Shestakovo-3: Barremian - Aptian, Russian Federation
collected 1995–2012, 2014–

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Tritylodontidae
Tritylodontidae indet. Cope 1884
Podobina et al. 2002
Mammalia - Amphidontidae
Amphidontidae indet. Simpson 1925
Leshchinskiy et al. 2000 1 individual
"genus et sp. nov."
Mammalia - Zhangheotheriidae
Kiyatherium cardiodens n. gen., n. sp. Maschenko and Lopatin 2002
Maschenko and Lopatin 2002 1 specimen
PM TGU no. 16/2-50, holotype maxilla
Mammalia - Eutriconodonta - Gobiconodontidae
Gobiconodon sp. Trofimov 1978
Leshchinskiy et al. 2000
Gobiconodon hoburensis (Trofimov 1978)
Podobina et al. 2002
Gobiconodon borissiaki Trofimov 1978
Podobina et al. 2002
Reptilia - Shartegosuchidae
Kyasuchus saevi n. gen., n. sp. Efimov and Leschinskii 1998
Efimov and Leshchinskiy 2000 2 specimens
Reptilia
Tagarosuchus kulemzini Alifanov et al. 1999
Podobina et al. 2002 1 individual
Psittacosaurus sp. Osborn 1923
Averianov et al. 2004
similar in tooth morphology to P. xinjiangensis
Psittacosaurus sibiricus n. sp. Voronkevich and Averianov 2000
Leshchinskiy et al. 2000 14 individuals
PM TSU 120-Sh3-125, 16/0-54 to 56; KOKM 4652/1. 9–11, 13, 16;plus hundreds of isolated specimens
Reptilia - Prismatoolithidae
Prismatoolithus ilekensis n. sp. Skutschas et al. 2017
Skutschas et al. 2017 1 specimen
LMCCE 4/6
Reptilia
Coelurosauria informal indet. A (Huene 1914)
Averianov et al. 2019 5 specimens
LMCCE 006/1 (Dromaeosauridae or Tyrannosauroidea)
Coelurosauria informal indet. D (Huene 1914)
Averianov et al. 2019 1 specimen
LMCCE 006/6 (Dromaeosauridae or Tyrannosauroidea)
Reptilia - Troodontidae
Troodontidae indet. Gilmore 1924
Leshchinskiy et al. 2000 1 individual
Reptilia
Evgenavis nobilis n. gen., n. sp. O'Connor et al. 2014
O'Connor et al. 2014 1 specimen
ZIN PH 1/154, an isolated right tarsometatarsus missing only the proximal half of metatarsal IV.
Sibirotitan astorsacralis
Averianov et al. 2023
KOKM A - 5142 and 5200
see common names

Geography
Country:Russian Federation State/province:Kemerovo County:Chebulinsky
Coordinates: 55.9° North, 88.0° East (view map)
Paleocoordinates:54.2° North, 84.7° East
Basis of coordinate:stated in text
Altitude:154 meters
Geographic resolution:outcrop
Time
Period:Cretaceous Epoch:Early/Lower Cretaceous
Key time interval:Barremian - Aptian
Age range of interval:129.40000 - 113.00000 m.y. ago
Stratigraphy
Formation:Ilek
Stratigraphic resolution:group of beds
Stratigraphy comments: "Ilekskaya Suite"; several bone-bearing layers within the main "Layer 1"

Skutschas et al. 2017: The dating of the Shestakovo 3 locality within the Lower Cretaceous remains controversial: it has been dated as Berriasian (Alifanov et al. 1999), Aptian–Albian (Averianov et al. 2006), and Barremian–Aptian (Kurochkin et al. 2011) based on vertebrates.

Age of Ilek Formation is poorly constrained but is currently believed to be Barremian-Aptian (see Averianov et al. 2018). The fossiliferous layer at Shestakovo 1 locality was dated to Aptian (Bugadeva et al. 2017). The overlying Kiya Formation has been assigned Albian age based on plant remains (Golovneva and Shchepetov 2010).
Lithology and environment
Primary lithology:concretionary,gray,green,yellow calcareous sandstone
Secondary lithology:planar lamination,brown,gray,green,red siltstone
Lithology description: "greenish-grey and yellowish-green sands, with calcareous sandstone concretions, interbedded rhythmically with packets of thin-laminated greenish-grey and brown-red siltstones and clay marls."
Environment:fluvial-lacustrine indet.
Geology comments: Skutschas et al. 2017: The Ilek Formation was deposited under continental environments in the fluvial-lacustrine semi-desert plain (Nesterov 1976).
Taphonomy
Modes of preservation:body
Degree of concentration:-single event
Size of fossils:macrofossils
Preservation of anatomical detail:good
Articulated whole bodies:some
Associated major elements:some
Disassociated major elements:some
Spatial resolution:parautochthonous
Collection methods and comments
Collection methods:selective quarrying,mechanical,field collection
Reason for describing collection:taxonomic analysis
Collection dates:1995–2012, 2014–
Metadata
Also known as:LMCCE 006
Database number:52808
Authorizer:M. Carrano, P. Mannion Enterer:M. Carrano, J. Tennant, P. Mannion
Modifier:M. Carrano Research group:vertebrate
Created:2005-08-22 07:26:00 Last modified:2023-11-14 15:58:25
Access level:the public Released:2005-08-22 07:26:00
Creative Commons license:CC BY
Reference information

