Laño L1A: Late/Upper Campanian, Spain

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Actinopteri - Elopiformes - Phyllodontidae
Phyllodontidae indet. Sauvage 1875
Pereda-Suberbiola et al. 2000
Actinopteri - Amiiformes - Palaeolabridae
? Palaeolabrus sp. Casier 1967
Pereda-Suberbiola et al. 2000
Actinopteri - Lepisosteiformes - Lepisosteidae
Lepisosteus sp. Agassiz 1843
Astibia et al. 1990
    = Atractosteus sp. Rafinesque 1820
Pereda-Suberbiola et al. 2000
Albanerpetontidae
cf. Albanerpeton sp. Estes and Hoffstetter 1976
Astibia et al. 1990
    = Albanerpetontidae indet. Fox and Naylor 1982
Duffaud and Rage 1999
five incomplete dentaries, distal part of 3 humeri
Salientia
Anura indet. (Fischer von Waldheim 1813)
Duffaud and Rage 1999
distal part of 2 humeri, 8 ilia, fragmentary maxillaries, 2 broken sphenethmoids
Salientia - Palaeobatrachidae
Palaeobatrachidae indet. Cope 1865
Duffaud and Rage 1999
43 prearticulars, 1 presacral vert, 2 synsacrums, 6 distal parts of humeri
Salientia - Discoglossidae
Discoglossidae indet. Günther 1859
Astibia et al. 1990
6 prearticulars, 5 presacral verts, 7 sacral verts, 1 urostyle, 7 ilia, 2 humeri
Caudata
Salamandridae indet. Gray 1825
Astibia et al. 1990
may be Salamandra
    = Caudata indet. Scopoli 1777
Duffaud and Rage 1999
1 atlantal centrum, 13 fragmentary vertebrae, 1 humerus
Reptilia - Testudines
de Lapparent and Murelaga 1999
Reptilia - Testudines - Bothremydidae
Polysternon atlanticum Lapparent de Broin and Murelaga 1996
de Lapparent and Murelaga 1999
? Elochelys sp. Nopsca 1931
de Lapparent and Murelaga 1999
right partial epiplastron
Reptilia
Cryptodira indet. Cope 1868
Astibia et al. 1987
    = Dermatemydidae indet. Baur 1888
Astibia et al. 1990
"Dermatemyididae" sic
    = Solemys vermiculata Lapparent de Broin and Murelaga 1996
Joyce 2017
Solemys vermiculata n. sp. Lapparent de Broin and Murelaga 1996
de Lapparent and Murelaga 1999
hypodigme around 260 shell fragments, most of them are peripherals and pleural fragments, around 25 fragments of plastron including the hologype
Helochelydra nopcsai Lapparent de Broin and Murelaga 1999
de Lapparent and Murelaga 1999
partial shell, lots of fragments
Titanosaurinae indet. Lydekker 1893
461 specimens
orig. 1 tooth
    = Titanosauridae indet. Lydekker 1885
Astibia et al. 1990
    = Lirainosaurus astibiae n. gen., n. sp. Sanz et al. 1999
Pereda-Suberbiola et al. 2000
Holotype: MCNA 7458, anteriormost caudal vertebra (Sanz et al., 1999: pl. 2; Díez Díaz et al., 2013: fig. 4). Paratypes: an incomplete braincase (MCNA 7439), two isolated teeth (MCNA 7440, 7441), ten dorsal vertebrae (MCNA 7442-7450, 7456), seven caudal vertebrae (MCNA 1812, 7451-7457), one scapula (MCNA 7459), one coracoid (MCNA 7460), one sternal plate (MCNA 7461), four humeri (MCNA 7462-7465), a fragment of ilium (MCNA 7466), a fragment of a pubis (MCNA 7467), three femora (MCNA 3160, 7468, 7470), a tibia (MCNA 7471), a fibula (MCNA 7472) and two osteoderms (MCNA 14473, 14474) (figured in Sanz et al., 1999).
