Berezovsk Quarry: Bathonian, Russian Federation
collected by S. Krasnolutskii et al. 2000-2005, 2010–2012

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Chondrichthyes - Hybodontiformes - Hybodontidae
Hybodus sp. Agassiz 1834
Averianov et al. 2016
Sinamiidae
Sinamiidae indet. Berg 1940
Averianov et al. 2016
Acipenseriformes
Acipenseriformes indet. Berg 1940
Averianov et al. 2016
Dipnoi
Dipnoi indet. Müller 1884
Lopatin and Averianov 2006
    = Ferganoceratodus sp. Nessov and Kaznyshkin 1985
Averianov et al. 2016
Amphibia - Temnospondyli - Discoglossidae
? Eodiscoglossus sp. Villalta 1957
Skutschas et al. 2016 1 specimen
ZIN PH 27/144
Amphibia - Temnospondyli
Caudata informal indet. A (Scopoli 1777)
Averianov et al. 2016
Berezovsk salamander A
Caudata indet. Scopoli 1777
Lopatin and Averianov 2006 1 specimen
    = Urupia monstrosa Skutschas and Krasnolutskii 2011
Averianov et al. 2016
Urupia monstrosa n. gen., n. sp. Skutschas and Krasnolutskii 2011
Skutschas and Krasnolutskii 2011 6 specimens
ZIN PH 1/144 (holotype atlas), 3/144, 4/144 (dorsal vertebrae); PM TGU 200.10, ZIN PH 2/144 (anterior dentary fragments); PM TGU 200/11 (femur)
Egoria malashichevi n. gen., n. sp. Skutschas et al. 2020
Skutschas et al. 2020 6 specimens
Amphibia - Temnospondyli - Urodela
Kiyatriton krasnolutskii n. sp. Skutchas 2016
Averianov et al. 2016
Amphibia - Temnospondyli - Karauridae
cf. Karauridae indet. Ivachnenko 1978
Skutschas 2006 1 specimen
Tritylodontidae
Tritylodontidae indet. Cope 1884
Lopatin and Averianov 2006 18 specimens
    = Stereognathus sp. Charlesworth 1854
Averianov et al. 2017
ZIN PH 19/117–38/117
Mammalia
Mammalia indet. Linnaeus 1758
Mammalia - Multituberculata
Tagaria antiqua n. gen., n. sp. Averianov et al. 2020
Averianov et al. 2020
Holotype: PIN 5087/108, right P5; LMCCE 004/10, left M2; PIN 5087/50, right M2
Tashtykia primaeva n. gen., n. sp. Averianov et al. 2020
Averianov et al. 2020
Holotype: PIN 5087/52, left p4; PIN 5087/25, right I2; LMCCE 010/2, right upper deciduous premolar; PIN 5087/46, right upper anterior premolar; PIN 5087/21, left P5
Mammalia
Sineleutherus issedonicus n. sp. Averianov et al. 2011
Averianov et al. 2011 10 specimens
PIN 5087/9 to 5087/18
Mammalia - Amphitheriidae
Amphibetulimus krasnolutskii n. gen., n. sp. Lopatin and Averianov 2007
Lopatin and Averianov 2007 7 specimens
PIN, no. 5087/3, right dentary fragment with M3 and alveoli and roots of M1, M2, M4, and M5; PIN 5087/34, right upper molar. PIN 5087/39, left dentary fragment with p1, alveoli or roots for i1–4, double rooted canine, and p2. PIN 5087/44, right dentary fragment with alveoli for two middle molars. PIN 5087/43, right dentary fragment with alveoli for ultimate and penultimate molars. PIN 5087/40, right dentary fragment with ultimate molar and roots of the penultimate molar. PIN 5087/45, right dentary fragment with alveolus for ultimate molar
Mammalia - Dryolestidae
Dryolestidae indet. Marsh 1879
Lopatin and Averianov 2006 11 specimens
    = Anthracolestes sergeii n. gen., n. sp. Averianov et al. 2014
Averianov et al. 2014
Holotype—PIN 5087/22, left lower molar. Referred Specimens—PIN 5087/23, damaged right lower molar; PIN 5087/36, right lower molar missing talonid; PIN 5087/27, left dentary fragment with alveoli for i1–4, double-rooted canine, and p1–3; PIN 5087/29, right dentary fragment with alveoli for i1–4, double-rooted canine, and p1–3; PIN 5087/32, left dentary with alveoli for double-rooted canine and p1–4; PM TGU 200/3- BR-3, left dentary fragment with alveoli for four last molars; PIN 5087/24, 5087/30, and 5087/31, left dentary fragments with alveoli for three last molars; PIN 5087/33, left dentary fragment with alveoli for two middle molars.
