El Kohol: Ypresian, Algeria
collected 1980, 1990

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Charales - Feistiellaceae
Nitellopsis dutemplei
Mahboubi et al. 1984
"N. (T.) dutemplei"
Nitellopsis aff. thaleri
Mahboubi et al. 1984
"N. (T.) aff. thaleri"
Nitellopsis (Tectochara) dutemplei
Nitellopsis (Tectochara) aff. thaleri
Angiospermae - Rosales - Ulmaceae
Celtis sp. Linnaeus 1753
Dipnomorpha - Lepidosirenidae
Protopterus sp. Agassiz 1843
Reptilia
Alethinophidia indet. (Nopcsa 1923)
Aves indet. (Linnaeus 1758)
Mesosuchia indet. Huxley 1875
Buffetaut 1982
"ziphodont mesosuchian"
    = Eremosuchus elkoholicus n. gen., n. sp. Buffetaut 1989
Buffetaut 1989
KB-301 - holotype (almost complete right dentary); hypodigm: KA-401 (posterior part of left mandibular ramus); KA-117, 118, 402-404 (teeth); KA-405 & 406 (thoracic vertebrae); KA-407 & 408 (caudal vertebrae), and KA-114 (fibula)
Mammalia - Creodonta
Creodonta indet. (Cope 1875)
Mammalia - Creodonta - Hyaenodontidae
Koholia atlasense Crochet 1988
Crochet 1988
Mammalia
Lipotyphla indet. (Haeckel 1866)
synonym of Eulipotyphla
Mammalia - Proboscidea - Numidotheriidae
Proboscidea indet. Illiger 1811
Mahboubi et al. 1984
new genus
    = Numidotherium koholense n. gen., n. sp. Mahboubi et al. 1986
Court and Mahboubi 1993
Mammalia - Hyracoidea
Hyracoidea indet. (Huxley 1869)
Mammalia - Hyracoidea - Pliohyracidae
Seggeurius amourensis n. gen., n. sp. Crochet 1986
Court and Mahboubi 1993
Mammalia - Adapisoriculidae
Garatherium mahboubii n. gen., n. sp. Crochet 1984
Crochet 1984
Mammalia - Alphadontidae
aff. Alphadon sp. Simpson 1927
Amphibia - Temnospondyli
Neobatrachia indet. (Reig 1958)
Actinopteri - Characiformes - Characidae
Characidae indet.
Ostracoda - Podocopida - Candonidae
? Candona sp. Baird 1845
Ostracoda - Podocopida - Cytherideidae
Cytherideidae indet. Sars 1925
Gastropoda - Naticidae
Ampullaria cf. assermoensis
spelled with current rank as Natica (Ampullaria)
Gastropoda - Hydrobiidae
Hydrobiidae indet. Troschel 1857
Gastropoda - Thiaridae
Melanoides sp. Olivier 1804
n. sp.
see common names

Geography
Country:Algeria
Coordinates: 33.1° North, 1.3° East (view map)
Paleocoordinates:26.1° North, 0.9° West
Basis of coordinate:based on nearby landmark
Time
Period:Paleogene Epoch:Eocene
Stage:Ypresian 10 m.y. bin:Cenozoic 2
Key time interval:Ypresian
Age range of interval:56.00000 - 47.80000 m.y. ago
Stratigraphy
Stratigraphy comments: "Cuisian" or "late early Eocene" based on charophyte gryogonites (Mahboubi et al. 1984); Ypresian age after Coster et al (2012)
Lithology and environment
Primary lithology: marl
Lithology description: "lacustrine marl" (Court and Mahboubi 1993)
Environment:lacustrine indet.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Collection methods and comments
Reason for describing collection:taxonomic analysis
Collection dates:1980s, 1990
Collection method comments: material belongs to the Université d'Oran
Taxonomic list comments:fauna includes "a lungfish, a crocodilian, and various mammals (hyracoids and a form related to Barytherium)" according to Buffeteaut 1982
Metadata
Also known as:Brezina
Database number:71984
Authorizer:J. Alroy, M. Uhen, P. Coster, P. Mannion, P. Holroyd Enterer:J. Alroy, P. Coster, P. Mannion, P. Holroyd, M. Uhen
Modifier:P. Mannion Research group:vertebrate
Created:2007-05-18 21:14:55 Last modified:2016-06-30 06:58:15
Access level:the public Released:2007-05-18 21:14:55
Creative Commons license:CC BY
Reference information

Primary reference:
43794. M. Mahboubi, R. Ameur, J.-Y. Crochet and J.-J. Jaeger. 1984. Implications Paleobiogeographiques de la decouverte d'une nouvelle localite Eocene a vertebres continentaux en Afrique Nord-Occidentale: El Kohol (Sud-Oranis, Algerie). Geobios 17(5):625-629 [M. Uhen/M. Uhen/P. Wagner]

Secondary references:

59620 J. Benoit, J.-V. Crochet, M. Mahboubi, J.-J. Jaeger, M. Bensalah, M. Adaci, and R. Tabuce. 2016. New material of Seggeurius amourensis (Paenungulata, Hyracoidea), including a partial skull with intact basicranium. Journal of Vertebrate Paleontology 36(1):e1034358:1-18 [P. Mannion/P. Mannion/P. Mannion]
24347 E. Buffetaut. 1982. A ziphodont mesosuchian crocodile from the Eocene of Algeria and its implications for vertebrate dispersal. Nature 300:176-1786 [J. Alroy/J. Alroy]
46912 E. Buffetaut. 1989. A new ziphodont mesosuchian crocodile from the Eocene of Algeria. Palaeontographica 208:1-10 [P. Mannion/P. Mannion]
47472 P. Coster, M. Benammi, M. Mahboubi, R. Tabuce, M. Adaci, L. Marivaux, M. Bensalah, S. Mahoubi, A. Mhboubi, F. Mebrouk, C. Mameri and J.-J. Jaeger. 2012. Chronology of the Eocene continental deposits of Africa: Magnetostratigraphy and biostratigraphy of the El Kohol and Glib Zegdou Formations, Algeria. GSA Bulletin 124(9/10):1590-1606 [M. Uhen/M. Uhen]
29524 N. Court and M. Mahboubi. 1993. Reassessment of Lower Eocene Seggeurius amourensis: aspects of primitive dental morphology in the mammalian order Hyracoidea. Journal of Paleontology 67(5):889-893 [J. Alroy/J. Alroy/J. Alroy]
57485 J. Y. Crochet. 1984. Garatherium mahboubii nov. gen., nov. sp., marsupial de l'Eocene inférieur d'El Kohol (Sud-Oranais, Algérie). Annales de Paléontologie 70:275-294 [P. Coster/P. Coster/P. Mannion]
57658 J. Y. Crochet. 1988. Le plus ancien Créodonte africain: Koholia atlasense nov. gen., nov. sp. (Eocène inférieur d’El Kohol, Atlas saharien, Algérie). Comptes Rendus de l’Académie des Sciences, Paris 307:1795-1798 [P. Coster/P. Coster/M. Uhen]
62975 G. Cuny, J.-J. Jaeger, M. Mahboubi and J.-C. Rage. 1990. Les plus anciens serpents (Reptilia, Squamata) connus. Mise au point sur l'âge géologique des serpents de la partie moyenne du Crétacé. Comptes rendus des séances de l'Académie des Sciences 311:1267-1272 [P. Barrett/T. Cleary]
24346 M. Mahboubi, R. Ameur, J. Y. Crochet and J. J. Jaeger. 1984. Earliest known proboscidean from the early Eocene of north-west Africa. Nature 308:543-544 [J. Alroy/J. Alroy]