus Schweigger 1812 side-necked turtle of Geoffroy
Aves
 Cariamae - Phorusrhacidae
"Phororhacidae indet." = Phorusrhacidae, "? Onactornis sp." = Devincenzia, Andalgalornis sp.
"Phororhacidae indet." = Phorusrhacidae Ameghino 1889 terror bird
"? Onactornis sp." = Devincenzia Kraglievich 1932 terror bird
Andalgalornis sp. Patterson and Kraglievich 1960 terror bird
 Gruiformes - Rallidae
Rallidae indet. Vigors 1825 rail
 Rheiformes - Rheidae
Rheidae indet. Bonaparte 1849 rhea
 Phoenicopteriformes - Palaelodidae
Megapaloelodus sp. Miller 1944 swimming flamingo
 Phoenicopteriformes - Phoenicopteridae
Phoenicopteridae indet. Bonaparte 1838 flamingo
Reptilia
 Loricata -
Crocodylia indet. crocodilian
 Crocodylia - Alligatoridae
"Carandaisuchus nativus n. gen. n. sp." = Mourasuchus arendsi9, Caiman jacare, Caiman cf. yacare1, Caiman australis, "Caiman latirostris" = Caiman latirostris, "Caiman lutescens n. sp." = Caiman latirostris1, Caiman gasparinae n. sp.2
"Carandaisuchus nativus n. gen. n. sp." = Mourasuchus arendsi9 Bocquentin Villanueva 1984 crocodilian
MLP 73-IV-15-8 - holotype (posterior portion of skull); hypodigm: MLP 73-IV-15-9 (posterior portion of skull)
Caiman cf. yacare1 Daudin 1802 caiman
MLP 73-IV-15-5, MLP 73-IV-15-6, MLP 73-IV-5-17, MACN PV 5417
Caiman australis Bravard 1858 caiman
"Caiman latirostris" = Caiman latirostris Daudin 1801 broud-snouted caiman
"Caiman lutescens n. sp." = Caiman latirostris1 Daudin 1801 broud-snouted caiman
MACN-PV 13551
Caiman gasparinae n. sp.2 Bona and Carabajal 2013 caiman
MLP-73-IV-15-1 - holotype (skull fragments represented by a rostrum with articulated fragments of premaxillae, maxillae, nasals, left lacrymal and a partial braincase lacking the basicranium); referred material - MACN PV 5555 (fragment of right premaxilla)
 Crocodylia -
Gryposuchus neogaeus Rusconi 1933 crocodilian
 Squamata -
Mammalia
 Notoungulata - Toxodontidae
Stenotephanos plicidens Ameghino 1885 notoungulate
Eutomodus elautus Ameghino 1886 notoungulate
? Berroia sp. Kraglievich 1931 notoungulate
Dilobodon lutarius Ameghino 1886 notoungulate
Xotodon foricurvatus Ameghino 1885 notoungulate
Xotodon doellojuradi Frenguelli 1920 notoungulate
Dinotoxodon paranensis Burmeister 1885 notoungulate
? Palaeotoxodon protoburmeisteri Ameghino 1887 notoungulate
? Palaeotoxodon virgatus Ameghino 1886 notoungulate
Palaeotoxodon paranensis Laurillard 1842 notoungulate
 Notoungulata - Haplodontheriidae
"Toxodontherium compressum" = Toxodontherium compressum Ameghino 1883 notoungulate
"Haplodontherium wildei" = Toxodontherium compressum Ameghino 1883 notoungulate
Toxodontherium reverendum Ameghino 1889 notoungulate
Haplodontherium limun notoungulate
Pachynodon modicus Burmeister 1891 notoungulate
 Notoungulata - Interatheriidae
Munizia paranensis Kraglievich 1931 notoungulate
 Notoungulata - Mesotheriidae
