Nammoura quarry (BMNH collection): Middle Cenomanian, Lebanon

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Actinopteri - Albuliformes - Albulidae
Lebonichthys namourensis n. sp. Forey et al. 2003
4 specimens
BMNH P.63260, P.63251, P.63252, P.63294
Actinopteri - Anguilliformes
Luenchelys minimus Belouze et al. 2003
1 specimen
BMNH P.62692a/b
Actinopteri - Elopiformes - Elopidae
Ctenodentelops striatus n. gen., n. sp. Forey et al. 2003
36 specimens
BMNH P. 63228, P.63227, P.63237-P.63239, P.63253-P.63258, P.63287-P.63292, P.63295, P.63299, P.63301, P.63304, P.63326-P.63331, P.63601, MNHN NRA-54a, NRA-22, NRA-15, 1991-3-4, 1989-12-1, NRA-16, 1991-3-5, MNHN 1991-1-7b
Actinopteri
Ornategulum sardinioides (Pictet 1850)
1 specimen
BMNH P.63293
Scombroclupea macrophthalma (Heckel 1849)
8 specimens
BMNH P.62525, P.62526, P.63143, P.63152, P.63153, P.63157, P.63165, P.63247
Scombroclupea diminuta n. sp. Forey et al. 2003
9 specimens
BMNH P.63136, P.63150, P.63158, P.63159, P.63248, P.63168, P.63272, P.63303, P.63230
Actinopteri - Ellimmichthyiformes - Armigatidae
Armigatus namourensis n. sp. Forey et al. 2003
23 specimens
BMNH P.59808, P.62527-29, P.63135, P.63137-42, P.63145-47, P.63148a/b, P.63149a/b, P.63151a/b, P.63156, P.63161-64, P.63598a/b
Armigatus alticorpus n. sp. Forey et al. 2003
2 specimens
BMNH P.63134, P.63144
Actinopteri - Ellimmichthyiformes - Paraclupeidae
Triplomystus noorae n. gen., n. sp. Forey et al. 2003
4 specimens
BMNH P.62517, P.62519, P.62520, P.63325
Triplomystus oligoscutatus n. gen., n. sp. Forey et al. 2003
2 specimens
BMNH P.63155, P.63154
Actinopteri - Aulopiformes
Serrilepis prymnostrigos n. sp. Forey et al. 2003
6 specimens
BMNH P.62523, P.62968, P.62969, P.62970, P.62971, P.63322
Serrilepis minor n. sp. Forey et al. 2003
1 specimen
BMNH P.62676
Actinopteri - Aulopiformes - Eurypholidae
Eurypholis boissieri Pictet 1850
1 specimen
BMNH P.63323
Actinopteri - Aulopiformes - Aulopidae
Nematonotus bottae (Pictet and Humbert 1866)
1 specimen
BMNH P.63319
Actinopteri - Aulopiformes - Enchodontidae
Enchodus mecoanalis n. sp. Forey et al. 2003
2 specimens
BMNH P.62522, P.63250
Actinopteri - Aulopiformes - Apateopholidae
Apateopholis laniatus (Davis 1887)
1 specimen
BMNH P.63263
Actinopteri - Aulopiformes - Dercetidae
Rhynchodercetis cf. gracilis Chalifa 1989
4 specimens
recombined as Hastichthys gracilis
BMNH P. 63235, P. 63246, P. 63332, P. 62677
Rhynchodercetis cf. yovanovitchi Arambourg 1954
5 specimens
BMNH P.63236, P.62678, P.62690, P.63262, P.63599
Actinopteri - Beryciformes
Paracentrus lebanonensis n. gen., n. sp. Forey et al. 2003
1 specimen
BMNH P.62518a/b
Actinopteri
Gigapteryx tethyestris n. gen., n. sp. Forey et al. 2003
1 specimen
BMNH P.62521
Gaudryella gaudryi Pictet and Humbert 1866
54 specimens
BMNH P.63205-26, P.63264-67, P.63273-84, P.63297-8, P.63302, P.63306-18
Actinopteri - Ichthyodectiformes - Cladocyclidae
Eubiodectes libanicus (Pictet and Humbert 1866)
9 specimens
recombined as Ichthyodectes libanicus
BMNH P.62691, P.63234, P.63194, P.63600, AUB 100821, MNHN 2000-11-17, MNHN 1993-1-2, MNHN 1998-11, MNHN 1993-1-1
Actinopteri - Aspidorhynchiformes - Aspidorhynchidae
Belonostomus sp. Agassiz 1834
6 specimens
BMNH P.62524, P.63242a/b, P.62673a/b, P.63285, P63241, P63286
Actinopteri - Pycnodontiformes
Pycnodontiformes indet. Berg 1937
1 specimen
BMNH P.62516
Actinopteri - Pycnodontiformes - Pycnodontidae
Nursallia sp. Blot 1987
2 specimens
BMNH P.63244a/b, P.63245a/b
    = Nursallia tethysensis Capasso et al. 2009
Capasso et al. 2009
? Proscinetes sp. Gistel 1848
1 specimen
BMNH P.63320a/b
Chondrichthyes - Trygonidae
Cyclobatis oligodactylus Egerton 1844
1 specimen
BMNH P.63193
Chondrichthyes - Lamniformes - Anacoracidae
Squalicorax falcatus (Agassiz 1843)
1 specimen
BMNH P.63243
see common names

