Ullujaya, Chilcatay Formation (Miocene of Peru)

Where: Peru (14.6° S, 75.6° W: paleocoordinates 15.5° S, 71.5° W)

• coordinate stated in text

When: Chilcatay Formation, Burdigalian (20.4 - 16.0 Ma)

• Chilcatay Formation, latest Oligocene to early Miocene (Dunbar et al. 1990; DeVries 1998, 2001). Although the precise locality of the holotype is unknown, the referred specimen MUSM 1401 originates from within the Chilcatay Formation, several meters under the Chilcatay Formation - Pisco Formation disconformity, at Ullujaya, a few kilometers north to the type locality. Based on a published section a short distance from Ullujaya, the Chilcatay Formation layers of this locality (which preserves the fossilized remains of other marine vertebrates including mysticetes, kentriodontids, the squalodelphinids Huaridelphis raimondii Lambert, Bianucci & Urbina, 2014 and Notocetus vanbenedeni Moreno, 1892, and a stem physeteroid) are tentatively dated to the early Miocene (Alván De la Cruz 2008; Lambert et al. 2014; Bianucci et al. 2015).

• bed-level stratigraphic resolution

Environment/lithology: marine; diatomaceous siltstone and sandstone

• See Di Celma et al 2018.

•The Chilcatay Formation comprises marine diatomaceous and tuffaceous sandy siltstones, bioclastic sandstones and, along the base, massive sandstones, and large boulders of crystalline basement rocks.

Size class: macrofossils

Primary reference: O. Lambert, C. de Muizon, and G. Bianucci. 2015. A new archaic homodont toothed cetacean (Mammalia, Cetacea, Odontoceti) from the early Miocene of Peru. Geodiversitas 37(1):79-108 [M. Uhen/M. Uhen/M. Uhen]more details

Purpose of describing collection: general faunal/floral analysis

PaleoDB collection 167732: authorized by Mark Uhen, entered by Mark Uhen on 30.03.2015

Creative Commons license: CC BY (attribution)

Taxonomic list

Gnathostomata
  -
Reptilia
 Testudines -
 Testudines - Dermochelyidae
Dermochelyidae indet.3 Lydekker 1889 leatherback
Mammalia
 Cetacea -
Odontoceti indet.1 Flower 1867 toothed whale
aff. Odontoceti indet.1 Flower 1867 toothed whale
 Cetacea - Squalodelphinidae
Huaridelphis raimondii3 Lambert et al. 2014 toothed whale
cf. Notocetus sp.2 Moreno 1892 toothed whale
Notocetus vanbenedeni3 Moreno 1892 toothed whale
Notocetus vanbenedeni2 Moreno 1892 toothed whale
 Cetacea - Kentriodontidae
Kentriodontidae indet.3 Slijper 1936 toothed whale
Kentriodon sp.1 Kellogg 1927 toothed whale
aff. Kentriodon pernix3 Kellogg 1927 toothed whale
 Cetacea -
Chilcacetus cavirhinus n. gen. n. sp. Lambert et al. 2015 toothed whale
Physeteroidea indet.3 Gray 1868 toothed whale
 Cetacea - Physeteridae
cf. Diaphorocetus sp.1 Ameghino 1894 sperm whale
Actinopteri
 Istiophoriformes - Istiophoridae
cf. Makaira sp.3 Lacépède 1802
aff. Makaira sp.1 Lacépède 1802
 Scombriformes - Scombridae
aff. Thunnus sp.1 South 1845
Chondrichthyes
 Elasmobranchii -
Elasmobranchii indet.3 elasmobranch
 Odontaspidida - Jaekelotodontidae
Anotodus agassizii1 Le Hon 1871 elasmobranch
 Euselachii -
Carcharhiniformes indet.3 Compagno 1973 ground shark
 Carcharhiniformes - Hemigaleidae
Hemigaleidae indet.3 ground shark
Hemipristis serra1 Agassiz 1835 ground shark
 Carcharhiniformes - Sphyrnidae
Sphyrnidae indet.3 Gill 1872 hammerhead shark
Sphyrna zygaena1 Linnaeus 1758 hammerhead shark
 Carcharhiniformes - Carcharhinidae
Carcharhinidae indet.3 Jordan and Evermann 1896 requiem shark
Physogaleus contortus1 Gibbes 1847 requiem shark
Negaprion brevirostris1 Poey 1868 lemon shark
Carcharhinus cf. leucas1 Valenciennes 1839 requiem shark
Carcharhinus brachyurus1 Gunther 1870 requiem shark
 Euselachii -
Lamniformes indet.3 Berg 1958 mackerel shark
 Lamniformes - Lamnidae
Lamnidae indet.3, Isurus oxyrinchus1, "Cosmopolitodus hastalis" = Carcharodon hastalis1, "Carcharocles chubutensis" = Carcharodon subauriculatus1
Lamnidae indet.3 Müller and Henle 1838 mackerel shark
Isurus oxyrinchus1 Rafinesque 1810 mackerel shark
"Cosmopolitodus hastalis" = Carcharodon hastalis1 Agassiz 1838 white shark
"Carcharocles chubutensis" = Carcharodon subauriculatus1 Agassiz 1839 white shark
 Lamniformes - Odontaspididae
"Odontaspidae indet." = Odontaspididae3
"Odontaspidae indet." = Odontaspididae3 Müller and Henle 1839 sand tiger shark
 Lamniformes - Alopiidae
Alopiidae indet.3 Bonaparte 1838 thresher shark
Alopias superciliosus1 Lowe 1840 thresher shark
 Lamniformes - Otodontidae
Otodontidae indet.3 mackerel shark
Megalolamna paradoxodon1 Shimada et al. 2017 mackerel shark
 Lamniformes - Carchariidae
Carcharias sp.1 Rafinesque 1810 sand shark
 Euselachii -
Rhinopristiformes indet.3 Last et al. 2016 elasmobranch
 Rhinopristiformes - Pristidae
Anoxypristis sp.1 White and Moy-Thomas 1941 sawfish
 Myliobatiformes -
Myliobatiformes indet.3 Compagno 1973 stingray