Santa Cruz Formation (Miocene of Argentina)

Where: Santa Cruz, Argentina (50.0° S, 70.0° W: paleocoordinates 50.6° S, 64.4° W)

• coordinate estimated from map

• basin-level geographic resolution

When: Santa Cruz Formation, Santacrucian (17.5 - 16.3 Ma)

Environment/lithology: terrestrial; lithology not reported

Size class: macrofossils

Primary reference: A. M. Albino. 1996. Snakes from the Miocene of Patagonia (Argentina) Part II: The Colubroidea. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen 200(3):353-360 [J. Head/J. Head/J. Alroy]more details

Purpose of describing collection: general faunal/floral analysis

PaleoDB collection 27313: authorized by Jason Head, entered by Jason Head on 01.12.2002, edited by Mark Uhen, Jelle Zijlstra, Miranta Kouvari and Edward Medeiros

Creative Commons license: CC BY (attribution)

Taxonomic list

Reptilia
 Squamata - Colubridae
Colubridae indet. Oppel 1811 colubrid snake
Aves
 Suliformes - Anhingidae
"Liptornis hesternus" = Anhinga hesterna5
"Liptornis hesternus" = Anhinga hesterna5 Ameghino 1894 darter
BMNH-A 509, cervical vertebra
 Pelecaniformes - Threskiornithidae
Protibis cnemialis5 Ameghino 1891 ibis
BMNH-A598, distal end of right tibiotarsus
 Sphenisciformes - Spheniscidae
"Palaeospheniscus antarcticus" = Paraptenodytes antarcticus13
"Palaeospheniscus antarcticus" = Paraptenodytes antarcticus13 Moreno and Mercerat 1891 penguin
 Rheiformes - Rheidae
Opisthodactylus patagonicus5 Ameghino 1891 rhea
BMNH-A 586, distal tarsometatarsi and phalanges; BMNH-A 587, symphysis, distal tibiotarsi; BMNH-A 588, distal tarsometarsus fragments
Opisthodactylus patagonicus6 Ameghino 1891 rhea
MPM-PV 18903 and MPM-PV 18904 (postcranial elements; distal ends of left tibiotarsi)
 Tinamiformes - Tinamidae
Tinamidae gen6 tinamou
MPM-PV 18900, (postcranial elements; proximal end of left carpometacarpus)
 Anseriformes -
Brontornis burmeisteri13 Moreno and Mercerat 1891 waterfowl
 Anseriformes - Anatidae
Eoneornis australis5 Ameghino 1894 waterfowl
BMNH-A595, distal end of radius
Eutelornis patagonicus5 Ameghino 1894 waterfowl
BMNH-A596, distal end of humerus and proximal tibiotarsus
 Gruiformes - Phorusrhacidae
"Phororhacos longissimus" = Phorusrhacos longissimus13, "Darwinornis socialis n. sp." = Phorusrhacos longissimus13, "Darwinornis copei n. gen. n. sp." = Phorusrhacos longissimus13, "Owenornis affinis n. gen. n. sp." = Phorusrhacos longissimus13, "Mesembriornis quatrefragesi n. sp." = Phorusrhacos longissimus13, "Darwinornis zitteli n. sp." = Phorusrhacos longissimus13, "Owenornis lydekkeri n. sp." = Phorusrhacos longissimus13, "Stereornis rollieri n. gen. n. sp." = Phorusrhacos longissimus13, Psilopterus sp.6, "Psilopterus lemoinei" = Psilopterus lemoinei6, "Psilopterus australis n. sp." = Psilopterus lemoinei13, Psilopterus intermedius n. sp.13, Psilopterus communis n. gen. n. sp.13, "Patagornis bachmanni n. sp." = Psilopterus bachmanni13, "Patagornis marshi n. gen. n. sp." = Patagornis marshi13, "Palaeociconia cristata n. sp." = Patagornis marshi13
