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Titanochelon

Osteichthyes - Testudines - Testudinidae

Discussion

Titanochelon is a new genus created to included mainly the Neogene large testudinids from Europe, which were previously included in the genus Cheirogaster Bergounioux (1935)

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Synonymy list
YearName and author
2014Titanochelon Pérez-García and Vlachos
2017Titanochelon Pérez-García et al.
2018Titanochelon Vlachos and Rabi p. 663
2019Titanochelon Vlachos et al.
2021Titanochelon Perez-Garcia et al.
2021Titanochelon Vlachos
2023Titanochelon Pappa et al.

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
classOsteichthyes
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
RankNameAuthor
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
Sauropsida
classReptilia
Testudinata(Oppel 1811)
orderTestudinesBatsch 1788
suborderCryptodira
Pantestudinoidea
superfamilyTestudinoidea
familyTestudinidaeBatsch 1788
tribeGeochelonini
genusTitanochelon

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. †Titanochelon Pérez-García and Vlachos 2014
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Titanochelon bacharidisi Vlachos et al. 2014
Titanochelon bolivari Hernandez-Pacheco 1917
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Invalid names: Testudo richardi Bergounioux 1938 [synonym]
Titanochelon ginsburgi Broin 1977
Titanochelon kayadibiensis Karl et al. 2021
Titanochelon perpiniana Depéret 1885
Titanochelon punicum Arambourg 1979
Titanochelon schafferi Szalai 1931
Titanochelon schleichi Pappa et al. 2023
Titanochelon vitodurana Biedermann 1862
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Invalid names: Testudo picteti Biedermann 1862 [synonym]
Diagnosis
ReferenceDiagnosis
E. Vlachos 2021The members of Geochelonini represented in the Greek fossil record can be easily distinguished from those belonging to Testudinini (see below) based on several characters from the shell: large size, with individuals that reach or exceed one meter in carapace length; absence of cervical scute; contact between marginal 6 and pleural 3; and absence of a plastral hinge.
I. Pappa et al. 2023belongs to Geochelonini based on the contact between the pleural 3 and marginal 6
Measurements
No measurements are available
Composition: phosphaticsubp
Environment: terrestrialf
Locomotion: actively mobilec
Life habit: ground dwellingf
Diet: herbivoref
Diet 2: herbivoref
Reproduction: oviparousf
Created: 2005-08-26 07:16:55
Modified: 2005-09-09 16:16:46
Source: f = family, c = class, subp = subphylum
References: Ernst and Barbour 1989, Carroll 1988, Hendy et al. 2009

Age range: base of the MN 4 to the top of the Gelasian or 16.90000 to 1.80000 Ma

Collections (73 total)


