PBDB Collection

Parana, Pueblo Brugo to Diamante, Ituzaingo Fm. (Neogene of Argentina)

Also known as “Conglomerado osífero” (= “Mesopotamiense”, sensu Frenguelli, 1920)

Where: Entre Rios, Argentina (31.5° S, 60.2° W: paleocoordinates 31.6° S, 58.1° W)

• coordinate estimated from map

• local area-level geographic resolution

When: Huayquerian mammal zone, Ituzaingo Formation, Huayquerian (8.0 - 5.0 Ma)

• Schmidt 2015: fossils from the base of the Ituzaingo Fm.

Environment/lithology: marine; sandstone and mudstone

Size classes: macrofossils, mesofossils

Preservation: original phosphate

Primary reference: A. L. Cione, M. M. Azpelicueta, M. Bond, A. A. Carlini, J. R. Casciotta, M. A. Cozzuol, M. Fuente, Z. Gasparini, F. J. Goin, J. Noriega, G. J. Scillato-Yane, L. Soibelzon, E. P. Tonni, D. H. Verzi, and M. G. Vucetich. 2000. Miocene vertebrates from Entre Rios province, eastern Argentina. Serie Correlacion Geologica 14:191-237 [M. Uhen/M. Uhen/O. Alfaro]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 55600: authorized by Mark Uhen, entered by Mark Uhen on 01.11.2005, edited by Philip Mannion, Juan Carrillo, Miranta Kouvari, Evangelos Vlachos, Grace Varnham and Kateryn Pino

