Guanling, Lower Xiaowa Formation fossil lagerstätte: Tuvalian, China

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Similingula cf. lipoldi
Wang et al. 2008
Reptilia - Testudines
Odontochelys semitestacea n. gen., n. sp. Li et al. 2008
Li et al. 2008 3 individuals
IVPP V 13240 (paratype), V 15639 (holotype), V 15653
Reptilia - Shastasauridae
Guanlingichthyosaurus liangae (Yin et al. 2000)
Wang et al. 2008
original and current combination Guanlingsaurus liangae
Ghuizhouichthyosaurus tangae (Cao et al. 2000)
Wang et al. 2008
original and current combination Guizhouichthyosaurus tangae
Reptilia - Toretocnemidae
Qianichthyosaurus zhoui Li 1999
Wang et al. 2008
Reptilia - Thalattosauria - Askeptosauridae
Anshunsaurus huangguoshuensis Liu 1999
Wang et al. 2008
Reptilia - Thalattosauria
Xinpusaurus bamaolinensis Cheng 2003
Wang et al. 2008
Xinpusaurus suni Yin 2000
Wang et al. 2008
Reptilia - Placodontia - Cyamodontidae
Sinocyamodus xinpuensis Li 2000
Wang et al. 2008
Cyamodus orientalis n. sp. Wang et al. 2019
Wang et al. 2019 1 individual
Zhejiang Museum of Natural History, Hangzhou, ZMNH M8820 (holotype), "an articulated and nearly complete skeleton with three-dimensionally preserved skull and lacking only the forelimbs, representing an adult individual based on overall body size"
Reptilia - Placodontia - Placochelyidae
Psephochelys polyosteoderma n. gen., n. sp. Li and Rieppel 2002
1 individual
IVPP V 12442
Conodonta - Ozarkodinida - Gondolellidae
Metapolygnathus polygnathiformis (Budurov and Stefanov 1965)
Wang et al. 2008
recombined as Paragondolella polygnathiformis
Metapolygnathus nodosus (Hayashi 1968)
Wang et al. 2008
recombined as Hayashiella nodosa
Crinoidea - Roveacrinida - Somphocrinidae
Osteocrinus cf. virgatus Kristan-Tollmann 1970
Wang et al. 2008
Osteocrinus cf. spinosus Kristan-Tollmann 1970
Wang et al. 2008
Crinoidea - Encrinida - Traumatocrinidae
Traumatocrinus sp. Woehrmann 1889
Wang et al. 2008
Holothuroidea
Holothuroidea indet. (de Blainville 1834)
Wang et al. 2008
Craniata - Craniida - Craniidae
? Crania sp. Retzius 1781
Wang et al. 2008
Bivalvia - Ostreida - Pergamidiidae
Krumbeckiella sp. Ichikawa 1958
Wang et al. 2008
Bivalvia - Ostreida - Bakevelliidae
Angustella sp. Waagen 1907
Wang et al. 2008
Bivalvia - Ostreida - Halobiidae
Daonella bifurcata
Wang et al. 2008
Halobia subcomata Kittl 1912
Wang et al. 2008
Cephalopoda - Ceratitida - Clionitidae
Clionites cf. zeilleri
Wang et al. 2008
replaced by Clionitites
Cephalopoda - Ceratitida - Tibetitidae
Hauerites cf. himalayanus
Wang et al. 2008
Cephalopoda - Ceratitida - Trachyceratidae
Protrachyceras cf. douvillei
Wang et al. 2008
Protrachyceras longiangense
Wang et al. 2008
Protrachyceras deprati (Mansuy 1912)
Wang et al. 2008
Protrachyceras ladinum (Mojsisovics 1882)
Wang et al. 2008
Protrachyceras costulatum Mansuy 1912
Wang et al. 2008
Paratrachyceras cf. hofmanni (Boekh 1873)
Wang et al. 2008
Trachyceras sinensis
Wang et al. 2008
Trachyceras multituberculatum Hsu 1940
Wang et al. 2008
Trachyceras cf. aon (Munster 1834)
Wang et al. 2008
see common names

Geography
Country:China State/province:Guizhou
Coordinates: 25.9° North, 105.4° East (view map)
Paleocoordinates:26.9° North, 87.4° East
Basis of coordinate:based on nearby landmark
Geographic resolution:outcrop
Time
Period:Triassic Epoch:Late/Upper Triassic
Stage:Carnian 10 m.y. bin:Triassic 3
Key time interval:Tuvalian
Age range of interval:233.40000 - 227.00000 m.y. ago
Stratigraphy
Formation:Xiaowa Member:Lower
Stratigraphic resolution:group of beds
Stratigraphy comments: 12 m thick, previously known as Wayao Member of Falang Formation or as Wayao Formation
JA: originally entered as Carnian; the Metapolygnathus polygnatiformis and M. nodosus zones are both Tuvalian according to Lucas et al. 2012
Lithology and environment
Primary lithology: lithified "shale"
Secondary lithology: lithified marl
Lithology description: lower unit (5 m thick) of medium- to thick-bedded, grey bioclastic micrites with yellow-green shale interbeds; upper unit (7 m thick) of greyish to black, thin- to medium-bedded laminated mudstone with argillaceous limestone interbeds and marlstones
Environment:deep-water indet.
Geology comments: deep and quiet water, the lower unit with generally oxygenated bottom water; the upper unit shaly interval with anoxic conditions and the greatest water depth
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Preservation of anatomical detail:excellent
Spatial resolution:parautochthonous
Collection methods and comments
Collection methods:selective quarrying,mechanical,field collection
Reason for describing collection:general faunal/floral analysis
Metadata
Database number:85166
Authorizer:W. Kiessling, B. Allen Enterer:U. Merkel, B. Allen
Modifier:J. Alroy Research group:marine invertebrate,vertebrate
Created:2008-12-04 04:10:03 Last modified:2012-09-05 23:51:02
Access level:the public Released:2008-12-04 04:10:03
Creative Commons license:CC BY
Reference information

Primary reference:

39730. C. Li and O. Rieppel. 2002. A new cyamodontid placodont from Triassic of Guizhou, China. Chinese Science Bulletin 47(5):403-407 [M. Carrano/M. Carrano]

Secondary references:

28659 C. Li, X. C. Wu, O. Rieppel, L. T. Wang, and L. J. Zhao. 2008. An ancestral turtle from the Late Triassic of southwestern China. Nature 456:497-501 [W. Kiessling/U. Merkel]
80944 W. Wang, C. Li, T. M. Scheyer and L. J. Zhao. 2019. A new species of Cyamodus (Placodontia, Sauropterygia) from the early Late Triassic of south-west China. Journal of Systematic Palaeontology 17(17):1457-1476 [B. Allen/B. Allen]
28658 X. Wang, G. H. Bachmann, H. Hagdorn, P. M. Sanders, G. Cuny, X. Chen, C. Wang, L. Chen, L. Cheng, F. Meng, and G. Xu. 2008. The Late Triassic black shales of the Guanling area, Guizhou province, south-west China: a unique marine reptile and pelagic crinoid fossil lagerstätte. Palaeontology 51(1):27-61 [W. Kiessling/U. Merkel]