Primary reference:

36988. E. N. Maschenko and A. V. Lopatin. 1998. First record of an Early Cretaceous triconodont mammal from Siberia. Bulletin de l'Institute Royal des Sciences Naturelles de Belgique, Sciences de la Terre 68:233-236 [M. Carrano/M. Carrano]

Secondary references:

71017 A. O. Averianov, S. V. Ivantsov, and P. P. Skutschas. 2019. Theropod teeth from the Lower Cretaceous Ilek Formation of Western Siberia, Russia. Proceedings of the Zoological Institute, Russian Academy of Sciences 323(2):65-84 [M. Carrano/M. Carrano]
14367ETE A. O. Averianov, S. Leshchinskiy, P. Skutschas, A. Fayngertz, and A. Rezvyi. 2004. Dinosaurs from the Early Cretaceous Ilek Formation in West Siberia, Russia. Second European Association of Vertebrate Palaeontologists Meeting, Moravian Museum, Brno. Abstracts of Papers 6 [M. Carrano/M. Carrano/M. Carrano]
86461 A. O. Averianov and A. V. Lopatin. 2023. Dinosaurs of Russia: a review of the localities. Herald of the Russian Academy of Sciences 93(4) [M. Carrano/M. Carrano/M. Carrano]
87538 A. O. Averianov, A. Podlesnov, D. Slobodin, P. Skutschas, O. Feofanova and O. Vladimirova. 2023. First sauropod dinosaur remains from the Early Cretaceous Shestakovo 3 locality, Western Siberia, Russia. Biological Communications 68(4):236-252 [P. Mannion/P. Mannion]
71292 A. O. Averianov, P. P. Skutschas, R. Schellhorn, A. V. Lopatin, P. N. Kolosov, V. V. Kolchanov, D. D. Vitenko, D. V. Grigoriev, and T. Martin. 2019. The northernmost sauropod record in the Northern Hemisphere. Lethaia 53:362-368 [P. Mannion/P. Mannion/M. Carrano]
19092ETE A. O. Averianov, A. V. Voronkevich, S. V. Lechchinskiy and A. Fayngertz. 2006. A ceratopsian dinosaur Psittacosaurus sibiricus from the Early Cretaceous of west Siberia, Russia and its phylogenetic relationships. Journal of Systematic Palaeontology 4(4):359-395 [M. Carrano/M. Carrano]
86243 N. V. Demidenko and E. N. Maschenko. 2016. Results of paleontological excavations 2014–2015 held by the Kemerovo Regional Museum at the Early Cretaceous Shestakovo vertebrate localities. In O. S. Dzyuba, E. B. Pestchevitskaya, & B. N. Shurygin (eds.), Cretaceous Ecosystems and Their Responses to Paleoenvironmental Changes in Asia and the Western Pacific. Short papers for the Fourth International Symposium of International Geoscience Programme, IGCP Project 608 114-116 [M. Carrano/M. Carrano]
43810 M. B. Efimov and S. B. Leschinskii. 1998. Pervaya nakhodka cherepa iskopaemogo krokodila v Siberi [The first finding of a fossil crocodile skull in Siberia]. Paleontologiya i Stratigrafiya 3:361-363 [P. Mannion/J. Tennant/J. Tennant]
14468ETE M. B. Efimov and S. V. Leshchinskiy. 2000. [First discovery of a fossil crocodile skull in Siberia]. In A.V. Komarov (ed.), Materialy regionalanoj konferencii geologov Sibiri, Dalshchnego Vostoka i Severoâ ’Vostoka Rossii. Gala Press, Tomsk 2:361-363 [M. Carrano/M. Carrano/M. Carrano]
86245 S. V. Ivantsov, A. V. Fayngerts, S.V. Leshchinskiy, A.S. Rezviy, P.P. Skutschas and A.O. Averianov. 2016. The Lower Cretaceous continental vertebrate fauna from the Shestakovo locality (West Siberia): Results of 20-year research. In O. S. Dzyuba, E. B. Pestchevitskaya, & B. N. Shurygin (eds.), Cretaceous Ecosystems and Their Responses to Paleoenvironmental Changes in Asia and the Western Pacific. Short papers for the Fourth International Symposium of International Geoscience Programme, IGCP Project 608 121-123 [M. Carrano/M. Carrano]