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

unclassified
-	Macrahinga paranensis, Dentrocheninae indet., Mycteriini indet. Macrahinga paranensis Dentrocheninae indet. Mycteriini indet.
Actinopteri
Characiformes - Cynodontidae	Cynodontidae indet. Cynodontidae indet.
Characiformes - Characidae	Colossoma macropomus Colossoma macropomus
Aves
Gruiformes - Rallidae	Rallidae indet. Rallidae indet. Vigors 1825 rail
Gruiformes - Phorusrhacidae	"Phororhacidae indet." = Phorusrhacidae, "? Onactornis sp." = Devincenzia, Andalgalornis sp. "Phororhacidae indet." = Phorusrhacidae Ameghino 1889 terror bird "? Onactornis sp." = Devincenzia Kraglievich 1932 terror bird Andalgalornis sp. Patterson and Kraglievich 1960 terror bird
Rheiformes - Rheidae	Rheidae indet. Rheidae indet. Bonaparte 1849 rhea
Phoenicopteriformes - Phoenicopteridae	Phoenicopteridae indet. Phoenicopteridae indet. Bonaparte 1838 flamingo
Phoenicopteriformes - Palaelodidae	Megapaloelodus sp. Megapaloelodus sp. Miller 1944 swimming flamingo
Reptilia
Loricata -	Crocodylia indet. Crocodylia indet. crocodilian
Crocodylia - Alligatoridae	Caiman jacare, Caiman gasparinae n. sp.², "Caiman latirostris" = Caiman latirostris, "Caiman lutescens n. sp." = Caiman latirostris¹, Caiman australis, Caiman cf. yacare¹, "Carandaisuchus nativus n. gen. n. sp." = Mourasuchus arendsi¹⁰ Caiman jacare caiman Caiman gasparinae n. sp.² Bona and Carabajal 2013 caiman MLP-73-IV-15-1 - holotype (skull fragments represented by a rostrum with articulated fragments of premaxillae, maxillae, nasals, left lacrymal and a partial braincase lacking the basicranium); referred material - MACN PV 5555 (fragment of right premaxilla) "Caiman latirostris" = Caiman latirostris Daudin 1801 broud-snouted caiman "Caiman lutescens n. sp." = Caiman latirostris¹ Daudin 1801 broud-snouted caiman MACN-PV 13551 Caiman australis Bravard 1858 caiman Caiman cf. yacare¹ Daudin 1802 caiman MLP 73-IV-15-5, MLP 73-IV-15-6, MLP 73-IV-5-17, MACN PV 5417 "Carandaisuchus nativus n. gen. n. sp." = Mourasuchus arendsi¹⁰ Bocquentin Villanueva 1984 crocodilian MLP 73-IV-15-8 - holotype (posterior portion of skull); hypodigm: MLP 73-IV-15-9 (posterior portion of skull)
Crocodylia - Gavialidae	Gryposuchus neogaeus Gryposuchus neogaeus Rusconi 1933 crocodilian
Testudines - Cheloniidae	Cheloniidae indet.⁹ Cheloniidae indet.⁹ Bonaparte 1832 sea turtle MACN Pv 14224, nearly complete right mandible
Testudines - Chelidae	Phrynops cf. geoffroanus Phrynops cf. geoffroanus Schweigger 1812 side-necked turtle of Geoffroy
Squamata -	Tupinambis cf. merianae Tupinambis cf. merianae tegu
Mammalia
Didelphimorphia - Didelphidae	Chironectes sp.¹¹, Philander entrerianus¹¹, Zygolestes paranensis¹¹ Chironectes sp.¹¹ Illiger 1811 water opossum Philander entrerianus¹¹ Ameghino 1899 four-eyed opossum Zygolestes paranensis¹¹ Ameghino 1898 opossum
Sparassodonta -	Stylocynus paranensis, "Notictis ortizi" = Notictis ortizii Stylocynus paranensis Mercerat 1917 metatherian "Notictis ortizi" = Notictis ortizii Ameghino 1889 metatherian
Sparassodonta - Thylacosmilidae	Thylacosmilus atrox¹¹ Thylacosmilus atrox¹¹ Riggs 1933 metatherian
Cetacea - Iniidae	Iniidae indet.¹², "Saurodelphis gigas n. sp." = Saurocetes gigas⁸ Iniidae indet.¹² Gray 1846 toothed whale "Saurodelphis gigas n. sp." = Saurocetes gigas⁸ Cozzuol 1988 toothed whale