"Phororhacos longissimus" = Phorusrhacos longissimus13 Ameghino 1887 terror bird
"Darwinornis socialis n. sp." = Phorusrhacos longissimus13 Ameghino 1887 terror bird
"Darwinornis copei n. gen. n. sp." = Phorusrhacos longissimus13 Ameghino 1887 terror bird
"Owenornis affinis n. gen. n. sp." = Phorusrhacos longissimus13 Ameghino 1887 terror bird
"Mesembriornis quatrefragesi n. sp." = Phorusrhacos longissimus13 Ameghino 1887 terror bird
"Darwinornis zitteli n. sp." = Phorusrhacos longissimus13 Ameghino 1887 terror bird
"Owenornis lydekkeri n. sp." = Phorusrhacos longissimus13 Ameghino 1887 terror bird
"Stereornis rollieri n. gen. n. sp." = Phorusrhacos longissimus13 Ameghino 1887 terror bird
Psilopterus sp.6 Moreno and Mercerat 1891 terror bird
MPM-PV 18901 (postcranial elements; fragments of tibiotarsus, tarsometatarsus)
"Psilopterus lemoinei" = Psilopterus lemoinei6 Moreno and Mercerat 1891 terror bird
MPM-PV 18897 (postcranial elements including long-bone fragments)
"Psilopterus australis n. sp." = Psilopterus lemoinei13 Moreno and Mercerat 1891 terror bird
Psilopterus intermedius n. sp.13 Moreno and Mercerat 1891 terror bird
Psilopterus communis n. gen. n. sp.13 Moreno and Mercerat 1891 terror bird
"Patagornis bachmanni n. sp." = Psilopterus bachmanni13 Moreno and Mercerat 1891 terror bird
"Patagornis marshi n. gen. n. sp." = Patagornis marshi13 Moreno and Mercerat 1891 terror bird
BMNH-A516, partial skeleton (ref. 58178)
"Palaeociconia cristata n. sp." = Patagornis marshi13 Moreno and Mercerat 1891 terror bird
originally stated to be from the Eocene, but almost certainly Miocene
 Gruiformes -
Anisolornis excavatus5 bird
BMNH-A 594, distal tarsometatarsus
Mammalia
 Notoungulata - Hegetotheriidae
Hegetotherium mirabile4 Ameghino 1887 notoungulate
AMNH 9159, cranium and mandible; YPM-VPPU 15298, mandible, tibia; 15395, tibia
 Notoungulata - Interatheriidae
Protypotherium sp.4 Ameghino 1882 notoungulate
MACN-A 9657–63, humerus
Protypotherium praeutilium15 notoungulate
P12058
Protypotherium australe4 Moreno 1882 notoungulate
AMNH 9149, humerus, ulna, femur, tibia
Protypotherium australe15 Moreno 1882 notoungulate
YPM-PU 15203, 15350, 15551
"Interatherium robustum" = Interatherium extensus4 Fernández et al. 2023 notoungulate
YPM-VPPU 15100, cranium, mandible; 15108, humerus, ulna, tibia; 15554, mandible; 15401, cranium, mandible, humerus, ulna, femur, tibia
 Notoungulata - Homalodotheriidae
Homalodotherium cunninghami4 Flower 1884 notoungulate
YPM-VPPU 15747, ulna
 Notoungulata - Toxodontidae
Adinotherium sp.4, Adinotherium sp.15, "Adinotherium ovinum" = Adinotherium ovinum15, "Adinotherium robustum" = Adinotherium ovinum4, Nesodon sp.4, Nesodon imbricatus4
Adinotherium sp.4 Ameghino 1887 notoungulate
AMNH 9532, cranium
Adinotherium sp.15 Ameghino 1887 notoungulate
