MAD93-33 (SUNY): Maastrichtian, Madagascar

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Reptilia - Cordylidae
Konkasaurus mahalana n. gen., n. sp. Krause et al. 2003
Krause et al. 2003 1 individual
Reptilia - Madtsoiidae
Madtsoia madagascariensis Hoffstetter 1961
LaDuke et al. 2010
Reptilia
Simosuchus clarki Buckley et al. 2000
Krause et al. 2010
FMNH PR 2591, 2592 (2 isolated teeth)
Araripesuchus sp. Price 1959
Krause et al. 2003 5 individuals
    = Araripesuchus tsangatsangana n. sp. Turner 2006
Turner 2006
UA 8720 - holotype; paratypes: large nearly complete skull, incl. L & R hyoid, 1st 12 presacral vertebrae w/ribs, partial L & R scapulae, proximal end of R humerus, L ilium and proximal head of L femur, R femur w/o head, partial pubis & 3 damaged dorsal ribs, 3 damaged dorsal ribs, R coracoid, part of shaft of R humerus, L calcaneum & astragalus (FMNH PR 2297); Partial skull, damaged R mandible, R jugal, L radius & ulna, partial R radius, nearly complete vertebral column, sacrum, portions of L & R hindlimbs (FMNH PR 2298); Partial skull, L palpebral, L surangular & angular, R ilium, R dentary, R splenial, R angular, partial maxilla, dorsal and caudal vertebrae, dorsal ribs (FMNH PR 2299); dentaries (UA 8750, UA 8751, UA 8763, FMNH PR 2318); splenial (FMNH PR 2316); angular (UA 8754); premaxillae (FMNH PR 2334); maxillae (UA 8756, UA 8760, UA 8761, UA 8762, FMNH PR 2321); jugals (UA 8752, FMNH PR 2317, FMNH PR 2332); nasals (UA 8764); frontals (UA 8765, FMNH PR 2322); postorbital (FMNH PR 2315); palpebrals (UA 8757, FMNH PR 2336); cervicals (UA 8728, UA 8739, FMNH PR 2307, FMNH PR 2323); dorsals (UA 8722, UA 8723–8727, UA 8742, UA 8769–8773, UA 8774, FMNH PR 2304–2306); sacrals (UA 8767, FMNH PR 2301, FMNH PR 2330); caudals (UA 8729– 8731, UA 8735, FMNH PR 2308, FMNH PR 2309, FMNH PR 2330); dorsal ribs (UA 8732, UA 8733, UA 8755, FMNH PR 2334); chevrons (UA 8755); osteoderms (FMNH PR 2333); scapulae (UA 8779, FMNH PR 2313, FMNH PR 2334); coracoids (UA 8740, UA 8741, FMNH PR 2313); humeri (UA 8721, FMNH PR 2302, FMNH PR 2325–2327, FMNH PR 2340); radii (FMNH PR 2324, FMNH PR 2321, FMNH PR 2327, FMNH PR 2329, FMNH PR 2339); ulnae (UA 8758, FMNH PR 2324, FMNH PR 2326, FMNH PR 2327, FMNH PR 2339); pisiform (FMNH PR 2324); radialia (UA 8736, UA 8737, FMNH PR 2310, FMNH PR 2324, FMNH PR 2339); ulnaria (UA 8736, UA 8759, FMNH PR 2324, FMNH PR 2339); metacarpals (FMNH PR 2312, FMNH PR 2328); manual phalanges (FMNH PR 2312, FMNH PR 2328); manual unguals (FMNH PR 2328); ilia (FMNH PR 2301, FMNH PR 2303, UA 8768, FMNH PR 2330); ischium (FMNH PR 2330); pubes (UA 8776, UA 8777, UA 8778, FMNH PR 2330, FMNH PR 2331); femora (UA 8734, FMNH PR 2300, FMNH PR 2330, FMNH PR 2337); tibiae (FMNH PR 2300,FMNH PR 2330, FMNH PR 2335, FMNH PR 2337); fibulae (UA 8781, FMNH PR 2300, FMNH PR 2330, FMNH PR 2335); astragalus (FMNH PR 2330); metatarsals (UA 8736, UA 8780, FMNH PR 2319, FMNH PR 2330, FMNH PR 2337, FMNH PR 2338); pedal phalanges (UA 8743, UA 8744–UA 8746, FMNH PR 2311, FMNH PR 2319, FMNH PR 2330, FMNH PR 2337); pedal unguals: (UA 8736, FMNH PR 2330); unidentified phalanges (UA 8747–UA 8749, FMNH PR 2314); unidentified unguals (UA 8747–UA 8749, FMNH PR 2314).
Reptilia - Noasauridae
Masiakasaurus knopfleri Sampson et al. 2001
Carrano et al. 2011 1 specimen
FMNH PR 2217
Reptilia - Abelisauridae
Majungasaurus crenatissimus (Depéret 1896)
2 specimens
FMNH PR 2008, 2009
Amphibia - Temnospondyli
Anura indet. (Fischer von Waldheim 1813)
Asher and Krause 1998
UA 8677 (right angulosplenial)
Beelzebufo ampinga Evans et al. 2008
Evans et al. 2014 2 specimens
FMNH PR 2497 – cranial or vertebral spine table fragment; UA 8677 – partial right angulosplenial
see common names