Panameriungulata - Macraucheniidae	"Mesorhinus piramydatus" = Oxyodontherium piramydatus, "Oxyodontherium zeballosi" = Oxyodontherium zeballozi, Paranauchenia denticulata, Cullinia sp.²⁰, Promacrauchenia antiqua, Scalabrinitherium bravardi, Scalabrinitherium rothii "Mesorhinus piramydatus" = Oxyodontherium piramydatus Ameghino 1885 placental "Oxyodontherium zeballosi" = Oxyodontherium zeballozi Ameghino 1883 placental Paranauchenia denticulata Ameghino 1891 placental Cullinia sp.²⁰ Cabrera and Kraglievich 1931 placental Promacrauchenia antiqua Ameghino 1887 placental Scalabrinitherium bravardi Ameghino 1883 placental Scalabrinitherium rothii Ameghino 1885 placental
Panameriungulata - Proterotheriidae	"Proterotherium gradatum" = Brachytherium gradatum, "Brachytherium cuspidatum" = Brachytherium cuspidatum, "Licaphrium mesopotamiense" = Brachytherium cuspidatum, "Epitherium paranensis" = Brachytherium cuspidatum, Neobrachytherium ameghinoi²⁰, Proterotherium cervioides, "Coelosoma eversa" = Diadiaphorus eversus "Proterotherium gradatum" = Brachytherium gradatum Ameghino 1891 placental "Brachytherium cuspidatum" = Brachytherium cuspidatum Ameghino 1883 placental "Licaphrium mesopotamiense" = Brachytherium cuspidatum Ameghino 1883 placental "Epitherium paranensis" = Brachytherium cuspidatum Ameghino 1883 placental Neobrachytherium ameghinoi²⁰ Soria 2001 placental Proterotherium cervioides Ameghino 1883 placental "Coelosoma eversa" = Diadiaphorus eversus Soria 2001 placental
Carnivora - Procyonidae	Cyonasua argentina Cyonasua argentina Ameghino 1885 procyonid carnivore
Rodentia - Dinomyidae	Rusconia crassidens⁶, Doellomys sp., Doellomys parcus⁶, Pseudosigmomys sp., Pseudosigmomys paranensis⁶, Eumegamysops sp., Eumegamysops praependens⁶, Briaromys sp., Briaromys trouessartianus⁶, Potamarchus sp., Potamarchus sigmodon⁶, Potamarchus murinus⁶, Olenopsis typicus⁶, Pentastylomys sp., Pentastylomys seriei⁶, Eumegamys sp., Eumegamys dubius⁶, Eumegamys depressidens⁶, Eumegamys contortus⁶, Eumegamys paranensis⁶, Eumegamys scalabrinianus⁶, Discolomys cuneus⁶, Colpostemma sp., Tetrastylus aguilari⁶, Tetrastylus diffusus⁶, Tetrastylus robustus⁶, Tetrastylus atavus⁶, Tetrastylus laevigatus⁶, Tetrastylus laevigatus n. sp.²³, Isostylomys sp., Isostylomys ameghinoi⁶, Isostylomys laurillardi⁶, Carlesia sp., Carlesia pendolai⁶, Diaphoromys sp., Diaphoromys compressidens⁶, Diaphoromys gamayensis⁶, Diaphoromys mesopotamicus⁶, Gyriabrus sp., Gyriabrus glutinatus⁶, Gyriabrus indivisus⁶, Gyriabrus rebagliattii⁶, Gyriabrus holmbergi⁶, Protomegamys sp., Protomegamys coligatus⁶, Pentastylodon racedi⁶, Tetrastylomys castellanosi⁶, Paranamys sp., "Drytomomys typicus" = Paranamys typicus⁷, Telodontomys compressidens⁶ Rusconia crassidens⁶ Kraglievich 1931 caviomorph Doellomys sp. Alvarez 1947 caviomorph Doellomys parcus⁶ caviomorph Pseudosigmomys sp. Kraglievich 1931 caviomorph Pseudosigmomys paranensis⁶ caviomorph Eumegamysops sp. Alvarez 1947 caviomorph Eumegamysops praependens⁶ Ameghino 1886 caviomorph Briaromys sp. Ameghino 1889 caviomorph Briaromys trouessartianus⁶ Ameghino 1889 caviomorph Potamarchus sp. Burmeister 1885 caviomorph Potamarchus sigmodon⁶ caviomorph Potamarchus murinus⁶ Burmeister 1885 caviomorph Olenopsis typicus⁶ Ameghino 1889 caviomorph Pentastylomys sp. Kraglievich 1926 caviomorph Pentastylomys