Mammalia - Eutriconodonta
Eutriconodonta indet. Kermack et al. 1973
Averianov et al. 2008 1 specimen
PIN 5087/4, left upper molariform tooth
Docodonta
Itatodon tatarinovi n. gen., n. sp. Lopatin and Averianov 2005
Lopatin and Averianov 2006 4 specimens
PIN 5087/2 (holotype, isolated right lower molar), 5087/7, PM TGU 200/3-BR-7
Docodonta - Tegotheriidae
Hutegotherium yaomingi n. gen., n. sp. Averianov et al. 2010
Averianov et al. 2010 3 specimens
PM TGU 200/5-BR-2; PIN 5087/6, 5087/8
Docodonta - Simpsonodontidae
Docodonta indet. Kretzoi 1946
Lopatin and Averianov 2006 2 specimens
PIN 5087/1, posterior fragment of a right denta− ry with alveoli for the ultimate molar, posterior alveolus for the penultimate molar and partially preserved coronoid and pseudangular processes. PM TGU 200/3−BR−1, posterior fragment of a right dentary with alveoli for the two last mo− lars and partially preserved coronoid process
    = Simpsonodon sibiricus n. sp. Averianov et al. 2010
Averianov et al. 2010
PIN 5087/5
Reptilia - Testudines
Testudines informal indet. 1 Batsch 1788
Averianov et al. 2016 1 specimen
ZIN PH, no. 1/186, left costal VIII.
Testudines informal indet. 2 Batsch 1788
Danilov et al. 2018 2 specimens
ZIN PH, no. 2/186, left xiphiplastron fragment; and specimen ZIN PH, no. 3/186, shell fragment.
Testudines informal indet. 3 Batsch 1788
Danilov et al. 2018 1 specimen
ZIN PH, no. 4/186, distal part of a right humerus.
Reptilia - Testudines - Xinjiangchelyidae
Xinjiangchelys sp. Yeh 1986
Skutschas 2006
    = Annemys sp. Sukhanov and Narmandakh 2006
Averianov et al. 2016
Danilov et al. 2018 8 specimens
ZIN PH, no. 431/50, costal VIII; ZIN PH, nos. 1/207 and 2/207, medial parts of costal VIII; ZIN PH, nos. 14/187 and 17/187, xiphiplastra; ZIN PH, no. 3/207, xiphiplastron fragment; ZIN PH, no. 6/185, right humerus; and ZIN PH, no. 4/207, distal part of the left humerus.
    = Annemys variabilis n. sp. Obraztsova et al. 2022
Obraztsova et al. 2022
Holotype. ZIN PH 26/184, braincase (Fig. 3E–H). Referred material. Thousands of isolated bones and several fragmentary specimens in the following collections: ZIN PH 50, more than 2000 isolated shell bones; ZIN PH 178, more than 350 basisphenoids; ZIN PH 183, vertebrae and cervical ribs; ZIN PH 184, three braincases, and isolated skull bones (in total more than 6500 specimens); ZIN PH 185, bones of girdles and limbs; ZIN PH 187, parts of the skeleton, big shell fragments and complete plates; SRM n/n, carapace and plastron restored based on fragments of several individuals (Krasnolutskii 2006, fig. 19; Skutschas et al. 2009, foto 1, fig. 4)
Reptilia
cf. Paramacellodidae indet. Estes 1983
Skutschas 2006
    = Scincomorpha indet. Camp 1923
Averianov et al. 2016
Reptilia - Goniopholididae
cf. Sunosuchus sp. Young 1948
Skutschas 2006 6 specimens
    = Goniopholididae indet. Cope 1875
Kuzmin et al. 2013
ZIN PH 1/176 (maxillary fragment); ZIN PH 2/176 (fragment of left nasal); ZIN PH 3/176 (anterior fragment of right splenial); ZIN PH 4-8/176, (teeth and 17 unnumbered teeth); ZIN PH 9/176 (fragment of the dorsal osteoderm); ZIN PH 10/176 (fragmentary ventral osteoderm)
Reptilia - Heterodontosauridae
Heterodontosauridae indet. Romer 1966
Skutschas 2006
Reptilia
Stegosauria indet. Marsh 1877
Skutschas 2006 2 individuals
ZIN PH 1–2/117; SHRM various
Reptilia - Proceratosauridae
cf. Dromaeosauridae indet. Colbert and Russell 1969
Skutschas 2006