Geography
Country:Madagascar State/province:Mahajanga
Coordinates: 15.9° South, 46.6° East (view map)
Paleocoordinates:30.1° South, 38.4° East
Basis of coordinate:stated in text
Altitude:150 meters
Geographic resolution:small collection
Time
Period:Cretaceous Epoch:Late/Upper Cretaceous
Stage:Maastrichtian 10 m.y. bin:Cretaceous 8
*Period:Late/Upper Cretaceous - Late/Upper Cretaceous *Epoch:Senonian - Senonian
*International age/stage:Campanian - Maastrichtian
Key time interval:Maastrichtian
Age range of interval:72.10000 - 66.00000 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Formation:Maevarano Member:Anembalemba
Stratigraphic resolution:bed
Stratigraphy comments: described as Campanian but now considered Maastrichtian
Lithology and environment
Primary lithology:yellow sandstone
Includes fossils?Y
Lithology description: "uppermost white sandstone unit" of the formation (now = Anembalemba Member)
Environment:"channel"
Geology comments: "shallow and very broad channel-belt system characterized by an ephemeral and flood-prone discharge regime."
Taphonomy
Modes of preservation:body
Degree of concentration:concentrated
Size of fossils:macrofossils
Preservation of anatomical detail:excellent
Articulated whole bodies:some
Associated major elements:some
Disassociated major elements:some
Disassociated minor elements:none
Fragmentation:occasional
Encrustation:none
Feeding/predation traces:arthropod boring
Temporal resolution:snapshot
Spatial resolution:autochthonous
Collection methods and comments
Collection methods:selective quarrying,mechanical,field collection
Reason for describing collection:taxonomic analysis
Collection method comments: many more fossils collected from this locality in 1993; none are listed in primary reference
Metadata
Database number:32366
Authorizer:M. Carrano, P. Mannion Enterer:M. Carrano, P. Mannion, J. Tennant
Modifier:M. Carrano Research group:vertebrate
Created:2003-06-26 14:33:50 Last modified:2019-11-07 17:18:44
Access level:the public Released:2003-06-26 14:33:50
Creative Commons license:CC BY
Reference information

Primary reference:
8731. S. D. Sampson, D. W. Krause, P. Dodson and C. A. Forster. 1996. The premaxilla of Majungasaurus (Dinosauria: Theropoda), with implications for Gondwanan paleobiogeography. Journal of Vertebrate Paleontology 16(4):601-605 [M. Carrano/M. Carrano/M. Carrano]

Secondary references:

37865 R. J. Asher and D. W. Krause. 1998. The first pre-Holocene (Cretaceous) record of Anura from Madagascar. Journal of Vertebrate Paleontology 18(4):696-699 [P. Mannion/P. Mannion]
44450 M. T. Carrano, M. A. Loewen, and J. J. W. Sertich. 2011. New materials of Masiakasaurus knopfleri Sampson, Carrano, and Forster, 2001, and implications for the morphology of the Noasauridae (Theropoda: Ceratosauria). Smithsonian Contributions to Paleobiology 95:1-53 [M. Carrano/M. Carrano/P. Mannion]
49974 S. E. Evans, J. R. Groenke, M. E. H. Jones, A. H. Turner, and D. W. Krause. 2014. New material of Beelzebufo, a hyperossified frog (Amphibia: Anura) from the Late Cretaceous of Madagascar. PLoS One 9(1):e87236 [P. Mannion/J. Tennant]
10609 D. W. Krause, S. E. Evans, and K.-Q. Gao. 2003. First definitive record of Mesozoic lizards from Madagascar. Journal of Vertebrate Paleontology 23(4):842-856 [M. Carrano/M. Carrano/M. Carrano]
24665ETE D. W. Krause, S. D. Sampson, M. T. Carrano and P. M. O'Connor. 2007. Overview of the history of discovery, taxonomy, phylogeny, and biogeography of Majungasaurus crenatissimus (Theropoda: Abelisauridae) from the Late Cretaceous of Madagascar. In S. D. Sampson & D. W. Krause (eds.), Majungasaurus crenatissimus (Theropoda: Abelisauridae) from the Late Cretaceous of Madagascar. Society of Vertebrate Paleontology Memoir 8. Journal of Vertebrate Paleontology 27(2, suppl.):1-20 [M. Carrano/M. Carrano/J. Alroy]
38067 D. W. Krause, J. J. W. Sertich, R. R. Rogers, S. C. Kast, A. H. Rasoamiaramanana and G. A. Buckley. 2010. Overview of the discovery, distribution, and geological context of Simosuchus clarki (Crocodyliformes: Notosuchia) from the Late Cretaceous of Madagascar. Society of Vertebrate Paleontology Memoir 10:4-12 [P. Mannion/P. Mannion]
32931ETE T. C. LaDuke, D. W. Krause, J. D. Scanlon and N. J. Kley. 2010. A Late Cretaceous (Maastrichtian) snake assemblage from the Maevarano Formation, Mahajanga Basin, Madagascar. Journal of Vertebrate Paleontology 30(1):109-138 [M. Carrano/M. Carrano]
23421ETE E. M. Roberts, R. R. Rogers, and B. Z. Foreman. 2007. Continental insect borings in dinosaur bone: examples from the Late Cretaceous of Madagascar and Utah. Journal of Paleontology 81(1):201-208 [M. Carrano/M. Carrano]
10610ETE R. R. Rogers, J. H. Hartman, and D. W. Krause. 2000. Stratigraphic analysis of Upper Cretaceous rocks in the Mahajanga Basin, northwestern Madagascar: implications for ancient and modern faunas. Journal of Geology 108:275-301 [M. Carrano/M. Carrano/M. Carrano]
28568ETE S. D. Sampson and L. M. Witmer. 2007. Craniofacial anatomy of Majungasaurus crenatissimus (Theropoda: Abelisauridae) from the Late Cretaceous of Madagascar. In S. D. Sampson & D. W. Krause (eds.), Majungasaurus crenatissimus (Theropoda: Abelisauridae) from the Late Cretaceous of Madagascar. Society of Vertebrate Paleontology Memoir 8. Journal of Vertebrate Paleontology 27(2, suppl.):32-102 [M. Carrano/M. Carrano]
37070 A. H. Turner. 2006. Osteology and phylogeny of a new species of Araripesuchus (Crocodyliformes: Mesoeucrocodylia) from the Late Cretaceous of Madagascar. Historical Biology 18(3):255-369 [P. Mannion/P. Mannion]