seriei⁶ caviomorph Eumegamys sp. Kraglievich 1926 caviomorph Eumegamys dubius⁶ caviomorph Eumegamys depressidens⁶ caviomorph "Eumegamys" depressidens Eumegamys contortus⁶ Ameghino 1889 caviomorph Eumegamys paranensis⁶ Kraglievich 1926 caviomorph Eumegamys scalabrinianus⁶ Kraglievich 1926 caviomorph Discolomys cuneus⁶ Ameghino 1889 caviomorph "Discolomys" Colpostemma sp. Ameghino 1891 caviomorph Tetrastylus aguilari⁶ caviomorph Tetrastylus diffusus⁶ caviomorph Tetrastylus robustus⁶ caviomorph Tetrastylus atavus⁶ caviomorph T. (Protelicomys) atavus Tetrastylus laevigatus⁶ Ameghino 1886 caviomorph Tetrastylus laevigatus n. sp.²³ Ameghino 1886 caviomorph Isostylomys sp. Kraglievich 1926 caviomorph Isostylomys ameghinoi⁶ caviomorph Isostylomys laurillardi⁶ Ameghino 1883 caviomorph Carlesia sp. Kraglievich 1926 caviomorph Carlesia pendolai⁶ caviomorph Diaphoromys sp. Kraglievich 1931 caviomorph Diaphoromys compressidens⁶ caviomorph Diaphoromys gamayensis⁶ Kraglievich 1931 caviomorph Diaphoromys mesopotamicus⁶ Kraglievich 1932 caviomorph Gyriabrus sp. Ameghino 1891 caviomorph Gyriabrus glutinatus⁶ caviomorph Gyriabrus indivisus⁶ caviomorph Gyriabrus rebagliattii⁶ caviomorph Gyriabrus holmbergi⁶ Ameghino 1885 caviomorph Protomegamys sp. Kraglievich 1932 caviomorph Protomegamys coligatus⁶ caviomorph Pentastylodon racedi⁶ caviomorph Tetrastylomys castellanosi⁶ caviomorph Paranamys sp. Kraglievich 1934 caviomorph "Drytomomys typicus" = Paranamys typicus⁷ Ameghino 1889 caviomorph Telodontomys compressidens⁶ Kraglievich 1931 caviomorph
Rodentia - Neoepiblemidae	Phoberomys sp., Phoberomys sp.⁶, "Phoberomys burmeisteri" = Phoberomys burmeisteri¹⁴, "Phoberomys lozanoi n. sp." = Phoberomys burmeisteri¹⁶, "Phoberomys insolita n. sp." = Phoberomys burmeisteri¹⁶, "Phoberomys minima n. sp." = Phoberomys burmeisteri¹⁶, "Phoberomys minima" = Phoberomys burmeisteri⁶, "Megamys burmeisteri" = Phoberomys burmeisteri¹³, "Phoberomys praecursor n. sp." = Phoberomys burmeisteri¹⁵, Neoepiblema sp., Epiblema horridula¹³, "Neoepiblema ambrosettianus" = Neoepiblema horridula¹⁸, Perimys scalabrinianus⁶ Phoberomys sp. Kraglievich 1926 caviomorph Phoberomys sp.⁶ Kraglievich 1926 caviomorph "Phoberomys burmeisteri" = Phoberomys burmeisteri¹⁴ Kraglievich 1926 caviomorph "Phoberomys lozanoi n. sp." = Phoberomys burmeisteri¹⁶ Kraglievich 1926 caviomorph "Phoberomys insolita n. sp." = Phoberomys burmeisteri¹⁶ Kraglievich 1926 caviomorph "Phoberomys minima n. sp." = Phoberomys burmeisteri¹⁶ Kraglievich 1926 caviomorph "Phoberomys minima" = Phoberomys burmeisteri⁶ Kraglievich 1926 caviomorph "Megamys burmeisteri" = Phoberomys burmeisteri¹³ Kraglievich 1926 caviomorph collected by F. Ameghino 1886 "Phoberomys praecursor n. sp." = Phoberomys burmeisteri¹⁵ Kraglievich 1926 caviomorph Neoepiblema sp. Ameghino 1889 caviomorph Epiblema horridula¹³ Ameghino 1886 caviomorph upper right molar - collected by F. Ameghino 1886 "Neoepiblema ambrosettianus" = Neoepiblema horridula¹⁸ Ameghino 1886 caviomorph Perimys scalabrinianus⁶ Ameghino 1889 caviomorph
Rodentia - Erethizontidae	Paradoxomys sp., Paradoxomys cancrivorus⁶ Paradoxomys sp. Ameghino 1885 porcupine Paradoxomys cancrivorus⁶ Ameghino 1885 porcupine