    = Kileskus aristotocus Averianov et al. 2010
Averianov et al. 2016
Kileskus aristotocus n. gen., n. sp. Averianov et al. 2010
Averianov et al. 2010
Reptilia - Mamenchisauridae
Titanosauriformes indet. Salgado et al. 1997
Skutschas 2006
    = Mamenchisauridae indet. Chao 1965
Averianov et al. 2016
PM TGU 200/5-BR-308-322; SM 4557; ZIN PH 41-62/117 (teeth); SM 4551-4554, 4556; ZIN PH 39, 40/117 (caudal vertebrae)
Reptilia - Rhamphorhynchidae
Pterodactyloidea indet. Plieninger 1901
Skutschas 2006
    = Rhamphorhynchidae indet. Seeley 1870
Averianov et al. 2016
Reptilia - Choristodera
? Choristodera indet. Cope 1876
Lopatin and Averianov 2006
Palaeonisciformes
Palaeonisciformes indet. Hay 1902
Skutschas 2006
see common names

Geography
Country:Russian Federation State/province:Krasnoyarsk County:Sharypovo
Coordinates: 55.7° North, 89.2° East (view map)
Paleocoordinates:58.2° North, 86.7° East
Basis of coordinate:stated in text
Geographic resolution:small collection
Time
Period:Jurassic Epoch:Middle Jurassic
Stage:Bathonian 10 m.y. bin:Jurassic 4
Key time interval:Bathonian
Age range of interval:168.30000 - 166.10000 m.y. ago
Stratigraphy
Formation:Itat Member:Upper
Stratigraphic resolution:group of beds
Stratigraphy comments: upper part of formation
The age of the Upper Mem- ber of the Itat Formation is Bathonian according to the pollen complex, which consists of Cyathidites minor, Piceapollenites sp., Eboracia torosa, Quadraeculina limbata, and Classopollis sp. (Raevskaya et al. 1993)
Lithology and environment
Primary lithology:gray claystone
Includes fossils?Y
Lithology description: "grey clays" Fossil rich layer tops a thick coal seam
Environment:fluvial indet.
Geology comments: By mid-Jurassic times, a low-profile alluvial plain had developed in this area that was characterized by anas- tomosing rivers and well-defined floodplains (Le Heron et al. 2008). Due to the fluvial origin of the sediments, vertebrate remains are mostly disarticulated and often water worn. However, the predominantly good preserva- tion of vertebrate remains demonstrates that transport occurred over short distances.
Taphonomy
Modes of preservation:body
Lagerst�tten type:concentrate
Size of fossils:macrofossils,mesofossils,microfossils
Preservation of anatomical detail:good
Disassociated major elements:many
Disassociated minor elements:many
Fragmentation:occasional
Collection methods and comments
Collection excludes:some genera,species names
Collection methods:bulk,selective quarrying,surface (in situ),mechanical,sieve,field collection
Reason for describing collection:general faunal/floral analysis
Museum repositories:PIN
Collectors:S. Krasnolutskii et al. Collection dates:2000-2005, 2010–2012
Collection method comments: Itatodon tatarinovi material housed in the Paleontological Museum of Tomsk State
University, Tomsk, Russia (PM TGU), and in the Paleontological Institute of the Russian Academy of Sciences, Moscow, Russia (PIN)
2010-2012 collection by a Joint Russian-German expedition: "The newly collected mammal specimens were obtained during screen-washing of ~12.5 tons of matrix. The jaw fragments were found in the field by picking the coarse fraction of fossiliferous concentrate. The teeth were recovered during sorting of the fine fraction in the laboratory.

The specimens described here are housed in the Borissiak Paleontological Institute of the Russian Academy of Sciences, Moscow (PIN).