P13229, P13232, P13245
"Adinotherium ovinum" = Adinotherium ovinum15 Owen 1846 notoungulate
P12066, P13098, P13104, P13107, P13108, P13110, P13231, UC1508, YPM-PU 16033
"Adinotherium robustum" = Adinotherium ovinum4 Owen 1846 notoungulate
AMNH 9497, cranium
Nesodon sp.4 Owen 1846 notoungulate
AMNH 9128, cranium, mandible
Nesodon imbricatus4 Owen 1846 notoungulate
AMNH 9192, mandible, humerus, ulna; YPM-VPPU 15132, ulna; 15141, cranium; 15256, cranium, mandible, humerus, ulna; 15260, mandible
 Primates - Homunculidae
"Homunculus patagonicus" = Homunculus patagonicus8, "Anthropops perfectus n. gen. n. sp." = Homunculus patagonicus8
"Homunculus patagonicus" = Homunculus patagonicus8 Ameghino 1891 monkey
MLP 66-V-2-2, mandibular fragment with i2, c, p2-3 roots, broken p4, broken m1; MLP 11-121, left dp4-m1, unerupted p4
"Anthropops perfectus n. gen. n. sp." = Homunculus patagonicus8 Ameghino 1891 monkey
MACN-A 635, symphysis with roots of left and right i1-2 and right p2-3, alveoli for left and right canines, right p4 in situ, holotype. Ref. 58226 state that the specimen comes from one of the coastal localities of the Santa Cruz Fm; ref. 47173 lists it as from near the city of Río Gallegos
 Rodentia -
Acarechimys sp.2 Patterson 1965 caviomorph
MACN-A 1895
Acarechimys minutissimus2 Ameghino 1887 caviomorph
MACN-A 256 (syntype of Acaremys minutissimus) MACN-A 257 (syntype of Acaremys minutissimus) MACN-A 258 (syntype of Acaremys minutissimus), MLP 15-1, MLP 15-1, MACN-A 4089, MACN-A 4088, MACN-A 4079 MACN-A 4080 MACN-A4081 MACN-A 4083 MACN-A 4084 MACN-A 4085 MACN-A 4086 MACN-A 4087, MACN-A 4076 MACN-A 4077
Acarechimys minutus2 Patterson and Pascual 1967 caviomorph
MACN-A 237 (syntype of Acaremys minutus) MACN-A 238 (syntype of Acaremys minutus), MACN-A 4082, , MLP 82-XII-1-31
 Rodentia - Abrocomidae
"Acarechimys constans" = Ameghinomys constans2
"Acarechimys constans" = Ameghinomys constans2 Ameghino 1887 chinchilla rat
MLP 15-57 (syntype of Stichomys constans), MACN-A 247 (syntype of Stichomys constans)
 Panameriungulata - Proterotheriidae
Tetramerorhinus fortis n. sp.4 Ameghino 1894 placental
MACN-A 5971, cranium, type
Diadiaphorus majusculus4 Ameghino 1887 placental
AMNH 9196, mandible; 9291, cranium, mandible
 Cingulata - Peltephilidae
Peltephilus pumilus16 Ameghino 1887 edentate
MACN-A 7784–7791, skull, mandible, postcrania
 Cingulata - Chlamyphoridae
Proeutatus sp.16 Ameghino 1891 edentate
MACN-A 1063, humerus; MLP 69-IX-8-11, near-complete skeleton
Stenotatus sp.16 Ameghino 1891 edentate
MLP 69-IX-5-11, femur
Stenotatus patagonicus16 Ameghino 1887 edentate
MACN-A 898–900, osteoderms
Stenotatus hesternus16 Ameghino 1889 edentate
MACN-A 871–878, osteoderms
Prozaedyus proximus16 Ameghino 1887 edentate
MACN-A 2201, partial carapace
 Cingulata - Propalaehoplophoridae
Propalaehoplophorus australis16 Moreno 1882 edentate
MLP 16–15, skull, mandible, skeleton
Metopotoxus laevatus16 Ameghino 1889 edentate