Eutypotherium sp.18 Roth 1901 notoungulate
 Notoungulata - Protypotheriidae
Protypotherium antiquum Ameghino 1882 notoungulate
 Rodentia - Simimyidae
Eumysops parodii Kraglievich 1945 rodent
Eumysops parodii6 Kraglievich 1945 rodent
“Eumysops” parodii
 Rodentia - Neoepiblemidae
Phoberomys sp., Phoberomys sp.6, "Phoberomys burmeisteri" = Phoberomys burmeisteri13, "Phoberomys lozanoi n. sp." = Phoberomys burmeisteri15, "Phoberomys minima n. sp." = Phoberomys burmeisteri15, "Phoberomys minima" = Phoberomys burmeisteri6, "Phoberomys praecursor n. sp." = Phoberomys burmeisteri14, "Phoberomys insolita n. sp." = Phoberomys burmeisteri15, Perimys scalabrinianus6, Neoepiblema sp., "Neoepiblema ambrosettianus" = Neoepiblema horridula17, Epiblema horridula12
Phoberomys sp. Kraglievich 1926 caviomorph
Phoberomys sp.6 Kraglievich 1926 caviomorph
"Phoberomys burmeisteri" = Phoberomys burmeisteri13 Kraglievich 1926 caviomorph
"Phoberomys lozanoi n. sp." = Phoberomys burmeisteri15 Kraglievich 1926 caviomorph
"Phoberomys minima n. sp." = Phoberomys burmeisteri15 Kraglievich 1926 caviomorph
"Phoberomys minima" = Phoberomys burmeisteri6 Kraglievich 1926 caviomorph
"Phoberomys praecursor n. sp." = Phoberomys burmeisteri14 Kraglievich 1926 caviomorph
"Phoberomys insolita n. sp." = Phoberomys burmeisteri15 Kraglievich 1926 caviomorph
Perimys scalabrinianus6 Ameghino 1889 caviomorph
Neoepiblema sp. Ameghino 1889 caviomorph
"Neoepiblema ambrosettianus" = Neoepiblema horridula17 Ameghino 1886 caviomorph
Epiblema horridula12 Ameghino 1886 caviomorph
upper right molar - collected by F. Ameghino 1886
 Rodentia - Chinchillidae
"Lagostomopsis sp." = Lagostomus (Lagostomopsis), "Lagostomopsis pallidens" = Lagostomus (Lagostomopsis)6, "Lagostomopsis antiquus" = Lagostomus (Lagostomopsis)6
"Lagostomopsis sp." = Lagostomus (Lagostomopsis) Kraglievich 1926 viscacha
"Lagostomopsis pallidens" = Lagostomus (Lagostomopsis)6 viscacha
"Lagostomopsis antiquus" = Lagostomus (Lagostomopsis)6 viscacha
 Rodentia - Dinomyidae
Rusconia crassidens6 Kraglievich 1931 caviomorph
Tetrastylus sp. Ameghino 1886 caviomorph
Tetrastylus atavus6 caviomorph
T. (Protelicomys) atavus
Isostylomys sp. Kraglievich 1926 caviomorph
Isostylomys laurillardi6 Ameghino 1883 caviomorph
Diaphoromys sp. Kraglievich 1931 caviomorph
Diaphoromys mesopotamicus6 Kraglievich 1932 caviomorph
Olenopsis typicus6 Ameghino 1889 caviomorph
Paranamys sp. Kraglievich 1934 caviomorph
"Drytomomys typicus" = Paranamys typicus7 Ameghino 1889 caviomorph
Protomegamys sp. Kraglievich 1932 caviomorph
Doellomys sp. Alvarez 1947 caviomorph
Doellomys parcus6 caviomorph
Eumegamysops sp. Alvarez 1947 caviomorph
Eumegamysops praependens6 Ameghino 1886 caviomorph
Pentastylomys sp. Kraglievich 1926 caviomorph
Carlesia sp. Kraglievich 1926 caviomorph
Carlesia pendolai6 caviomorph
Briaromys sp. Ameghino 1889 caviomorph
