PBDB Collection

Lo Hueco, Fuentes (G1): Late/Upper Campanian - Early/Lower Maastrichtian, Spain
collected 2007

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

Gastropoda

	Gastropoda indet. Cuvier 1795	Barroso-Barcenilla et al. 2009
Bivalvia

	Bivalvia indet. Linnaeus 1758	Barroso-Barcenilla et al. 2009
Bivalvia - Unionida - Unionidae

	Unionidae indet. Rafinesque 1820	Barroso-Barcenilla et al. 2009
unclassified

	Pisces indet. Linnaeus 1758
	= Osteichthyes indet. Huxley 1880	Barroso-Barcenilla et al. 2009
Reptilia

	Scincomorpha indet. (Camp 1923)	Torices et al. 2010

	? Lacertoidea indet. (Camp 1923)	Torices et al. 2010

	Lepidosauromorpha indet. (Benton 1983)
	= ? Iguania indet. Cuvier 1817	Barroso-Barcenilla et al. 2009
	"probably Iguania indet."
Reptilia - Testudines

	Foxemydina indet. Gaffney et al. 2006	Pérez-García et al. 2013	1 specimen
	HUE-1220
	= Foxemys mechinorum Tong et al. 1998	Pérez-García and Ortega 2018
	Two unpublished specimens, corresponding to an almost complete and disarticulated carapace, preserving the articulate left ilium, HUE-8532 (Figs. 1, 2(A-H)), and an isolated nuchal plate, HUE-10218 (Fig. 2(I, J)), as well as an almost complete plastron, HUE-1220 (Fig. 2(K, L)), previously attributed to Foxemydina indet. by Pérez-García et al. (2013).

	Bothremydidae indet. Baur 1891		1 specimen
	= Elochelys convenarum Laurent et al. 2002	Pérez-García et al. 2010
	synonym of Iberoccitanemys atlanticum
	HUE-4913

	Testudines indet. Batsch 1788
	= Pancryptodira indet. Joyce et al. 2004	Barroso-Barcenilla et al. 2009
Reptilia

	Crocodylia indet. (Owen 1842)
	= Musturzabalsuchus sp. Buscalioni et al. 1997	Puértolas-Pascual et al. 2016

	Allodaposuchus sp. Nopcsa 1928
	= Lohuecosuchus megadontos n. gen., n. sp. Narváez et al. 2015	Narváez et al. 2015
	recombined as Allodaposuchus megadontos
	HUE-04498 - holotype (a complete skull and right lower jaw); paratypes: HUE-02920, a nearly complete skull lacking most of the pterygoids; HUE-04263, the palatal region of an incomplete skull which preserves most of its elements; HUE-04378, a left lower jaw; HUE-05161, a right lower jaw

	Agaresuchus fontisensis n. gen., n. sp. Narváez et al. 2016	Narváez et al. 2016
	recombined as Allodaposuchus fontisensis
	HUE-02502 - holotype (complete skull, right lower jaw and cervical vertebra)
Reptilia - Rhabdodontidae

	Ornithopoda indet. (Marsh 1881)
	= Rhabdodon sp. Matheron 1869	Barroso-Barcenilla et al. 2009
Reptilia

	? Ankylosauria indet. (Osborn 1923)	Barroso-Barcenilla et al. 2009

	Neotheropoda indet. Bakker 1986	Baiano et al. 2016	1 specimen
	metatarsal IV (HUE-01818)
Reptilia - Dromaeosauridae

	? Velociraptorinae indet. Barsbold 1983	Torices et al. 2008	1 specimen
	tooth type 3
	= cf. Richardoestesia sp. Currie et al. 1990	Torices et al. 2010

	Velociraptorinae indet. Barsbold 1983	Barroso-Barcenilla et al. 2009	12 specimens
	tooth type 2; HUE-02696 (tibia + proximal tarsals)

