Pesciara (Alveolina dainelli Zone) (Eocene of Italy)

Also known as Monte Bolca

Where: Italy (45.6° N, 11.2° E: paleocoordinates 37.3° N, 10.3° E)

• coordinate based on nearby landmark

When: Alveolina dainelli foram zone, Ypresian (56.0 - 47.8 Ma)

• "Upper Ypresian (uppermost Lower or lowermost Middle Eocene)"

• group of beds-level stratigraphic resolution

Environment/lithology: lagoonal or restricted shallow subtidal; lithified limestone

• "shallow marine basin characterized by restricted circulation on the seafloor, exposed to pelagic, seagrass and coral reef biotic influences"
• "micritic limestone"

Size class: macrofossils

Preservation: mold/impression

Reposited in the FMNH

Collection methods: quarrying, mechanical,

• FMNH and Museo Civico di Storia Naturale, Verona collections

•"prepared using thin needles"

Primary reference: A. F. Bannikov and G. Carnevale. 2009. A new percoid fish from the Eocene of Monte Bolca, Italy: Hendrixella grandei gen. & sp nov. Swiss Journal of Geosciences 102(3):481-488 [J. Alroy/J. Alroy]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 94830: authorized by John Alroy, entered by John Alroy on 29.03.2010, edited by Mark Uhen

Creative Commons license: CC BY (attribution)