Briaromys trouessartianus6 Ameghino 1889 caviomorph
Pseudosigmomys sp. Kraglievich 1931 caviomorph
Potamarchus sp. Burmeister 1885 caviomorph
Potamarchus murinus6 Burmeister 1885 caviomorph
Telodontomys compressidens6 Kraglievich 1931 caviomorph
Eumegamys sp. Kraglievich 1926 caviomorph
Eumegamys dubius6 caviomorph
Eumegamys depressidens6 caviomorph
"Eumegamys" depressidens
Eumegamys paranensis6 Kraglievich 1926 caviomorph
Megamys burmeisteri12 Ameghino 1886 caviomorph
collected by F. Ameghino 1886
Eumegamys scalabrinianus6 Kraglievich 1926 caviomorph
Eumegamys contortus6 Ameghino 1889 caviomorph
Colpostemma sp. Ameghino 1891 caviomorph
Gyriabrus sp. Ameghino 1891 caviomorph
Gyriabrus holmbergi6 caviomorph
Gyriabrus indivisus6 caviomorph
 Rodentia - Hydrochoeridae
Anatochoerus sp., Cantracavia sp.6, "Protohydrochoerus sp." = Phugatherium6, "Anchimysops sp." = Phugatherium, Phugatherium cataclisticum, "Cardiatherium sp." = Cardiatherium, "Plexochoerus sp." = Cardiatherium, "Kiyutherium sp." = Cardiatherium, "Procardiatherium sp." = Cardiatherium, "Anchimys sp." = Cardiatherium, Cardiatherium paranense n. sp.16, Contracavia sp.
Anatochoerus sp. Vucetich and Mones 1991 caviomorph
Cantracavia sp.6 Burmeister 1885 caviomorph
"Protohydrochoerus sp." = Phugatherium6 Ameghino 1887 caviomorph
"Anchimysops sp." = Phugatherium Ameghino 1887 caviomorph
Phugatherium cataclisticum Ameghino 1887 caviomorph
"Cardiatherium sp." = Cardiatherium Ameghino 1883 caviomorph
"Plexochoerus sp." = Cardiatherium Ameghino 1883 caviomorph
"Kiyutherium sp." = Cardiatherium Ameghino 1883 caviomorph
"Procardiatherium sp." = Cardiatherium Ameghino 1883 caviomorph
"Anchimys sp." = Cardiatherium Ameghino 1883 caviomorph
Cardiatherium paranense n. sp.16 Ameghino 1883 caviomorph
Contracavia sp. Burmeister 1885 caviomorph
 Rodentia - Caviidae
Caviodon sp. Ameghino 1885 caviomorph
Cardiomys sp. Ameghino 1885 caviomorph
Palaeocavia sp. Ameghino 1889 caviomorph
Pliodolichotis sp.5 Kraglievich 1927 mara
Prodolichotis sp. Kraglievich 1932 caviomorph
Parodimys sp. Kraglievich 1932 caviomorph
 Rodentia - Erethizontidae
Paradoxomys sp. Ameghino 1885 porcupine
Paradoxomys cancrivorus6 Ameghino 1885 porcupine
 Rodentia - Abrocomidae
Protabrocoma sp. Kraglievich 1927 chinchilla rat
Protabrocoma paranensis6 chinchilla rat
 Rodentia - Echimyidae
Myocastor sp. Kerr 1782 spiny rat
Myocastor obesus6 spiny rat
Myocastor sinuata6 spiny rat
Haplostropha sp. Ameghino 1891 spiny rat
 Rodentia - Myocastoridae
Strophostephanos sp.5 Ameghino 1891 caviomorph
 Rodentia - Echinomyidae
Discolomys cuneus6 Ameghino 1889 rodent
"Discolomys"
 Carnivora - Procyonidae
Cyonasua argentina Ameghino 1885 procyonid carnivore
 Cingulata - Dasypodidae
Proeuphractus limpidus Ameghino 1886 armadillo
Macroeuphractus retusus19 Ameghino 1887 armadillo
but also ref 15016