MACN-A 1042, fragment of carapace
Cochlops muricatus16 Ameghino 1889 edentate
MACN-A 2120, mandible, right femur
 Theriamorpha -
Argyromanis patagonicus n. gen. n. sp.12, Orthoarthrus mixtus n. gen. n. sp.12
Argyromanis patagonicus n. gen. n. sp.12 edentate
MACN A-11687, left astragalus
Orthoarthrus mixtus n. gen. n. sp.12 Ameghino 1904 edentate
MLP 69-IX-5-16, left astragalus
 Theriamorpha - Myrmecophagidae
Myrmecophagidae indet.3, "Protamandua rothi n. gen. n. sp." = Protamandua rothi12, "Promyrmephagus euryarthrus n. gen. n. sp." = Protamandua rothi12, "Promyrmephagus dolichoarthrus n. sp." = Protamandua rothi12
Myrmecophagidae indet.3 edentate
MLP 69-IX-8-8a, left humerus
"Protamandua rothi n. gen. n. sp." = Protamandua rothi12 Ameghino 1904 edentate
MACN-A 10901b, calcaneum
"Promyrmephagus euryarthrus n. gen. n. sp." = Protamandua rothi12 Ameghino 1904 edentate
MACN-A 11530, astragalus
"Promyrmephagus dolichoarthrus n. sp." = Protamandua rothi12 Ameghino 1904 edentate
MLP ? right astragalus (now lost), cast = MACN A-4621b
 Megatherioidea - Megalonychidae
Eucholoeops ingens3 Ameghino 1887 edentate
MACN-A 11614, left half of skull and dentary
 Sparassodonta - Borhyaenidae
Prothylacynus patagonicus9, "Borhyaena tuberata" = Borhyaena tuberata14, "Borhyaena zitteli n. sp." = Borhyaena tuberata7, Arctodictis munizi7
Prothylacynus patagonicus9 Ameghino 1891 metatherian
BM(NH) M8075, partial maxilla with teeth; MACN 11540, dentary left and right m2-4
"Borhyaena tuberata" = Borhyaena tuberata14 Ameghino 1887 metatherian
Ref. 58202: YPM-VPPU 15701, from "South of Río Coyle". Ref. 58303: MACN A 6238, partial skull, mandible, and postcrania; 9342-3, jaw fragments; 9344, skull; MACN unnumbered, maxillary fragment; MACN unnumbered, both hemimandibles
"Borhyaena zitteli n. sp." = Borhyaena tuberata7 Ameghino 1887 metatherian
MACN A 6238, skull fragment, holotype
Arctodictis munizi7 Mercerat 1891 metatherian
MLP 11-65, skull and mandible; MLP 11-81, left m4; MLP 11-89, right m2
 Sparassodonta -
Cladosictis patagonica10 Ameghino 1887 metatherian
MLP 11-100, dentary p2; MLP 11-13, dentary p1-m4; MLP 11-63, dentary m1-4; MLP 11-68, dentary p1; MLP 11-87, maxilla M2-3
Sipalocyon gracilis10 Ameghino 1887 metatherian
MLP 11-105, c and m1-2; MLP 11-76, dentary m2; MLP 11-77, dentary m3; MLP 11-41, dentary p1, p3
Acyon tricuspidatus14 Ameghino 1887 metatherian
MACN-A 9
 Microbiotheria - Microbiotheriidae
Microbiotherium tehuelchum n. sp.11 Ameghino 1887 marsupial
MLP 11-36, right dentary with p2-m, neotype; PU 15038, maxilla M2-3, dentary c-m3, postcrania, skull fragment; MLP 11-35, dentary p2-m3; MLP 58-IX-3-4, partial skull
 Paucituberculata - Abderitidae
Abderites meridionalis1 Ameghino 1887 marsupial
MACN-A 2031, dentary m2; 2032, dentary m2-4; 2033, dentary m2-4; 2034, dentary m1-3; MLP 11-109, dentary m1-2; 11-133, dentary m2-3; 55-XII-13-144, dentary m2; MACN 11651, dentary p3-m4