86246 A. E. Kostyunin. 2016. The history of discovery and research of the Shestakovo locality of Early Cretaceous vertebrates. In O. S. Dzyuba, E. B. Pestchevitskaya, & B. N. Shurygin (eds.), Cretaceous Ecosystems and Their Responses to Paleoenvironmental Changes in Asia and the Western Pacific. Short papers for the Fourth International Symposium of International Geoscience Programme, IGCP Project 608 124-126 [M. Carrano/M. Carrano]
38196 E. N. Kurochkin, N. V. Zelenkov, A. O. Averianov and S. V. Leshchinskiy. 2011. A new taxon of birds (Aves) from the Early Cretaceous of Western Siberia, Russia. Journal of Systematic Palaeontology 9(1):109-117 [P. Mannion/P. Mannion]
40059 S. V. Leshchinskiy, A. V. Faingerts, A. V. Voronkevich, E. N. Mashchensko, and A. O. Averianov. 2000. Predvaritel'nye rezul'taty izucheniy mestonakhozhdenii Shestakovskogo kompleksa rannemelovykh pozvonochnykh [Preliminary results of the investigation of the Shestakovo localities of Early Cretaceous vertebrates]. In A.V. Komarov (ed.), Materialy Regional'noy Konferentsii Geologov Sibiri Dal'nego Vostoka i Severo-Vostoka Rossii. Gala Press, Tomsk 2:363-366 [M. Carrano/M. Carrano/M. Carrano]
17882ETE A. V. Lopatin and A. O. Averianov. 2006. Mesozoic mammals of Russia. 9th International Symposium on Mesozoic Terrestrial Ecosystems and Biota, Abstracts and Proceedings Volume 67-70 [M. Carrano/M. Carrano/M. Carrano]
45810 A. V. Lopatin, A. O. Averianov, E. N. Maschenko and S. V. Leschinskiy. 2010. Early Cretaceous mammals of western Siberia: 3. Zhangeotheriidae. Paleontological Journal 44(5):573-583 [P. Mannion/J. Tennant/J. Tennant]
59918 A. V. Lopatin, E. N. Mashchenko, K. K. Tarasenko, A. V. Podlesnov, N. V. Demidenko and E. A. Kuzmina. 2015. A unique burial site of Early Cretaceous vertebrates in western Siberia (the Shestakovo 3 locality, Kemerovo Province, Russia). Doklady Biological Sciences 462:148-151 [M. Carrano/M. Carrano]
39703 E. N. Maschenko and A. V. Lopatin. 2002. A new Early Cretaceous mammal from Western Siberia. Doklady Biological Sciences 386:475-477 [R. Benson/R. Benson/R. Benson]
51098 J. K. O'Connor, A. O. Averianov, and N. V. Zelenkov. 2014. A confuciusornithiform (Aves, Pygostylia)-like tarsometatarsus from the Early Cretaceous of Siberia and a discussion of the evolution of avian hind limb musculature. Journal of Vertebrate Paleontology 34(3):647-656 [P. Mannion/J. Tennant]
14209ETE V. M. Podobina, G. M. Tatyanin, S. V. Leschchinskiy, A. V. Voronkevich, and A. V. Faingerts. 2002. A review of the Early Cretaceous fossil vertebrates of the Sehstakovo assemblage (south-east western Siberia, Russia). M. Wagreich (ed.), Aspects of Cretaceous Stratigraphy and Palaeobiogeography, Proceedings of the 6th International Cretaceous Symposium. Osterreichische Akademie der Wissenschaften Schriftenreihe der Erdwissenschaftlichen Kommissionen 15:239-247 [M. Carrano/M. Carrano/M. Carrano]
72250 P. P. Skutschas, V. D. Markova, E. A. Boitsova, S. V. Leshchinskiy, S. V. Ivantsov, E. N. Maschenko, and A. O. Averianov. 2017. The first dinosaur egg from the Lower Cretaceous of Western Siberia, Russia. Historical Biology 31:836-844 [G. Lloyd/G. Lloyd/G. Varnham]
77054 P. P. Skutschas, S. S. Morozov, A. O. Averianov, S. V. Leshchinskiy, S. V. Ivantsov, A. V. Fayngerts, O. A. Feofanova, O. N. Vladimirova, and D. A. Slobodin. 2021. Femoral histology and growth patterns of the ceratopsian dinosaur Psittacosaurus sibiricus from the Early Cretaceous of Western Siberia. Acta Palaeontologica Polonica 66 [M. Carrano/M. Carrano]