Taxonomic list

unclassified
  -
Chondrichthyes
 Rajiformes -
 Rajiformes - Rhinobatidae
Rhinobatus primaevus8 de Zigno 1874 guitarfish
Rhinobatus dezignoi8 Heckel 1853 guitarfish
 Myliobatiformes - Urolophidae
"Urolophus" crassicaudatus8 de Blainville 1818 round ray
 Myliobatiformes - Dasyatidae
"Dasyatis" dezignoi8 Molin 1861 whiptail stingray
"Dasyatis" muricata8 Volta 1796 whiptail stingray
 Myliobatiformes - Platyrhinidae
Platyrhina gigantea8 de Blainville 1818 stingray
Platyrhina egertoni8 de Zigno 1878 stingray
Platyrhina bolcensis8 Heckel 1851 stingray
 Myliobatiformes - Myliobatidae
Promyliobatis gazolae8 de Zigno 1885 eagle ray
 Euselachii -
Torpediniformes indet.8 electric ray
n. gen. n. sp.
 Torpediniformes - Narcinidae
Titanonarke molini8 Jaekel 1894 numbfish
 Carchariniformes - Carcharhinidae
Physogaleus cuvieri8 Agassiz 1835 requiem shark
Eogaleus bolcensis8 Capetta 1975 requiem shark
Alopiopsis plejodon8 Lioy 1865 requiem shark
 Orectolobiformes -
Mesiteia emiliae8 Gorjanovic-Kramberger 1886 carpet shark
Osteichthyes
  -
Protaulopsis bolcensis2 Woodward 1901
Actinopteri
 Anguilliformes - Chlopsidae
Whitapodus breviculus2 Agassiz 1835 eel
 Anguilliformes -
 Anguilliformes - Anguillidae
Eoanguilla leptoptera2 Agassiz 1835 eel
 Anguilliformes - Patavichthyidae
Patavichthys bolcensis2 Bassani 1897 eel
 Anguilliformes - Paranguillidae
Dalpiaziella brevicauda2 Cadrobbi 1962 eel
Paranguilla tigrina2 Agassiz 1839 eel
 Anguilliformes - Proteomyridae
Proteomyrus ventralis2 Agassiz 1839 eel
 Anguilliformes - Milananguillidae
Milananguilla lehmani2 Blot 1978 eel
 Anguilliformes - Ophichthidae
Goslinophis acuticaudus2 Agassiz 1835 eel
 Anguilliformes - Anguilloididae
Veronanguilla ruffoi2 Blot 1978 eel
 Anguilliformes -
Gazolapodus homopterus2 Blot 1980 eel
 Anguilliformes - Congridae
Voltaconger latispinus2 Agassiz 1835 conger eel
Paracongroides heckeli2 Blot 1978 conger eel
Bolcyrus bajai2 Blot 1978 conger eel
Bolcyrus formosissimus2 Eastman 1905 conger eel
 Teleostei -
Rhamphosus biserratus2 Bassani 1876
Ramphosus biserratus8 Bassani 1876
Ramphosus rastrum8 Volta 1796
Rhamphosus rastrum2 Volta 1796
 Dactylopteriformes - Pterygocephalidae
 Acanthomorphata -
Xiphopterus falcatus2 Agassiz 1835
 Lampridiformes - Veliferidae
Veronavelifer sorbinii2 Bannikov 1990
Velifer sp.8 Temminck and Schlegel 1850
n. sp.
Velifer carosii2 Carnevale and Bannikov 2014
 Lampridiformes -
"Pegasus" volans2 Volta 1796
 Lampridiformes - Bajaichthyidae
Bajaichthys elegans2 Sorbini 1983
 Acanthomorphata -
Pietschellus aenigmaticus2 Bannikov and Carnevale 2011
 Beloniformes - Exocoetidae
Rhamphexocoetus volans2 Bannikov et al. 1985
"Engraulis" evolans2 Blainville 1818
 Beloniformes - Hemiramphidae
Hemiramphus edwardsi2 Bassani 1876
 Atheriniformes - Rhamphognathidae
Rhamphognathus paralepoides2 Agassiz 1835 silverside
 Atheriniformes - Mesogasteridae
Mesogaster sphyraenoides2 Agassiz 1835 silverside
Latellagnathus teruzzii2 Bannikov 2008 silverside
 Atheriniformes - Atherinidae
Atherina ? macrocephala2 Woodward 1901 silverside
Atherina ? macrocephala8 Woodward 1901 silverside
 Acanthomorphata - Pomacentridae
Lorenzichthys olihan2 Bellwood 1999
Sorbinichromis francescoi n. gen. n. sp.3 Bannikov and Bellwood 2014
Palaeopomacentrus orphae n. gen. n. sp.7 Bellwood and Sorbini 1996
 Acanthomorphata - Menidae
Mene rhombea8 Volta 1796 moonfish