Reptilia - Avetheropoda
Coelurosauria indet. Huene 1914
Canudo and Ruiz-Omeñaca 2003
Reptilia - Avetheropoda - Dromaeosauridae
Theropoda indet. Marsh 1881
Astibia et al. 1990
    = Dromaeosauridae indet. Colbert and Russell 1969
Pereda-Suberbiola et al. 2000
3 forms (A, B,and C)
Reptilia - Avetheropoda - Troodontidae
cf. Euronychodon sp. Antunes and Sigogneau-Russell 1991
Pereda-Suberbiola et al. 2000
nomen dubium belonging to Troodontidae
Reptilia - Avetheropoda
Ornithomimosauria indet. Barsbold 1976
Pereda-Suberbiola et al. 2000
    = ? Ornithomimosauria indet. Barsbold 1976
Canudo and Ruiz-Omeñaca 2003
Reptilia
cf. Tarascosaurus sp. Le Loeuff and Buffetaut 1991
Le Loeuff and Buffetaut 1991 2 specimens
femora, MHNV
    = Neoceratosauria indet. Novas 1991
Pereda-Suberbiola et al. 2000
Reptilia - Abelisauridae
? Abelisauridae indet. Bonaparte and Novas 1985
Astibia et al. 1990
Reptilia - Nodosauridae
Nodosauridae indet. Marsh 1890
Astibia et al. 1990 1 specimen
Struthiosaurus sp. Bunzel 1870
Pereda-Suberbiola 1999
a partial skeleton, including the synsacrum, portions of the pelvic girdle and hindlimbs, a dorsal vertebra, dorsal rib fragments and a dermal scute
    = cf. Struthiosaurus austriacus Bunzel 1871
Le Loeuff and Buffetaut 1995
Reptilia - Hadrosauridae
Hadrosauridae indet. Cope 1869
Pereda-Suberbiola et al. 2000
MCNA 10510, tooth
Reptilia - Rhabdodontidae
Ornithopoda indet. Marsh 1881
Astibia et al. 1987
    = Rhabdodon priscus Matheron 1869
Astibia et al. 1990
    = Rhabdodon sp. Matheron 1869
Pereda-Suberbiola and Sanz 1999
four isolated teeth, three from the maxilla and one from the dentary, two dorsal vertebrae, four caudal vertebrae, one left humerus, left and right femora
Reptilia - Azhdarchidae
Azhdarchidae indet. Nessov 1984
Astibia et al. 1990 295 specimens
    = cf. Azhdarcho sp. Nessov 1981
Pereda-Suberbiola et al. 2000
Reptilia
Crocodylidae indet. Cuvier 1807
Astibia et al. 1990 52 specimens
may be Allodaposuchus
    = aff. Allodaposuchus sp. Nopcsa 1928
Pereda-Suberbiola et al. 2000
    = Allodaposuchus precedens Nopcsa 1928
Buscalioni et al. 2001
? Trematochampsidae indet. Buffetaut 1974
Astibia et al. 1990
    = Musturzabalsuchus buffetauti Buscalioni et al. 1997
Pereda-Suberbiola et al. 2000
Alligatoridae indet. Gray 1844
Astibia et al. 1987
    = Acynodon iberoccitanus Buscalioni et al. 1997
Pereda-Suberbiola et al. 2000
Ischyrochampsa sp. Vasse 1995
Pereda-Suberbiola et al. 2000
nomen dubium belonging to Neosuchia
Reptilia - Squamata - Iguanidae
Scincomorpha informal indet. B Camp 1923
Astibia et al. 1990
    = Iguanidae indet. Oppel 1811
Rage 1999
one fragment of maxilla with four teeth, two fragments of unidentified bones with two teeth, and two fragments of unidentified bone with one tooth
Reptilia - Squamata - Madtsoiidae
Madtsoia sp. Simpson 1933
Astibia et al. 1990
    = Madtsoia laurasiae Rage 1996
Rage 1999
recombined as Menarana laurasiae
8 more or less complete vertebrae, and perhaps 2 fragments
Aniliidae indet. Fitzinger 1826
Astibia et al. 1990
    = Herensugea caristiorum Rage 1996
Rage 1999
17 vertebrae, perhaps one poorly preserved vertebra and perhaps one caudal vertebra