Chasicotatus spinozai20 Scillato-Yané et al. 2010 armadillo
? Zaedyus sp. Ameghino 1889 armadillo
Dasypus neogaeus Ameghino 1891 armadillo
 Phyllophaga - Nothrotheriidae
Neohapalops rothi Kraglievich 1923 edentate
Pronothrotherium mirabilis Kraglievich 1925 edentate
 Xenarthra - Megalonychidae
Amphiocnus paranense, Protomegalonyx doellojuradoi, Protomegalonyx praecursor, "Ortotherium laticurvatum" = Ortotherium laticurvatum, "Ortotherium scrofum" = Ortotherium laticurvatum, "Amphiocnus seneus" = Ortotherium laticurvatum, "Ortotherium schlosseri" = Ortotherium laticurvatum, Ortotherium robustum, Pliomorphus robustus, "Pliomorphus mutilatus" = Pliomorphus mutilatus, "Pliomorphus ameghinoi" = Pliomorphus mutilatus, "Pliomorphus brevis" = Pliomorphus mutilatus, "Ortotherium brevirostrum n. sp." = Mesopotamocnus brevirostrum, Megalonychops primigenius, Torcellia paranense
Amphiocnus paranense Kraglievich 1922 edentate
Protomegalonyx doellojuradoi Kraglievich 1925 edentate
Protomegalonyx praecursor Kraglievich 1925 edentate
"Ortotherium laticurvatum" = Ortotherium laticurvatum Ameghino 1885 edentate
"Ortotherium scrofum" = Ortotherium laticurvatum Ameghino 1885 edentate
"Amphiocnus seneus" = Ortotherium laticurvatum Ameghino 1885 edentate
"Ortotherium schlosseri" = Ortotherium laticurvatum Ameghino 1885 edentate
Ortotherium robustum Ameghino 1881 edentate
Pliomorphus robustus Ameghino 1885 edentate
"Pliomorphus mutilatus" = Pliomorphus mutilatus Ameghino 1885 edentate
"Pliomorphus ameghinoi" = Pliomorphus mutilatus Ameghino 1885 edentate
"Pliomorphus brevis" = Pliomorphus mutilatus Ameghino 1885 edentate
"Ortotherium brevirostrum n. sp." = Mesopotamocnus brevirostrum Bordas 1941 edentate
Holotype: MACN Pv 13656, type of Ortotherium brevirostrum Bordas, 1942
Megalonychops primigenius Kraglievich 1926 edentate
Torcellia paranense Kraglievich 1923 edentate
 Xenarthra - Megatheriidae
Eomegatherium nanum4 Burmeister 1891 edentate
"Pliomegatherium lelongi" = Pliomegatherium lelongi Kraglievich 1930 edentate
"Pliomegatherium paranensis" = Pliomegatherium lelongi3 Kraglievich 1930 edentate
Megatherium antiquum3 Ameghino 1885 edentate
Pyramiodontherium sp.4 Rovereto 1914 edentate
"Promegatherium remulsum" = Pyramiodontherium remulsum3 Ameghino 1886 edentate
Promegatherium smaltatum Ameghino 1883 edentate
Promegatherium parvulum Ameghino 1891 edentate
 Xenarthra - Mylodontidae
"Sphenotherus paranensis" = Sphenotherus zavaletianus Ameghino 1891 edentate
Strabosodon acuticavus Ameghino 1891 edentate
Strabosodon obtusicavus Ameghino 1891 edentate
Promylodon paranensis Ameghino 1883 edentate
Ranculcus scalabrinianus Ameghino 1891 edentate
Megabradys darwini Scillato-Yané 1981 edentate
Prolestodon antiquus Ameghino 1885 edentate
Prolestodon paranensis Ameghino 1889 edentate
Octomylodon aversus Ameghino 1904 edentate
Scelidotheriinae indet. Ameghino 1889 edentate