"...other specimens were collected in 2010–2012 during a joint Russian-German expedition to Berezovsk Quarry. The edentulous jaw fragments were found in the field while picking the coarse fraction of fossiliferous concentrate obtained from screenwashing of ∼12.5 tons of matrix" (Avaerianov et al. 2014)"
Metadata
Also known as:Stegosaur Quarry; Berezovsk coal mine
Database number:61891
Authorizer:M. Carrano, E. Vlachos, P. Mannion, R. Benson, O. Rauhut Enterer:M. Carrano, J. Tennant, G. Varnham, O. Rauhut, P. Mannion, R. Benson, E. Vlachos
Modifier:G. Varnham Research group:vertebrate
Created:2006-07-06 14:56:38 Last modified:2022-03-30 07:45:38
Access level:the public Released:2006-07-06 14:56:38
Creative Commons license:CC BY
Reference information

Primary reference:

44653. A. O. Averianov, A. V. Lopatin, P. P. Skutschas, N. V. Matynovich, S. V. Leschinkskiy, A. S. Rezvyi, S. A. Krasnolutskii and A. V. Fayngertz. 2005. Discovery of Middle Jurassic mammals from Siberia. Acta Palaeontologica Polonica 50(4):789-797 [P. Mannion/J. Tennant]

Secondary references:

68147 A. Averianov, S. Krasnolutskii, S. Ivantsov, P. Skutschas, R. Schellhorn, J. Schultz, and T. Martin. 2019. Sauropod remains from the Middle Jurassic Itat Formation of West Siberia, Russia. Paläontologische Zeitschrift [P. Mannion/P. Mannion]
55679 A. Averianov, T. Martin, A. Lopatin and S. Krasnolutskii. 2015. Stem therian mammal Amphibetulimus from the Middle Jurassic of Siberia. Palaeontologische Zeitschrift 89:197-206 [P. Mannion/J. Tennant]
61822 A. Averianov, T. Martin, P. Skutschas, I. Danilov, J. Schultz, R. Schellhorn, E. Obraztsova, A. Lopatin, E. Sttchevskaya, I. Kuzmin, S. Krasnotutskii and S. Ivantsov. 2016. Middle Jurassic vertebrate assemblage of Berezovsk coal mine in western Siberia (Russia). Global Geology 19(4):187-204 [E. Vlachos/E. Vlachos/M. Carrano]
34709ETE A. O. Averianov and S. A. Krasnolutskii. 2009. Stegosaur remains from the Middle Jurassic of west Siberia. Proceedings of the Zoological Institute RAS 313(2):153-167 [M. Carrano/M. Carrano]
36119 A. O. Averianov, S. A. Krasnolutskii, and S. V. Ivantsov. 2010. A new basal coelurosaur (Dinosauria: Theropoda) from the Middle Jurassic of Siberia. Proceedings of the Zoological Institute, Russian Academy of Science 314(1):42-57 [O. Rauhut/O. Rauhut]
44655 A. O. Averianov and A. V. Lopatin. 2006. Itatodon tatarinovi (Tegotheriidae, Mammalia), a docodont from the Middle Jurassic of Western Siberia and phylogenetic analysis of Docodonta. Paleontological Journal 40(6):668-677 [P. Mannion/J. Tennant]
44847 A. O. Averianov, A. V. Lopatin, and S. A. Krasnolutskii. 2008. An amphilestid-grade eutriconodontan from the Middle Jurassic of Russia. Russian Journal of Theriology 7(1):1-4 [P. Mannion/J. Tennant/J. Tennant]
34572ETE A. O. Averianov, A. V. Lopatin, S. A. Krasnolutskii and S. V. Ivantsov. 2010. New docodontans from the Middle Jurassic of Siberia and reanalysis of Docodonta interrelationships. Proceedings of the Zoological Institute RAS 314(2):121-148 [M. Carrano/M. Carrano]
36940ETE A. O. Averianov, A. V. Lopatin, and S. A. Krasnolutskii. 2011. The first haramiyid (Mammalia, Allotheria) from the Jurassic of Russia. Doklady Biological Sciences 437:103-106 [M. Carrano/M. Carrano/M. Carrano]
86461 A. O. Averianov and A. V. Lopatin. 2023. Dinosaurs of Russia: a review of the localities. Herald of the Russian Academy of Sciences 93(4) [M. Carrano/M. Carrano/M. Carrano]