Mene oblonga8 Agassiz 1833 moonfish
 Carangiformes - Carangidae
Ceratoichthys pinnatiformis2 Blainville 1818
Seriola prisca2 Agassiz 1834
Eastmanalepes primaevus2 Eastman 1904
Lichia veronensis2 Bannikov and Sorbini 1990
Vomeropsis triurus2 Volta 1796
 Acanthomorphata - Sphyraenidae
Sphyraena bolcensis2 Agassiz 1844 barracuda
 Pleuronectiformes - Amphistiidae
Amphistium paradoxum8 Agassiz 1835 flatfish
 Pleuronectiformes -
Eobothus minimus8 Agassiz 1839 flatfish
Heteronectes chaneti8 Friedman 2008 flatfish
 Syngnathiformes - Syngnathidae
"Syngnathus" heckeli2 de Zigno 1874
"Syngnathus" bolcensis2 de Zigno 1887
 Syngnathiformes -
Calamostoma breviculum2 Agassiz 1835
 Syngnathiformes - Parasynarcualidae
 Syngnathiformes - Aulorhamphidae
Pesciarhamphus carnevalei2 Bannikov and Tyler 2011
Aulorhamphus bolcensis2 de Zigno 1887
Aulorhamphus chiarasorbiniae2 Bannikov and Tyler 2011
Aulorhamphus capellinii2 de Zigno 1887
 Syngnathiformes - Aulostomidae
"Macraulostomus veronensis" = Macroaulostomus veronensis2 Blot 1980
Eoaulostomus bolcensis2 Blainville 1818
"Jungensenichthys elongatus" = Jungersenichthys elongatus2 Blot 1980
Tyleria necopinata2 Parin 1993
 Syngnathiformes -
 Syngnathiformes - Paraeoliscidae
 Syngnathiformes - Fistularioididae
 Syngnathiformes - Solenostomidae
 Syngnathiformes - Urosphenidae
Urosphen dubius2 Blainville 1818
 Scombriformes - Scombridae
Auxides propterygius8 Agassiz 1835
Godsilia lanceolata8 Agassiz 1835
 Scombriformes - Centrolophidae
Zorzinia postalensis8 Bannikov 2000
 Scombriformes - Pomatomidae
Carangopsis brevis2 Blainville 1818
Carangopsis dorsalis2 Agassiz 1835
 Kurtiformes - Apogonidae
Apogonidae indet.8 Bonaparte 1846 Cardinalfish
Apogoniscus pauciradiatus2 Bannikov 2005 Cardinalfish
Eosphaeramia pygopterus2 Agassiz 1836 Cardinalfish
Eoapogon fraseri2 Bannikov 2005 Cardinalfish
Bolcapogon johnsoni2 Bannikov 2005 Cardinalfish
 Ophidiiformes - Ophidiidae
"Ophidium" voltianum2 Massalongo 1859
 Acanthomorphata - Leiognathidae
Eoleiognathus dorsalis2 Agassiz 1838
 Acanthuriformes - Zanclidae
Eozanclus brevirostris8 Agassiz 1835
 Acanthuriformes - Acanthuridae
Tauichthys aspesae8 Tyler and Bannikov 2000
Pesciarichthys punctatus8 Blot and Tyler 1990
Eorandallius elegans8 Blot and Tyler 1990
Eorandallius rectifrons8 Agassiz 1842
Gazolaichthys vestenanovae8 Blot and Tyler 1990
Proacanthurus tenuis8 Agassiz 1842
Proacanthurus bonatoi8 Blot and Tyler 1990
Proacanthurus ovalis8 Agassiz 1842
Proacanthurus elongatus8 Blot and Tyler 1990
Metaspisurus emmanueli8 Blot and Tyler 1990
Tylerichthys nuchalis8 Agassiz 1842
Tylerichthys milani8 Blot and Tyler 1990
Metacanthurus veronensis8 Blot and Tyler 1990
Protozebrasoma bloti8 Sorbini and Tyler 1998
Padovathurus gaudryi8 de Zigno 1887
"Pesciarichthys baldwinae n. sp." = Frigosorbinia baldwinae12 Sorbini and Tyler 1998
Acanthuroides massalongoi8 Blot and Tyler 1990
Lehmanichthys lessiniensis8 Blot and Tyler 1990
 Perciformes - Quasimullidae
Quasimullus sorbinii2 Bannikov 1999 perch-like fish
 Perciformes -
Parapygaeus polyacanthus8 Pellégrin 1907 perch-like fish
 Perciformes - Zorzinichthyidae
Zorzinichthys annae8 Tyler and Bannikov 2002 perch-like fish
 Perciformes - Massalongiidae
Massalongius gazolai8 Massalongo 1859 perch-like fish
 Perciformes - Acanthonemidae
Acanthonemus subaureus8 Blainville 1818 perch-like fish
 Perciformes - Sorbinipercidae