Rodentia - Chinchillidae	"Lagostomopsis sp." = Lagostomus (Lagostomopsis), "Lagostomopsis pallidens" = Lagostomus (Lagostomopsis)⁶, "Lagostomopsis antiquus" = Lagostomus (Lagostomopsis)⁶ "Lagostomopsis sp." = Lagostomus (Lagostomopsis) Kraglievich 1926 viscacha "Lagostomopsis pallidens" = Lagostomus (Lagostomopsis)⁶ viscacha "Lagostomopsis antiquus" = Lagostomus (Lagostomopsis)⁶ viscacha
Rodentia - Echimyidae	Myocastor sp., Myocastor paranensis⁶, Myocastor obesus⁶, Myocastor sinuata⁶, Eumysops parodii, Eumysops parodii⁶, Haplostropha sp., Haplostropha sacabriniana⁶ Myocastor sp. Kerr 1782 spiny rat Myocastor paranensis⁶ spiny rat Myocastor obesus⁶ spiny rat Myocastor sinuata⁶ spiny rat Eumysops parodii Kraglievich 1945 spiny rat Eumysops parodii⁶ Kraglievich 1945 spiny rat “Eumysops” parodii Haplostropha sp. Ameghino 1891 spiny rat Haplostropha sacabriniana⁶ spiny rat
Rodentia - Myocastoridae	Strophostephanos sp.⁵, Strophostephanos jheringi⁶ Strophostephanos sp.⁵ Ameghino 1891 caviomorph Strophostephanos jheringi⁶ caviomorph
Rodentia - Abrocomidae	Protabrocoma sp., Protabrocoma paranensis⁶ Protabrocoma sp. Kraglievich 1927 chinchilla rat Protabrocoma paranensis⁶ chinchilla rat
Rodentia - Caviidae	Parodimys sp., Pliodolichotis sp.⁵, Palaeocavia sp., Prodolichotis sp., Cardiomys sp., Caviodon sp. Parodimys sp. Kraglievich 1932 caviomorph Pliodolichotis sp.⁵ Kraglievich 1927 mara Palaeocavia sp. Ameghino 1889 caviomorph Prodolichotis sp. Kraglievich 1932 caviomorph Cardiomys sp. Ameghino 1885 caviomorph Caviodon sp. Ameghino 1885 caviomorph
Rodentia - Hydrochoeridae	"Cardiatherium sp." = Cardiatherium, Cardiatherium paranense n. sp.¹⁷, "Anchimys sp." = Cardiatherium, "Procardiatherium sp." = Cardiatherium, "Kiyutherium sp." = Cardiatherium, "Plexochoerus sp." = Cardiatherium, Cantracavia sp.⁶, Anatochoerus sp., "Protohydrochoerus sp." = Phugatherium⁶, Phugatherium cataclisticum, "Anchimysops sp." = Phugatherium, Contracavia sp. "Cardiatherium sp." = Cardiatherium Ameghino 1883 caviomorph Cardiatherium paranense n. sp.¹⁷ Ameghino 1883 caviomorph "Anchimys sp." = Cardiatherium Ameghino 1883 caviomorph "Procardiatherium sp." = Cardiatherium Ameghino 1883 caviomorph "Kiyutherium sp." = Cardiatherium Ameghino 1883 caviomorph "Plexochoerus sp." = Cardiatherium Ameghino 1883 caviomorph Cantracavia sp.⁶ Burmeister 1885 caviomorph Anatochoerus sp. Vucetich and Mones 1991 caviomorph "Protohydrochoerus sp." = Phugatherium⁶ Ameghino 1887 caviomorph Phugatherium cataclisticum Ameghino 1887 caviomorph "Anchimysops sp." = Phugatherium Ameghino 1887 caviomorph Contracavia sp. Burmeister 1885 caviomorph
Notoungulata - Mesotheriidae	Eutypotherium sp.²⁰ Eutypotherium sp.²⁰ Roth 1902 notoungulate
Notoungulata - Interatheriidae	Protypotherium antiquum, Munizia paranensis Protypotherium antiquum Ameghino 1882 notoungulate Munizia paranensis Kraglievich 1931 notoungulate
Notoungulata - Toxodontidae	"Stenotephanos plicidens" = Stenotephanos plicidens, "? Palaeotoxodon virgatus" = Stenotephanos plicidens, "Eutomodus elautus" = Stenotephanos plicidens, Xotodon foricurvatus, Xotodon doellojuradi, ? Berroia sp., ? Adinotherium paranense, ? Palaeotoxodon protoburmeisteri, "Dinotoxodon paranensis" = Dinotoxodon paranensis, "Palaeotoxodon paranensis" = Dinotoxodon paranensis, Dilobodon lutarius "Stenotephanos plicidens" = Stenotephanos plicidens Ameghino 1885 notoungulate "? Palaeotoxodon virgatus" = Stenotephanos plicidens Ameghino 1885 