51915 A. O. Averianov, T. Martin, and A. Lopatin. 2014. The oldest dryolestid mammal from the Middle Jurassic of Siberia. Journal of Vertebrate Paleontology 34(4):924-931 [R. Benson/R. Benson]
65142 A. O. Averianov, T. Martin, A. V. Lopatin, J. A. Schultz, P. P. Skutschas, R. Schellhorn, and S. A. Krasnolutskii. 2017. A tritylodontid synapsid from the Middle Jurassic of Siberia and the taxonomy of derived tritylodontids. Journal of Vertebrate Paleontology 37(5):e1363767:1-10 [M. Carrano/M. Carrano]
81047 A. O. Averianov, T. Martin, A. V. Lopatin, J. A. Schultz, R. Schellhorn, S. Krasnolutskii, P. Skutschas and S. Ivantsov. 2020. Multituberculate mammals from the Middle Jurassic of western Siberia, Russia, and the origin of Multituberculata . Papers in Palaeontology 7(2):769-787 [P. Mannion/G. Varnham]
71292 A. O. Averianov, P. P. Skutschas, R. Schellhorn, A. V. Lopatin, P. N. Kolosov, V. V. Kolchanov, D. D. Vitenko, D. V. Grigoriev, and T. Martin. 2019. The northernmost sauropod record in the Northern Hemisphere. Lethaia 53:362-368 [P. Mannion/P. Mannion/M. Carrano]
64692 I. G. Danilov, E. M. Obraztsova, E. A. Boitsova and P. P. Skutchas. 2018. Diversity of Middle Jurassic Turtles from the Berezovsk Quarry Locality, Krasnoyarsk Region, Russia: Morphology and Histological Study. Paleontological Journal 52(1):58-70 [E. Vlachos/E. Vlachos]
49777 I. T. Kuzmin, P. P. Skutschas, O. I. Grigorieva and S. A. Krasnolutskii. 2013. Goniopholidid crocodylomorph from the Middle Jurassic Berezovsk Quarry Locality (Western Siberia, Russia). Proceedings of the Zoological Institute of the Russian Academy of Sciences 317(4):452-458 [P. Mannion/P. Mannion/P. Mannion]
45969 A. V. Lopatin and A. O. Averianov. 2005. A new docodont (Docodonta, Mammalia) from the Middle Jurassic of Siberia. General Biology 405(2):434-436 [P. Mannion/J. Tennant/J. Tennant]
17882ETE A. V. Lopatin and A. O. Averianov. 2006. Mesozoic mammals of Russia. 9th International Symposium on Mesozoic Terrestrial Ecosystems and Biota, Abstracts and Proceedings Volume 67-70 [M. Carrano/M. Carrano/M. Carrano]
44659 A. V. Lopatin and A. O. Averianov. 2007. The earliest Asiatic pretribosphenic mammal (Cladotheria, Amphitheriidae) from the Middle Jurassic of Siberia. Doklady Biological Sciences 417:432-434 [P. Mannion/J. Tennant]
82558 E. M. Obraztsova, S. A. Krasnolutskii, V. B. Sukhanov and I. G. Danilov. 2022. Xinjiangchelyid turtles from the Middle Jurassic of the Berezovsk coal mine (Krasnoyarsk Territory, Russia): systematics, skeletal morphology, variation, relationships and palaeobiogeographic implications. Journal of Systematic Palaeontology 20(1):1-61 [E. Vlachos/E. Vlachos/E. Vlachos]
17903ETE P. Skutschas. 2006. Mesozoic amphibians from Siberia, Russia. 9th International Symposium on Mesozoic Terrestrial Ecosystems and Biota, Abstracts and Proceedings Volume 123-126 [M. Carrano/M. Carrano]
78277 P. Skutschas, V. Kolchanov, S. Krasnolutskii, A. Averianov, R. Schellhorn, J. Schultz, and T. Martin. 2020. A new small-sized stem salamander from the Middle Jurassic of Western Siberia, Russia. PLoS One 15(2):e0228610 [M. Uhen/K. Eleazer]
38941 P. P. Skutschas and S. A. Krasnolutskii. 2011. A new genus and species of basal salamanders from the Middle Jurassic of western Siberia, Russia. Proceedings of the Zoological Institute of the Russian Academy of Sciences 315(2):167-175 [R. Benson/R. Benson/P. Mannion]
58264 P. P. Skutschas, T. Martin, and S. A. Krasnolutskii. 2016. First Jurassic frog from Siberia, Russia. Journal of Vertebrate Paleontology 36:e1101376:1-3 [P. Mannion/P. Mannion]