Sorbiniperca scheuchzeri8 Tyler 1999 perch-like fish
Sorbinicapros sorbiniorum8 Bannikov and Tyler 1999 perch-like fish
 Perciformes - Euzaphlegidae
Veronaphleges brunae8 Bannikov 2008 perch-like fish
 Perciformes - Palaeorhynchidae
Palaeorhynchus zorzini8 Fierstine et al. 2008 perch-like fish
 Perciformes - Tortonesidae
Tortonesia esilis2 Sorbini 1983 perch-like fish
 Perciformes -
"Labrus" valenciennesi2 Agassiz 1839 perch-like fish
Sorbinia caudopunctata8 Bellwood 1995 perch-like fish
 Perciformes - Blochiidae
Blochius macropterus8 de Zigno 1887 perch-like fish
Blochius longirostris8 Volta 1796 perch-like fish
 Perciformes - Carangodidae
Carangodes bicornis2 Volta 1796 perch-like fish
 Perciformes - Robertanniidae
Hendrixella grandei n. gen. n. sp. Bannikov and Carnevale 2009 perch-like fish
Hendrixella grandei2 Bannikov and Carnevale 2009 perch-like fish
Robertannia sorbiniorum2 Bannikov 2011 perch-like fish
 Perciformes - Callipterygidae
Callipteryx speciosus8 Agassiz 1839 perch-like fish
Callipteryx recticaudus8 Agassiz 1839 perch-like fish
 Perciformes -
Pavarottia lonardonii2 Bannikov and Zorzin 2011 perch-like fish
Squamibolcoides minciottii2 Bannikov and Zorzin 2013 perch-like fish
Jimtylerius temnopterus2 Agassiz 1836 perch-like fish
Bradyurus szajnochae2 de Zigno 1887 perch-like fish
Veronabrax schizurus2 Agassiz 1836 perch-like fish
Blotichthys coleanus2 Agassiz 1838 perch-like fish
Frippia labroiformis2 Bannikov and Carnevale 2012 perch-like fish
Gillidia antiqua2 Agassiz 1842 perch-like fish
Parapelates quindecimalis2 Agassiz 1836 perch-like fish
Pygaeus nobilis2 Agassiz 1842 perch-like fish
Pygaeus bolcanus2 Volta 1796 perch-like fish
Pygaeus nuchalis2 Agassiz 1842 perch-like fish
Malacopygaeus oblongus2 Agassiz 1838 perch-like fish
Frigoichthys margaritae2 Bannikov 2004 perch-like fish
Voltamulloides ceratorum2 Bannikov 2008 perch-like fish
Montepostalia annamariae2 Bannikov and Zorzin 2004 perch-like fish
Quasicichla mucistonaver8 Bannikov 2004 perch-like fish
 Perciformes - Exelliidae
Exellia velifer2 Agassiz 1842 perch-like fish
 Perciformes - Eocottidae
Bassanichthys pesciaraensis2 Bannikov 2004 perch-like fish
Eocottus veronensis2 Volta 1796 perch-like fish
 Perciformes - Ductoridae
Ductor vestenae2 Voltra 1796 perch-like fish
 Perciformes - Latidae
Eolates gracilis2 Agassiz 1833 perch-like fish
 Perciformes - Gobioidei
"Gobius" microcephalus8 Agassiz 1839 perch-like fish
 Acanthomorphata - Gerreidae
Aspesiperca ruffoi2 Bannikov 2008
 Acanthomorphata - Lutjanidae
Ottaviania mariae2 Sorbania 1983
Goujetia crassispina2 Agassiz 1839
Veranichthys ventralis2 Agassiz 1839
Lessinia sp.2 Bannikov and Zorzin 2014
Lessinia horrenda2 Bannikov and Zorzin 2014
 Acanthomorphata - Percichthyidae
Cyclopoma gigas2 Agassiz 1833
 Acanthomorphata - Siganidae
Ruffoichthys spinosus8 Sorbinia 1983
Ruffoichthys bannikovi8 Tyler and Sorbini 1990
Aspesiganus margaritae8 Bannikov and Tyler 2002
Acanthopygaeus agassizi8 Eastman 1904
 Spariformes - Sparidae
Abromasta microdon2 Agassiz 1839
Sparnodus vulgaris2 de Blainville 1818
Sparnodus elongatus2 Agassiz 1839
"Dentex" microdon2 Agassiz 1839
"Dentex" ventralis8 Agassiz 1839
Ellaserrata monksi2 Day 2002
 Lophiiformes - Brachionichthyidae
"Histionotophorus bassani" = Histionotophorus bassani9, "Histionotophorus bassanii" = Histionotophorus bassani2, Orrichthys longimanus n. gen. n. sp.9