Reptilia - Squamata
Lacertilia informal indet. 1 Owen 1842
Rage 1999
numerous remains (teeth)
Lacertilia informal indet. 2 Owen 1842
Rage 1999
teeth
Lacertilia informal indet. 3 Owen 1842
Rage 1999
teeth
Lacertilia indet. Owen 1842
Rage 1999
vertebrae
Scincomorpha informal indet. A Camp 1923
Astibia et al. 1990
    = Scincomorpha indet. Camp 1923
Rage 1999
two fragments of unidentified bones with two teeth, and one isolated tooth
Amphisbaenia indet. Gray 1844
Astibia et al. 1990
? Amphisbaenia indet. Gray 1844
Rage 1999
10 vertebrae
Mammalia
Eutheria indet. (Huxley 1880)
Gheerbrant and Astibia 1999
Mammalia - Zhelestidae
Lainodon informal new species
Gheerbrant and Astibia 2012
L1AT 18 (talonid of left M/3) and L1AT 13 and (talonid of right M/1 or M/2)
Lainodon orueetxebarriai n. gen., n. sp. Gheerbrant and Astibia 1994
Gheerbrant and Astibia 1994
? Labes sp. Pol et al. 1992
Gheerbrant and Astibia 1999
new species
    = Lainodon orueetxebarriai Gheerbrant and Astibia 1994
Gheerbrant and Astibia 2012
L1AT 17, talonid of left M/1 or M/2 (fig. 2); L1AT 4, disto-labial fragment of left upper molar; and possibly L1AT11, damaged right M/1 or M/2
Lainodon cf. orueetxebarriai Gheerbrant and Astibia 1994
Gheerbrant and Astibia 2012
L1AT 20 (distal part of right lower premolar)
Theria informal indet. A Parker and Haswell 1897
Astibia et al. 1990
may be Marsupialia
    = Lainodon sp. Gheerbrant and Astibia 1994
Gheerbrant and Astibia 1999
    = Lainodon ragei n. sp. Gheerbrant and Astibia 2012
Gheerbrant and Astibia 2012
L1AT 15 - type (right dP/5); referred specimens: L1AT 5: left dP/5; L1AT 15: right dP/5; L1AT 16: left dP5/; L1AT 8, left M/1 (trigonid); L1AT 9, damaged right upper molar (protocone)
Mammalia
Theria informal indet. B Parker and Haswell 1897
Astibia et al. 1990
see common names

Geography
Country:Spain State/province:Castilla y Leon County:Burgos
Coordinates: 42.7° North, 2.6° West (view map)
Paleocoordinates:33.0° North, 0.1° West
Basis of coordinate:estimated from map
Altitude:747 meters
Geographic resolution:outcrop
Time
Period:Cretaceous Epoch:Late/Upper Cretaceous
Stage:Campanian 10 m.y. bin:Cretaceous 7
*Period:Late/Upper Cretaceous *Epoch:Early/Lower Maastrichtian
Key time interval:Late/Upper Campanian
Age range of interval:83.50000 - 70.60000 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Formation:Vitoria
Local section:Lano section Local bed:S1U3
Local order:bottom to top
Stratigraphic resolution:bed
Stratigraphy comments: in a sector of the south limb of the Miranda-Trevino syncline, level is also called L1A
Lithology and environment
Primary lithology:"cross stratification",hardground conglomeratic sandstone
Secondary lithology:black siltstone
Includes fossils?Y
Includes fossils?Y
Lithology description: Multidirectional cross-laminated sands with microconglomerates in the bottom-sets, rarely more than 1m thick. Strata become massive toward the top, and are frequently intersected by abrasion surfaces showing an iron-mineralized crust or hard-ground. The most important bone accumulation is between two of these hardgrounds. The unit is topped by two black silt beds.
Environment:coastal indet.