 Xenarthra -
Protoglyptodon primiformis Ameghino 1885 edentate
Palaehoplophorus antiquus Ameghino 1883 edentate
Chlamyphractus pressulus Ameghino 1885 edentate
 Xenarthra - Panochthidae
Hoplophorus verus20 Ameghino 1889 edentate
 Xenarthra - Pampatheriidae
Scirrotherium carinatum n. sp.20 Gois et al. 2013 edentate
Kraglievichia paranense Ameghino 1883 edentate
 Xenarthra - Glyptodontidae
Plohophorus paranensis Ameghino 1891 glyptodon
Paraglyptodon paranensis20 Castellanos 1953 glyptodon
Eleutherocercus paranensis Castellanos 1927 glyptodon
Comaphorus concisus Ameghino 1886 glyptodon
 Xenarthra -
"Berthawyleria sp." = Pseudoplohophorus Castellanos 1926 edentate
Urotherium interundatum Ameghino 1885 edentate
Pseudoeuryurus lelongianus Ameghino 1889 edentate
? Trachycalyptus cingulatus Ameghino 1889 edentate
Parahoplophorus paranensis Ameghino 1883 edentate
 Ungulata - Macraucheniidae
Scalabrinitherium rothii Ameghino 1885 eutherian
Scalabrinitherium bravardi Ameghino 1883 eutherian
Paranauchenia denticulata Ameghino 1891 eutherian
Promacrauchenia antiqua Ameghino 1887 eutherian
Cullinia sp.18 Cabrera and Kraglievich 1931 eutherian
 Ungulata - Proterotheriidae
"Proterotherium gradatum" = Brachytherium gradatum, "Brachytherium cuspidatum" = Brachytherium cuspidatum, "Licaphrium mesopotamiense" = Brachytherium cuspidatum, "Epitherium paranensis" = Brachytherium cuspidatum, Neobrachytherium ameghinoi18, Proterotherium cervioides, "Coelosoma eversa" = Diadiaphorus eversus
"Proterotherium gradatum" = Brachytherium gradatum Ameghino 1891 eutherian
"Brachytherium cuspidatum" = Brachytherium cuspidatum Ameghino 1883 eutherian
"Licaphrium mesopotamiense" = Brachytherium cuspidatum Ameghino 1883 eutherian
"Epitherium paranensis" = Brachytherium cuspidatum Ameghino 1883 eutherian
Neobrachytherium ameghinoi18 Soria 2001 eutherian
Proterotherium cervioides Ameghino 1883 eutherian
"Coelosoma eversa" = Diadiaphorus eversus Soria 2001 eutherian
 Ungulata - Mesorhinidae
"Oxyodontherium zeballosi" = Oxyodontherium zeballozi, "Mesorhinus piramydatus" = Oxyodontherium piramydatus
"Oxyodontherium zeballosi" = Oxyodontherium zeballozi Ameghino 1883 eutherian
"Mesorhinus piramydatus" = Oxyodontherium piramydatus Ameghino 1885 eutherian
 Sparassodonta - Thylacosmilidae
Thylacosmilus atrox10 Riggs 1933 metatherian
 Sparassodonta -
"Notictis ortizi" = Notictis ortizii Ameghino 1889 metatherian
Stylocynus paranensis Mercerat 1917 metatherian
 Didelphimorphia - Didelphidae
Chironectes sp.10 Illiger 1811 water opossum
Philander entrerianus10 Ameghino 1899 four-eyed opossum
Zygolestes paranensis10 Ameghino 1898 opossum
 Cetacea - Iniidae
Iniidae indet.11, "Saurodelphis gigas n. sp." = Saurocetes gigas8
Iniidae indet.11 Gray 1846 toothed whale
"Saurodelphis gigas n. sp." = Saurocetes gigas8 Cozzuol 1988 toothed whale