"Histionotophorus bassani" = Histionotophorus bassani9 De Zigno 1887 anglerfish
"Histionotophorus bassanii" = Histionotophorus bassani2 De Zigno 1887 anglerfish
Orrichthys longimanus n. gen. n. sp.9 Carnevale and Pietsch 2010 anglerfish
 Lophiiformes - Lophiidae
Sharfia mirabilis2 Pietsch and Carnevale 2011 anglerfish
Sharfia mirabilis n. gen. n. sp.10 Pietsch and Carnevale 2011 anglerfish
Lophius brachysomus11 Agassiz 1835 anglerfish
Caruso brachysomus2 Agassiz 1835 anglerfish
 Lophiiformes - Ogcocephalidae
Tarkus squirei2 Carnevale and Pietsch 2011 anglerfish
 Lophiiformes - Antennariidae
Eophryne barbutii2 Carnevale and Pietsch 2009 anglerfish
 Labriformes - Labridae
Phyllopharyngodon longipinnis n. gen. n. sp.6, Bellwoodilabrus landinii n. gen. n. sp.5, Eocoris bloti n. gen. n. sp.1, Labrobolcus giorgioi n. gen. n. sp.4
Phyllopharyngodon longipinnis n. gen. n. sp.6 Bellwood 1990
Bellwoodilabrus landinii n. gen. n. sp.5 Bannikov and Carnevale 2010
Eocoris bloti n. gen. n. sp.1 Banikov and Sorbini 1990
Labrobolcus giorgioi n. gen. n. sp.4 Bannikov and Bellwood 2015
 Acanthomorphata - Priacanthidae
Pristigenys substriatus2 Blainville 1818
 Acanthomorphata - Scatophagidae
 Acanthomorphata - Acropomatidae
Acropoma lepidotum2 Agassiz 1836
 Tetraodontiformes - Aracanidae
Proaracana dubia2 Blainville 1818
 Tetraodontiformes - Ostraciidae
Eolactoria sorbinii2 Tyler 1975
 Tetraodontiformes - Eoplectidae
Eoplectus bloti2 Tyler 1975
 Tetraodontiformes - Bolcabalistidae
Bolcabalistes varii2 Tyler and Sorbini 1998
 Tetraodontiformes - Tetraodontidae
Eotetraodon tavernei2 Tyler and Bannikov 2013
Eotetraodon pygmaeus2 de Zigno 1887
 Tetraodontiformes - Diodontidae
Zignodon fornasieroae2 Tyler and Santini 2002
Prodiodon tenuispinus2 Agassiz 1844
Prodiodon erinaceus2 Agassiz 1844
Heptadiodon echinus2 Heckel 1854
 Tetraodontiformes - Protobalistidae
Spinacanthus cuneiformis2 Blainville 1818
Protobalistum imperiale2 Massalongo 1857
 Tetraodontiformes - Zignoichthyidae
Zignoichthys oblongus2 de Zigno 1874
 Tetraodontiformes - Triacanthidae
Protacanthodes nimesensis2 Tyler and Santini 2001
Protacanthodes ombonii8 de Zigno 1887
 Acanthomorphata - Monodactylidae
Psettopsis latellai2 Bannikov 2005
Psettopsis subarcuatus2 Blainville 1818
 Acanthomorphata - Caproidae
Eoantigonia veronensis8 Sorbini 1983
 Ephippiformes - Ephippidae
Eoplatax papilio2 Volta 1796
Archaephippus asper2 Volta 1796
 Holocentriformes - Holocentridae
 Acanthomorphata -
Oncolepis isseli2 Bassani 1897
 Aulopiformes - Paralepididae
Holosteus esocinus2 Agassiz 1835
 Clupeiformes - Clupeoidei
Platinx macropterus2 Blainville 1818 herring
Trollichthys bolcensis2 MarramĂ  and Carnevale 2014 hering
"Clupea" catopygoptera2 Woodward 1901 hering
 Ostariophysi -
Chanoides macropoma2 Agassiz 1835
Coelogaster leptostea2 Eastman 1905
"Chanos" forcipatus2 Heckel 1854
 Osteoglossiformes - Foreyichthyidae
Foreyichthys bolcensis2 Taverne 1979
 Osteoglossiformes -
Monopteros gigas2 Volta 1796
 Osteoglossiformes - Arapaimidae
 Gasterosteiformes - Centriscidae
Aeoliscoides longirostris2 Blainville 1818
Paramphisile weleri2 Blot 1980
 Pycnodontiformes - Pycnodontidae
Pycnodus apodus2 Volta 1796
Abdobalistum thyrsus2 Poyato-Ariza and Wenz 2002
 Pycnodontiformes - Nursallidae
Nursallia veronae2 Blot 1987
Palaeobalistum orbiculatum2 Blainville 1818