Glacial or sequence phase:regressive
Geology comments: upper foreshore and backshore environments to a subcontinental, plant-rich perhaps swampy environment. The vertebrate bearing levels indicate a continental depositional environment affected by high-energy episodes, a fluviatile environment.
Taphonomy
Modes of preservation:body,permineralized,replaced with pyrite
Size of fossils:macrofossils,mesofossils,microfossils
Disassociated minor elements:all
Collection methods and comments
Collection excludes:difficult macrofossils
Collection methods:sieve,field collection
Collection size:33 specimens
Reason for describing collection:taxonomic analysis
Collection method comments: mostly chelonian shell fragments (30)
Metadata
Also known as:Laño Quarry
Database number:28406
Authorizer:J. Alroy, P. Mannion, M. Carrano Enterer:E. Leckey, P. Mannion, M. Carrano
Modifier:M. Carrano Research group:vertebrate
Collections that are a subset of this one:37199
Created:2003-02-12 10:11:45 Last modified:2019-09-25 09:43:39
Access level:the public Released:2003-02-12 10:11:45
Creative Commons license:CC BY
Reference information

Primary reference:

30625.ETE J. L. Sanz. 1986. Nouveaux gisements de dinosaures dans le Crétacé espagnol [New dinosaur localities in the Spanish Cretaceous]. Les Dinosaures de La Chine à La France, Colloque International de Paléontologie, Toulouse, France, 2-6 Septembre 1985; Muséum d'Histoire Naturelle de Toulouse, Toulouse 81-88 [M. Carrano/M. Carrano/M. Carrano]

Secondary references:

7903 H. Astibia, E. Buffetaut, A. D. Buscalioni, H. Cappetta, C. Corral, R. Estes, F. Garcia-Garmilla, J. J. Jaeger, E. Jiminez-Fuentes, J. Le Loeuff, J. M. Mazin, X. Orue-Etexebarria, J. Pereda-Suberbiola, J. E. Powell, J. C. Rage, J. Rodriguez-Lazaro, J. L. Sanz and H. Tong. 1990. The fossil vertebrates from the Lano (Basque Country, Spain); new evidence on the composition and affinities of the Late Cretaceous continental faunas of Europe. Terra Nova 2:460-466 [J. Alroy/E. Leckey/E. Leckey]
7885 H. Astibia, F. Garcia-Garmilla, X. Orue-Etxebarria and J. Rodriguez-Lazaro. 1987. The Cretaceous-Tertiary boundary in a sector of the south limb of the Miranda-Trevino synclinal: the first appearance of Chelonia and Archosauria in the Basque Country. Cretaceous Research 8:15-27 [J. Alroy/E. Leckey/E. Leckey]
14444ETE A. D. Buscalioni, F. Ortega, D. B. Weishampel and C. M. Jianu. 2001. A revision of the crocodyliform Allodaposuchus precedens from the Upper Cretaceous of the Hateg Basin, Romania. Its relevance in the phylogeny of Eusuchia. Journal of Vertebrate Paleontology 21(1):74-86 [M. Carrano/M. Carrano/M. Carrano]
17647ETE J. I. Canudo and J. I. Ruiz-Omeñaca. 2003. Los restos directos de dinosaurios teropódos (excluyendo Aves) en España [Direct remains of theropod dinosaurs (excluding Aves) in Spain]. Ciencias de la Tierra 26:347-373 [M. Carrano/M. Carrano]
32749ETE J. Company, X. Pereda Suberbiola, and J. I. Ruiz-Omenaca. 2009. Nuevos restos fosiles del dinosaurio Lirainosaurus (Sauropoda, Titanosauria) en el Cretacico Superior (Campaniano-Maastrichtiano) de la Peninsula Iberica. Ameghiniana 46(2):391-405 [M. Carrano/M. Carrano/M. Carrano]