	Dromaeosauridae indet. Colbert and Russell 1969		6 specimens
	= Dromaeosaurinae indet. Matthew and Brown 1922	Torices et al. 2008
	tooth type 1
Reptilia

	Titanosauria informal indet. 2 Bonaparte and Coria 1993	Barroso-Barcenilla et al. 2009	1 individual
	HUE-EC-13 (humerus HUE-1647, femur HUE-1183, tibia and fibula HUE-1612)

	Titanosauria informal indet. A Bonaparte and Coria 1993	Díez Díaz et al. 2014	13 specimens
	morphotype A teeth; HUE-30, 90, 685, 702, 1125, 1212, 1636, 3793, 4723, 6155, 6809, 8013, 8743

	Titanosauria informal indet. B Bonaparte and Coria 1993	Díez Díaz et al. 2014	9 specimens
	morphotype B teeth; HUE-540, 2687, 3891, 4040, 4148, 4893, 5575, 6942, 7041

	Titanosauria indet. Bonaparte and Coria 1993	Díez Díaz et al. 2014	3 specimens
	unassignable teeth; HUE-5498, 5809, 6889

	Ampelosaurus sp. Le Loeuff 1995	Knoll et al. 2013	1 specimen
	MCCM-HUE-8741 (braincase)

	Titanosauria indet. Bonaparte and Coria 1993
	three forms
	= Titanosauria informal indet. 1 Bonaparte and Coria 1993	Barroso-Barcenilla et al. 2009
	= Lohuecotitan pandafilandi n. gen., n. sp. Díez Díaz et al. 2016	Díez Díaz et al. 2016
	HUE-EC-01 - holotype
Amphibia - Temnospondyli

	Lissamphibia indet. (Haeckel 1866)	Torices et al. 2010
Actinopterygii

	Actinopterygii indet. Cope 1887	Barroso-Barcenilla et al. 2009
Actinopteri - Amiiformes - Amiidae

	Amiidae indet. Huxley 1861	Torices et al. 2010
Actinopteri - Lepisosteiformes - Lepisosteidae

	Lepisosteidae indet. Cuvier 1825	Barroso-Barcenilla et al. 2009

	cf. Lepisosteidae indet. Cuvier 1825	Torices et al. 2010	1 specimen
	atlas
Actinopteri

	Pycnodontoidea indet.	Torices et al. 2010
Actinopteri - Albuliformes - Albulidae

	Albulidae indet. Bleeker 1859	Torices et al. 2010
unclassified

	Plantae indet. (Haeckel 1866)	Barroso-Barcenilla et al. 2009
see common names

Geography

Country:	Spain	State/province:	Castilla-La Mancha	County:	Cuenca
Coordinates:	40.0° North, 2.0° West (view map)
Paleocoordinates:	30.7° North, 0.6° West
Basis of coordinate:	based on nearby landmark
Geographic resolution:	outcrop

Time

Period:	Cretaceous	Epoch:	Late/Upper Cretaceous

Key time interval:	Late/Upper Campanian - Early/Lower Maastrichtian
Age range of interval:	83.60000 - 66.00000 m.y. ago

Stratigraphy

Formation:	Villalba de la Sierra
Local section:	Lo Hueco	Local bed:	3 m
Local order:	bottom to top
Stratigraphic resolution:	bed
Stratigraphy comments: "Garumnian", upper part of formation

Lithology and environment

Primary lithology:	planar lamination,gypsiferous,gray argillaceous mudstone
Secondary lithology:	sandstone
Includes fossils?	Y
Lithology description: marly mudstone with high proportion of clay minerals (> 85%), moderate gypsum (< 15%), and low calcite (5-10%). Mainly planar or parallel, locally wavy and massive bedding. Some thin sandy interbeds.
Environment:	wet floodplain
Geology comments: "proximal flooded muddy plain", "close to the distributary channels"