Time interval Ma Country or state Original ID and collection number
Early/Lower Miocene23.04 - 15.98Portugal (Lisbon) Testudo bolivari (184304)
Burdigalian20.45 - 15.98Spain (Lardero) T. bolivari (217466 217467)
Burdigalian20.45 - 15.98Portugal (Lisbon) T. bolivari (35989)
Burdigalian20.45 - 15.98Spain (Catalonia) T. bolivari (205815)
Burdigalian - Langhian20.45 - 13.82Germany (Baden-Wurttemberg) Geochelone sp. (106003)
Orleanian20.0 - 15.98Spain (Madrid) Geochelone bolivari (40747)
MN 416.9 - 15.98France (Centre) Testudo ginsburgi (69116)
Langhian15.98 - 13.82Portugal (Lisbon) T. bolivari (35986)
Langhian15.98 - 13.82Spain T. bolivari (187337 187341)
Langhian15.98 - 13.82Germany (Bavaria) Geochelone perpiniana (34615)
Langhian15.98 - 13.82Spain (Catalonia) T. sp. (180192)
MN 5 - MN 615.98 - 12.75Türkiye (Ankara) Cheirogaster bolivari (68561)
MN 5 - MN 615.98 - 12.75Portugal Testudo bolivari (65912)
Langhian - Serravallian15.98 - 11.63Spain T. bolivari (187338 187347)
Middle Miocene15.98 - 11.63Portugal (Azambuja Municipality) T. bolivari (184303)
Middle Miocene15.98 - 11.63Spain (Madrid) Cheirogaster bolivari (197674)
Middle Miocene15.98 - 11.63Türkiye (Mugla) Cheirogaster bolivari (184394)
Middle Miocene15.98 - 11.63Spain Cheirogaster bolivari (144667) Testudo bolivari (144666)
Middle Miocene15.98 - 11.63Spain (Segovia) Cheirogaster bolivari (208187)
Badenian13.82 - 12.8Czechia (Brno) T. sp. (207955)
Serravallian13.82 - 11.63Spain Testudo sp. (187336) T. bolivari (187339 187342 187343 187344 187345 187346)
Serravallian13.82 - 11.63Switzerland (Zurich) Testudo picteti (198617) Testudo vitodurana (198616)
Sarmatian12.8 - 11.63Türkiye Cheirogaster bolivari (56563)
MN 7 + 812.75 - 11.1Spain (Cataluña) Testudo richardi (92640)
Vallesian11.63 - 8.725Spain (Catalonia) T. richardi (193491 193496)
Vallesian11.63 - 8.725Spain Cheirogaster richardi (144670)
Tortonian11.63 - 7.246Spain T. bolivari (187340)
Tortonian11.63 - 7.246Germany (Bavaria) T. sp. (206345)
Late/Upper Miocene11.63 - 5.333Türkiye (Afyon) Cheirogaster bolivari (184398)
Late/Upper Miocene11.63 - 5.333Türkiye (Konya) Cheirogaster bolivari (184395)
Late/Upper Miocene11.63 - 5.333Greece (Aegean) Testudo schafferi (182751) Testudo sp. (202120)
Late/Upper Miocene11.63 - 5.333Greece (Attica) Testudo schafferi (182754) Testudo sp. (183121)
Late/Upper Miocene11.63 - 5.333Greece (Kassandra) T. sp. (202330)
Late/Upper Miocene11.63 - 5.333Greece (Chalkidona) Testudo sp. (182750)
Late/Upper Miocene11.63 - 5.333Greece (Pieria) T. sp. (226433 226434 226435 226437 226439 226440)
MN 109.67 - 8.725Spain (Madrid) Testudinidae indet. (197661)
MN 118.725 - 7.9Spain (Alicante) Cheirogaster bolivari (68914)
Turolian8.725 - 5.09Greece (Makedonia Thraki) Chelonia indet. (157582)
Messinian7.246 - 5.333Greece T. schafferi (95690 95691)
Early/Lower Pliocene5.333 - 3.6Tunisia (Bizerte ) Testudo punica (211247)
Early/Lower Pliocene5.333 - 3.6Bulgaria (Yambol) Testudo sp. (184410)
Early/Lower Pliocene5.333 - 3.6Spain (Murcia) T. sp. (185681)
Zanclean5.333 - 3.6France (Pyrénées-Orientales) Testudo perpiniana (178672)
Early/Lower Pliocene - Late/Upper Pliocene5.333 - 2.58Greece (Thessaloniki) Cheirogaster bacharidisi (182672 182673 182674)
Pliocene5.333 - 2.58France (Languedoc-Roussillon Region) Testudo perpiniana (36103)
Zanclean - Piacenzian5.333 - 2.58France (Languedoc-Roussillon Region) Cheirogaster perpiniana (36985)
Ruscinian5.09 - 3.55Greece Testudo sp. (36579)
Late/Upper Pliocene3.6 - 2.58Greece (Grevena) T. sp. (185859)
Late/Upper Pliocene3.6 - 2.58Spain (Alcolea de Calatrava) Cheirogaster sp. (224542)
Gelasian2.58 - 1.8Spain (Granada) T. sp. (77773)
Early/Lower Pleistocene2.58 - 0.774Greece (Lesvos) Testudinata indet. (183341)