19147ETE F. M. Dalla Vecchia. 2006. Telmatosaurus and the other hadrosaurids of the Cretaceous European Archipelago. An overview. Natura Nascosta 32:1-55 [M. Carrano/M. Carrano]
56054 F. M. Dalla Vecchia. 2014. An overview of the latest Cretaceous hadrosauroid record in Europe. In D. A. Eberth & D. C. Evans (ed.), Hadrosaurs 268-297 [M. Carrano/M. Carrano]
9511 F. de Lapparent and X. Murelaga. 1999. Turtles from the Upper Cretaceous of Lano (Iberian Peninsula). Estudios del Museo de Ciencias Naturales de Alava 14(1):135-211 [J. Alroy/E. Leckey/E. Leckey]
38464 V. Díez Díaz, X. Pereda Suberbiola, and J. L. Sanz. 2011. Braincase anatomy of the titanosaurian sauropod Lirainosaurus astibiae from the Late Cretaceous of the Iberian Peninsula. Acta Palaeontologica Polonica 56(3):521-533 [M. Carrano/M. Carrano]
43435 V. Díez Díaz, X. Pereda Suberbiola, and J. L. Sanz. 2012. Juvenile and adult teeth of the titanosaurian dinosaur Lirainosaurus (Sauropoda) from the Late Cretaceous of Iberia. Geobios 45:265-274 [M. Carrano/M. Carrano]
46308 V. Díez Díaz, X. Pereda Suberbiola, and J. L. Sanz. 2013. The axial skeleton of the titanosaur Lirainosaurus astibiae (Dinosauria: Sauropoda) from the latest Cretaceous of Spain. Cretaceous Research 43:145-160 [P. Mannion/J. Tennant/J. Tennant]
48052 V. Díez Díaz, X. Pereda Suberbiola, and J. L. Sanz. 2013. Appendicular skeleton and dermal armour of the Late Cretaceous titanosaur Lirainosaurus astibiae (Dinosauria: Sauropoda) from Spain. Palaeontologia Electronica 16(2):19A [P. Mannion/J. Tennant/P. Mannion]
9509 S. Duffaud and J.-C. Rage. 1999. Amphibians from the Upper Cretaceous of Lano (Basque Country, Spain). Estudios del Museo de Ciencias Naturales de Alava 14(1):111-120 [J. Alroy/E. Leckey/J. Head]
7914 E. Gheerbrant and H. Astibia. 1994. Un nouveau mammifere du Maastrichtien de Lano. Comptes Rendus de l'Academie des Sciences 318:1125-1131 [J. Alroy/E. Leckey/M. Uhen]
77915% 960E. Gheerbrant and H. Astibia. 1999. The Upper Cretaceous mammals from Lano (Spanish Basque Country). Estudios del Museo de Ciencias Naturales de Alava 14(1):295-323 [J. Alroy/E. Leckey/E. Leckey]
44341 E. Gheerbrant and H. Astibia. 2012. Addition to the Late Cretaceous Laño mammal faunule (Spain) and to the knowledge of European “Zhelestidae” (Lainodontinae nov.). Bulletin de la Société Géologique de France 183(6):537-546 [P. Mannion/P. Mannion]
62124 W. G. Joyce. 2017. A Review of the Fossil Record of Basal Mesozoic Turtles. Bulletin of the Peabody Museum of Natural History 58(1):65-113 [P. Holroyd/P. Holroyd/P. Mannion]
13022ETE J. Le Loeuff. 1993. European titanosaurids. Revue de Paléobiologie, Volume Spéciale 7:105-117 [M. Carrano/M. Carrano/M. Carrano]
36728ETE J. Le Loeuff. 1998. Les dinosaures du Crétacé supérieur de l'Île Ibéro-Armoricaine [The dinosaurs of the Upper Cretaceous of the Ibero-Armorican Island]. In A. M. G. de Carvalho, A. M. Andrade, V. F. dos Santos, J. Cascalho, R. Taborda, (eds.), I Encontro Internacional sobre Paleobiologia dos Dinossáurios. Museu Nacional de História Natural, Lisboa 49-64 [M. Carrano/M. Carrano]