Taphonomy

Modes of preservation:	body
Degree of concentration:	concentrated
Size of fossils:	macrofossils,mesofossils
Spatial orientation:	preferred
Preservation of anatomical detail:	good
Abundance in sediment:	common
Associated major elements:	many
Disassociated major elements:	many
Disassociated minor elements:	many
Size sorting:	medium
Fragmentation:	occasional
Spatial resolution:	parautochthonous

Collection methods and comments

Collection methods:	bulk,selective quarrying,surface (in situ),mechanical,sieve,field collection
Reason for describing collection:	general faunal/floral analysis
Collection dates:	May 2007

Metadata

Database number:	91781
Authorizer:	M. Carrano, P. Mannion, P. Holroyd, E. Vlachos	Enterer:	M. Carrano, P. Mannion, J. Tennant, E. Vlachos, P. Holroyd
Modifier:	P. Mannion	Research group:	vertebrate
Created:	2009-10-19 20:19:54	Last modified:	2023-03-02 08:55:46
Access level:	the public	Released:	2009-10-19 20:19:54
Creative Commons license:	CC BY

Reference information

Primary reference:

30927.

ETE

F. Ortega, J. L. Sanz, F. Barroso-Barcemilla, O. Cambra-Moo, F. Escaso, M. García-Oliva, F. Marcos Fernández, A. Pérez García, M. Segura and A. Torices. 2008. Macrovertebrados del Cretácico Superior de "Lo Hueco" (Fuentes, Cuenca) [Macrovertebrates from the Upper Cretaceous of "Lo Hueco" (Fuentes, Cuenca)]. In J. I. Ruiz-Omeñaca, L. Piñuela and J. C. García-Ramos (eds), XXIV Jornadas de la Sociedad Española de Paleontología, 15-18 October 2008, Museo del Jurásico de Asturias (MUJA), Colunga, Spain, Libro de Resúmenes 165-166 [M. Carrano/M. Carrano]

Secondary references:

83613		M. Aurell, D. Torromé, J. M. Gasca, P. Calvín, M. Pérez-Pueyo, J. Parrilla-Bel, E. Medrano-Aguado, C. Martín-Closas, A. Vicente, P. Sierra-Campos, and J. I. Canudo. 2022. Latest Cretaceous palaeogeographic evolution of northeast Iberia: Insights from the Campanian continental Montalbán subbasin (Spain). Earth-Science Reviews 235:10425 [M. Carrano/M. Carrano/P. Mannion]
76316		M. A. Baiano, F. Ortega-Coloma, and R. A. Coria. 2016. Un metatarsiano de un Neotheropoda basal del yacimiento de Lo Hueco, Formación “Margas, Arcillas y Yesos de Villalba de la Sierra” (Campaniano-Maastrichtiano, Cretácico Superior, Cuenca, España) [A metatarsal from a basal Neotheropoda from the Lo Hueco locality, “Margas, Arcillas y Yesos de Villalba de la Sierra” Formation (Campanian–Maastrichtian, Upper Cretaceous, Cuenca, Spain)]. 53(6 (suppl.)):6 [M. Carrano/M. Carrano]
32222	ETE	F. Barroso-Barcenilla, O. Cambra-Moo, F. Escaso, F. Ortega, A. Pascual, A. Pérez-García, J. Rodríguez-Lázaro, J. L. Sanz, M. Segura and A. Torices. 2009. New and exceptional discovery in the Upper Cretaceous of the Iberian Peninsula: the palaeontological site of ‘‘Lo Hueco’’, Cuenca, Spain. Cretaceous Research 30:1368-1378 [M. Carrano/M. Carrano]
41899		O. Cambra-Moo, F. Barroso-Barcenilla, A. Berreteaga, B. Carenas, F. Coruña, L. Domingo, M. S. Domingo, A. Elvira, F. Escaso, F. Ortega, A. Pérez-García, D. Peyrot, J. L. Sanz, M. Segura, A. Sopelana and A. Torices. 2012. Preliminary taphonomic approach to ‘‘Lo Hueco’’ palaeontological site (Upper Cretaceous, Cuenca, Spain). Geobios 45:157-166 [M. Carrano/M. Carrano]
74890		V. Díez Díaz, G. Garcia, X. Pereda Suberbiola, B. Jentgen-Ceschino, K. Stein, P. Godefroit, and X. Valentin. 2021. A new titanosaur (Dinosauria: Sauropoda) from the Upper Cretaceous of Velaux-La-Bastide Neuve (southern France). Historical Biology 33(11):2998-3017 [P. Mannion/P. Mannion/M. Carrano]
60088		V. Díez Díaz, P. Mocho, A. Páramo, F. Escaso, F. Marcos-Fernández, J. L. Sanz, and F. Ortega. 2016. A new titanosaur (Dinosauria, Sauropoda) from the Upper Cretaceous of Lo Hueco (Cuenca, Spain). Cretaceous Research 68:49-60 [P. Mannion/P. Mannion/P. Mannion]
79650		V. Díez Díaz, F. Ortega, and J. L. Sanz. 2014. Titanosaurian teeth from the Upper Cretaceous of “Lo Hueco” (Cuenca, Spain). Cretaceous Research 51:285-291 [M. Carrano/M. Carrano]
76794		V. Díez Díaz and X. Pereda Suberbiola. 2010. Titanosaurian fossil record of Europe: a review. 8th Meeting of the European Association of Vertebrate Palaeontologists. Abstract Volume 30 [M. Carrano/M. Carrano/M. Carrano]
83617		F. Escaso, R. A. Coria, and F. Ortega. 2021. Elementos del basicráneo de un orntópod del yacimiento Cretácico superior de Lo Hueco (Fuentes, Cuenca) [Elements of the basicranium of an ornithopod from the Upper Cretaceous Lo Hueco locality (Fuentes, Cuenca)]. Lucas Mallada 23:167-168 [M. Carrano/M. Carrano]
76434		F. Gascó, F. Ortega, D. Vidal, A. Paramo, P. Mocho and J. L. Sanz. 2016. Preliminary approach to the palaeohistology of the titanosaur sauropods from the Upper Cretaceous of Lo Hueco (Cuenca, Spain). In F. Torcida Fernández‐Baldor, J. I. Canudo, P. Huerta, & X. Pereda (eds.), VII Jornadas Internacionales sobre Paleontología de Dinosaurios y su Entorno, Libro de Resúmenes 73-74 [M. Carrano/M. Carrano]
80599		E. Isasmendi, A. Torices, J. I. Canudo, P. J. Currie, and X. Pereda-Suberbiola. 2022. Upper Cretaceous European theropod palaeobiodiversity, palaeobiogeography and the intra‐Maastrichtian faunal turnover: new contributions from the Iberian fossil site of Laño. Papers in Palaeontology 8(1):e1419:1-38 [M. Carrano/M. Carrano]