notoungulate "Eutomodus elautus" = Stenotephanos plicidens Ameghino 1885 notoungulate Xotodon foricurvatus Ameghino 1885 notoungulate Xotodon doellojuradi Frenguelli 1920 notoungulate ? Berroia sp. Kraglievich 1931 notoungulate ? Adinotherium paranense notoungulate ? Palaeotoxodon protoburmeisteri Ameghino 1887 notoungulate "Dinotoxodon paranensis" = Dinotoxodon paranensis Burmeister 1885 notoungulate "Palaeotoxodon paranensis" = Dinotoxodon paranensis Burmeister 1885 notoungulate Dilobodon lutarius Ameghino 1886 notoungulate
Notoungulata - Haplodontheriidae	"Haplodontherium wildei" = Haplodontherium wildei, "Toxodontherium reverendum" = Haplodontherium wildei, "Haplodontherium limun" = Haplodontherium wildei, "Toxodontherium compressum" = Haplodontherium wildei, Pachynodon modicus "Haplodontherium wildei" = Haplodontherium wildei Ameghino 1885 notoungulate "Toxodontherium reverendum" = Haplodontherium wildei Ameghino 1885 notoungulate "Haplodontherium limun" = Haplodontherium wildei Ameghino 1885 notoungulate "Toxodontherium compressum" = Haplodontherium wildei Ameghino 1885 notoungulate Pachynodon modicus Burmeister 1891 notoungulate
- Mylodontidae	Scelidotheriinae indet., "Sphenotherus paranensis" = Sphenotherus zavaletianus, Ranculcus scalabrinianus, Promylodon paranensis, Strabosodon obtusicavus, Strabosodon acuticavus, Prolestodon paranensis, Prolestodon antiquus, Megabradys darwini, Octomylodon aversus Scelidotheriinae indet. Ameghino 1889 edentate "Sphenotherus paranensis" = Sphenotherus zavaletianus Ameghino 1891 edentate Ranculcus scalabrinianus Ameghino 1891 edentate Promylodon paranensis Ameghino 1883 edentate Strabosodon obtusicavus Ameghino 1891 edentate Strabosodon acuticavus Ameghino 1891 edentate Prolestodon paranensis Ameghino 1889 edentate Prolestodon antiquus Ameghino 1885 edentate Megabradys darwini Scillato-Yané 1981 edentate Octomylodon aversus Ameghino 1904 edentate
Megatherioidea - Megalonychidae	Torcellia paranense, "Ortotherium brevirostrum n. sp." = Mesopotamocnus brevirostrum, Megalonychops primigenius, "Ortotherium laticurvatum" = Ortotherium laticurvatum, "Ortotherium schlosseri" = Ortotherium laticurvatum, "Ortotherium scrofum" = Ortotherium laticurvatum, "Amphiocnus seneus" = Ortotherium laticurvatum, Ortotherium robustum, Protomegalonyx doellojuradoi, Protomegalonyx praecursor, Amphiocnus paranense, Pliomorphus robustus, "Pliomorphus mutilatus" = Pliomorphus mutilatus, "Pliomorphus brevis" = Pliomorphus mutilatus, "Pliomorphus ameghinoi" = Pliomorphus mutilatus Torcellia paranense Kraglievich 1923 edentate "Ortotherium brevirostrum n. sp." = Mesopotamocnus brevirostrum Bordas 1941 edentate Holotype: MACN Pv 13656, type of Ortotherium brevirostrum Bordas, 1942 Megalonychops primigenius Kraglievich 1926 edentate "Ortotherium laticurvatum" = Ortotherium laticurvatum Ameghino 1885 edentate "Ortotherium schlosseri" = Ortotherium laticurvatum Ameghino 1885 edentate "Ortotherium scrofum" = Ortotherium laticurvatum Ameghino 1885 edentate "Amphiocnus seneus" = Ortotherium laticurvatum Ameghino 1885 edentate Ortotherium robustum Ameghino 1881 edentate Protomegalonyx doellojuradoi Kraglievich 1925 edentate Protomegalonyx praecursor Kraglievich 1925 edentate Amphiocnus paranense Kraglievich 1922 edentate Pliomorphus robustus Ameghino 1885 edentate "Pliomorphus mutilatus" = Pliomorphus mutilatus Ameghino 1885 edentate "Pliomorphus brevis" = Pliomorphus mutilatus Ameghino 1885 edentate "Pliomorphus ameghinoi" = Pliomorphus mutilatus Ameghino 1885 edentate