7842 J. Le Loeuff and E. Buffetaut. 1991. Tarascosaurus salluvicus nov. gen., nov. sp., dinosaure théropode du Crétacé supérieur du sud de la France [Tarascosaurus salluvicus nov. gen., nov. sp., a theropod dinosaur from the Upper Cretaceous of southern France]. Géobios 25(5):585-594 [M. Carrano/M. Carrano/M. Carrano]
67934 J. Le Loeuff and E. Buffetaut. 1995. The evolution of Late Cretaceous non-marine vertebrate fauna in Europe. Sixth Symposium on Mesozoic Terrestrial Ecosystems and Biota, Short Papers 181-184 [M. Carrano/M. Carrano]
46669 A. Osi and E. Prondvai. 2013. Sympatry of two ankylosaurs (Hungarosaurus and cf. Struthiosaurus) in the Santonian of Hungary. Cretaceous Research 44:58-63 [P. Mannion/J. Tennant/P. Mannion]
9513 X. Pereda-Suberbiola. 1999. Ankylosaurian dinosaur remains from the Upper Cretaceous of Lano (Iberian Peninsula). Estudios del Museo de Ciencias Naturales de Alava 14(1):273-288 [J. Alroy/E. Leckey/E. Leckey]
9559 X. Pereda-Suberbiola, H. Asibia, X. Murelaga, J. J. Elzorza, and J. J. Gomez-Alday. 2000. Taphonomy of the Late Cretaceous dinosaur-bearing beds of the Lano Quarry (Iberian Peninsula). Palaeogeography, Palaeoclimatology, Palaeoecology 157:247-275 [J. Alroy/E. Leckey/E. Leckey]
14565ETE X. Pereda-Suberbiola, H. Astibia, and E. Buffetaut. 1995. New remains of the armoured dinosaur Struthiosaurus from the Late Cretaceous of the Iberian peninsula (Laño locality, Basque-Cantabric basin). Bulletin de la Société Géologique de France 166(2):207-211 [M. Carrano/M. Carrano/M. Carrano]
9512 X. Pereda-Suberbiola and J. Sanz. 1999. The ornithopod dinosaur Rhabdodon from the Upper Cretaceous of Lano (Iberian Peninsula). Estudios del Museo de Ciencias Naturales de Alava 14(1):257-272 [J. Alroy/E. Leckey/E. Leckey]
41499 A. Pérez-García, T. M. Scheyer, and X. Murelaga. 2012. New interpretations of Dortoka vasconica Lapparent de Broin and Murelaga, a freshwater turtle with an unusual carapace. Cretaceous Research 36:151-161 [R. Benson/R. Benson]
9510 J. Rage. 1999. Squamates (Reptilia) from the Upper Cretaceous of Lano (Basque Country, Spain). Estudios del Museo de Ciencias Naturales de Alava 14(1):121-133 [J. Alroy/E. Leckey/E. Leckey]
70303 J. L. Sanz. 2000. Yacimientos ibéricos de dinosaurios [Iberian dinosaur localities]. Dinosaurios: Los Señores del Pasado [Dinosaurs: The Lords of the Land] 267-299 [M. Carrano/M. Carrano]
13503ETE J. L. Sanz, J. E. Powell, J. Le Loeuff, R. Martinez, and X. Pereda-Suberbiola. 1999. Sauropod remains from the Upper Cretaceous of Laño (northcentral Spain). Titanosaur phylogenetic relationships. Estudios del Museo de Ciencias Naturales de Alava 14(especial 1):235-255 [M. Carrano/M. Carrano/M. Carrano]
19710ETE F. Torcida Fernández-Baldor. 2005. Los dinosaurios de Castilla y León [The dinosaurs of Castilla y León]. Patrimonio Histórico de Castilla y León 6(23):23-34 [M. Carrano/M. Carrano/M. Carrano]
55315 B. Vila, M. Suñer, A. Santos-Cubedo, J. I. Canudo, B. Poza and A. Galobart. 2011. Saurischians through time. In A. Galobart, M. Suñer, & B. Poza (eds.), Dinosaurs of Eastern Iberia 130-168 [M. Carrano/M. Carrano]