44421		F. Knoll, R. C. Ridgely, F. Ortega, J. L. Sanz, and L. M. Witmer. 2013. Neurocranial osteology and neuroanatomy of a Late Cretaceous titanosaurian sauropod from Spain (Ampelosaurus sp.). PLoS ONE 8(1):e54991:1-11 [P. Mannion/P. Mannion/M. Carrano]
83916		E. Malafaia, F. Escaso, R. A. Coria and F. Ortega. 2023. An eudromaeosaurian theropod from Lo Hueco (Upper Cretaceous. Central Spain). Diversity 15(2):141 [P. Mannion/P. Mannion/M. Carrano]
56860		I. Narváez, C. A. Brochu, F. Escaso, A. Pérez-García, and F. Ortega. 2015. New Crocodyliforms from Southwestern Europe and Definition of a Diverse Clade of European Late Cretaceous Basal Eusuchians. PLoS ONE 10(11):e0140679:1-34 [P. Mannion/P. Mannion]
59221		I. Narváez, C. A. Brochu, F. Escaso, A. Perez-García, and F. Ortega. 2016. New Spanish Late Cretaceous eusuchian reveals the synchronic and sympatric presence of two allodaposuchids. Cretaceous Research 65:112-125 [P. Mannion/P. Mannion/P. Mannion]
81290		F. Ortega. 2013. Towards the last European ecosystems with dinosaurs: the Upper Cretaceous biota of Lo Hueco (Fuentes, Cuenca, Spain). In F. Torcida Fernández-Baldor & P. Huerta (ed.), VI Jornadas Internacionales sobre Paleontología de Dinosaurios y su Entorno. Libro de Resúmenes 27-29 [M. Carrano/M. Carrano]
75861		A. Páramo, P. Mocho, and F. Ortega. 2020. Three‑dimensional analysis of the titanosaurian limb skeleton: implications for systematic analysis. Journal of Iberian Geology 46:369-402 [M. Carrano/M. Carrano]
76438		A. Páramo, F. Ortega, P. Mocho and J. L. Sanz. 2016. Femoral variability in two titanosaur taxa from Lo Hueco (Cuenca, Spain): Insights for iberoarmorican titanosaur diversity assessment. In F. Torcida Fernández‐Baldor, J. I. Canudo, P. Huerta, & X. Pereda (eds.), VII Jornadas Internacionales sobre Paleontología de Dinosaurios y su Entorno, Libro de Resúmenes 107-108 [M. Carrano/M. Carrano]
67929		J. Párraga and A. Prieto-Márquez. 2019. Pareisactus evrostos, a new basal iguanodontian (Dinosauria: Ornithopoda) from the Upper Cretaceous of southwestern Europe. Zootaxa 4555(2):247-258 [M. Carrano/M. Carrano]
46020		A. Pérez-García, F. Ortega, and X. Murelaga. 2010. Ampliación de la distribución geográfica y temporal de Elochelys convenarum (Chelonii, Bothremydidae) en el Cretácico Superior de la Península Ibérica. Ameghiniana 47(3):307-315 [P. Mannion/J. Tennant/P. Mannion]
59664		A. Pérez-García, F. Ortega, and X. Murelaga. 2013. Two Synchronic and Sympatric Bothremydidae Taxa (Chelonii, Pleurodira) in the Late Cretaceous Site of ‘‘Lo Hueco’’ (Cuenca, Spain). In D. B. Brinkman, P. A. Holroyd, and J. D. Gardner (eds.), Morphology and Evolution of Turtles 251-259 [P. Holroyd/P. Holroyd]
65471		A. Pérez-García and F. Ortega. 2018. Identification of the French Upper Cretaceous bothremydid turtle Foxemys mechinorum in the Spanish record. Geobios 51(3):211-217 [E. Vlachos/E. Vlachos/P. Mannion]
55576		J. L. Sanz, F. Ortega, and M. Shibata. 2014. Dinosaurios Maravillosos de España [Marvelous Dinosaurs from Spain] 1-155 [M. Carrano/M. Carrano]
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76803		A. Torices, E. Díaz Berenguer, I. Narvaez, F. Ortega, S. Perez and H. Serrano. 2010. Preliminary analisys of the microvertebrate fossils of "Lo Hueco" (Upper Cretaceous, Cuenca, Spain). 8th Meeting of the European Association of Vertebrate Palaeontologists. Abstract Volume 81 [M. Carrano/M. Carrano]
52266		D. Vidal, F. Ortega, and J. L. Sanz. 2014. Titanosaur osteoderms from the Upper Cretaceous of Lo Hueco (Spain) and their implications on the armor of Laurasian titanosaurs. PLoS ONE 9(8):e102488:1-12 [M. Carrano/M. Carrano]
55315		B. Vila, M. Suñer, A. Santos-Cubedo, J. I. Canudo, B. Poza and A. Galobart. 2011. Saurischians through time. In A. Galobart, M. Suñer, & B. Poza (eds.), Dinosaurs of Eastern Iberia 130-168 [M. Carrano/M. Carrano]