Geography
Country:Lebanon
Coordinates: 34.1° North, 35.7° East (view map)
Paleocoordinates:12.9° North, 30.3° East
Basis of coordinate:stated in text
Time
Period:Cretaceous Epoch:Late/Upper Cretaceous
Stage:Cenomanian 10 m.y. bin:Cretaceous 5
Key time interval:Middle Cenomanian
Age range of interval:100.50000 - 93.90000 m.y. ago
Stratigraphy
Formation:Sannine
Stratigraphy comments: At Namoura the fish-bearing Lagerstatte of 'unit 3' are apparently without microfossils. However, Dalla Vecchia et al. (2002) record that in the overlying 'unit 4' (about 15 m above the fish-bearing layer) the following foraminifers are found: Pseudorhapidionina laurensis (de Castro, 1965), P. dubia (de Castro, 1965), Prealviolina group cretacea (d'Archiac, 1837), Pseudodomia drorimensis Reiss, Hamaoui & Ecker, 1964 and Conicorbitolina group conico (d'Archiac, 1837). These taxa are seen to coincide stratigraphically within the base of the Upper Cenomanian (Schroeder & Neumann 1985: Table 1). Thus, a conservative dating places the Namoura Lagerstatte, which lies just below the foraminfer-bearing 'unit 4' as being in the middle portion of the Middle Cenomanian.
Lithology and environment
Primary lithology:planar lamination,gray,green,white,yellow,blue cherty/siliceous wackestone
Secondary lithology: carbonaceous mudstone
Includes fossils?Y
Lithology description: The rock is a classic lithographic limestone: a pale, laminated micrite, resembling that at the classic Tithonian deposits at Solnhofen, Bavaria.
Environment:lagoonal/restricted shallow subtidal
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Collection methods and comments
Collection excludes:some macrofossils
Reason for describing collection:general faunal/floral analysis
Museum repositories:BMNH
Collection method comments: Our specimens were purchased from locals and fossil dealers.
Taxonomic list comments:In addition to fishes we have also seen a few turtles, crustaceans, an echinoid and abundant land plants from the Namoura plattenkalk. Dalla Vecchia et al. (2002) also recorded bivalves, snakes, a dolichosaur (marine lizard) and a theropod dinosaur. At least one articulated enanthiornine bird as well as isolated feathers have been found.
Metadata
Database number:124426
Authorizer:G. Lloyd, M. Clapham Enterer:G. Lloyd, M. Clapham
Modifier:M. Clapham Research group:vertebrate
Created:2012-02-15 03:39:22 Last modified:2021-08-18 00:17:42
Access level:the public Released:2012-02-15 03:39:22
Creative Commons license:CC BY
Reference information

Primary reference:
40127. P. L. Forey, L. Yi, C. Patterson and C. E. Davis. 2003. Fossil fishes from the Cenomanian (Upper Cretaceous) of Namoura, Lebanon. Journal of Systematic Palaeontology 1(4):227-330 [G. Lloyd/G. Lloyd]

Secondary references:

77370 L. L. Capasso, P. A. Saad, and L. Tavarne. 2009. Nursallia tethysensis sp. nov., a new pycnodont fish (Neopterygii: †Halecostomi) from the Cenomanian of Lebanon. 79:117-136 [M. Clapham/S. Hegde/S. Hegde]