Megatherioidea - Nothrotheriidae	Pronothrotherium mirabilis, Neohapalops rothi Pronothrotherium mirabilis Kraglievich 1925 edentate Neohapalops rothi Kraglievich 1923 edentate
Megatherioidea - Megatheriidae	Eomegatherium nanum⁴, "Pliomegatherium lelongi" = Pliomegatherium lelongi, "Pliomegatherium paranensis" = Pliomegatherium lelongi³, Megatherium antiquum³, Pyramiodontherium sp.⁴, "Promegatherium remulsum" = Pyramiodontherium remulsum³, Promegatherium parvulum, Promegatherium smaltatum Eomegatherium nanum⁴ Burmeister 1891 edentate "Pliomegatherium lelongi" = Pliomegatherium lelongi Kraglievich 1930 edentate "Pliomegatherium paranensis" = Pliomegatherium lelongi³ Kraglievich 1930 edentate Megatherium antiquum³ Ameghino 1885 edentate Pyramiodontherium sp.⁴ Rovereto 1914 edentate "Promegatherium remulsum" = Pyramiodontherium remulsum³ Ameghino 1886 edentate Promegatherium parvulum Ameghino 1891 edentate Promegatherium smaltatum Ameghino 1883 edentate
Cingulata - Dasypodidae	? Zaedyus sp., Proeuphractus limpidus, Macroeuphractus retusus²¹, Chasicotatus spinozai²², Dasypus neogaeus ? Zaedyus sp. Ameghino 1889 armadillo Proeuphractus limpidus Ameghino 1886 armadillo Macroeuphractus retusus²¹ Ameghino 1887 armadillo but also ref 15016 Chasicotatus spinozai²² Scillato-Yané et al. 2010 armadillo Dasypus neogaeus Ameghino 1891 armadillo
Cingulata -	"Berthawyleria sp." = Pseudoplohophorus, Palaehoplophorus antiquus, Protoglyptodon primiformis, Chlamyphractus pressulus, ? Trachycalyptus cingulatus, Pseudoeuryurus lelongianus, Urotherium interundatum "Berthawyleria sp." = Pseudoplohophorus Castellanos 1926 edentate Palaehoplophorus antiquus Ameghino 1883 edentate Protoglyptodon primiformis Ameghino 1885 edentate Chlamyphractus pressulus Ameghino 1885 edentate ? Trachycalyptus cingulatus Ameghino 1889 edentate Pseudoeuryurus lelongianus Ameghino 1889 edentate Urotherium interundatum Ameghino 1885 edentate
Cingulata - Glyptodontidae	Paraglyptodon paranensis²², Comaphorus concisus, Plohophorus paranensis, Eleutherocercus sp.¹⁹ Paraglyptodon paranensis²² Castellanos 1953 glyptodon Comaphorus concisus Ameghino 1886 glyptodon Plohophorus paranensis Ameghino 1891 glyptodon Eleutherocercus sp.¹⁹ Koken 1888 glyptodon
Cingulata - Pampatheriidae	Kraglievichia paranense, "Scirrotherium carinatum n. sp." = Kraglievichia carinatum²² Kraglievichia paranense Ameghino 1883 edentate "Scirrotherium carinatum n. sp." = Kraglievichia carinatum²² Gois et al. 2013 edentate
Cingulata - Panochthidae	Hoplophorus verus²² Hoplophorus verus²² Ameghino 1889 edentate
Cingulata -	Parahoplophorus paranensis Parahoplophorus paranensis Ameghino 1883 edentate

¹Bona and Barrios 2015; ²Bona and Carabajal 2013; ³Brandoni 2006; ⁴Brandoni 2013; ⁵Brandoni 2013; ⁶Candela 2005; ⁷Candela and Nasif 2006; ⁸Cozzuol 1988; ⁹Falbo and Agnolin 2020; ¹⁰Gasparini 1985; ¹¹Goin et al. 2013; ¹²Gutstein et al. 2014; ¹³Kerber et al. 2019; ¹⁴Kraglievich 1926; ¹⁵Kraglievich 1932; ¹⁶Kraglievich 1940; ¹⁷Mones 1991; ¹⁸Negri and Ferigolo 1999; ¹⁹Nuñez-Blasco et al. 2023; ²⁰Schmidt and Cerdeño 2013; ²¹Scillato-Yané 1975; ²²Scillato-Yané et al. 2013; ²³Sostillo et al. 2022