Classification of Lujan et al. 2016

G. Testudo Linnaeus 1758
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†Testudo brevitesta Vlachos and Tsoukala 2016
†Testudo kenitrensis Gmira 1993
Testudo kleinmanni Lortet 1883 [Egyptian tortoise]
Testudo marginata Schoepff 1792 [Marginated tortoise]
†Testudo marmorum Gaudry 1862
†Testudo oughlamensis Gmira et al. 2013
G. †Paleotestudo Lartet 1851
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†Paleotestudo canetotiana Lartet 1851
G. Agrionemys Khosatzky and Mlynarsky 1966
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†Agrionemys bessarabica Riabinin 1918
Subg. Testudo (Chersine) Merrem 1820
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