SH-20, Khodzhakul (Cretaceous of Uzbekistan)

Also known as Khodja-Kull; SKh-20, CX-20

Where: Navoi, Uzbekistan (40.1° N, 65.8° E: paleocoordinates 35.9° N, 60.5° E)

• coordinate based on nearby landmark

• small collection-level geographic resolution

When: Khodzhakul Formation, Early/Lower Cenomanian (99.6 - 93.5 Ma)

• lower/middle part of the formation. Upper Albian in Averianov (2001), but Early Cenomanian in more recent papers.

• bed-level stratigraphic resolution

Environment/lithology: coastal; pebbly, calcareous sandstone and pebbly, sandy claystone

• "limonite pebbles lying amongst a suite of calcareous sandstones and sandy clays"

Size classes: macrofossils, mesofossils

Collected by A. D. Arkhangelsky, L. A. Nessov in 1914; 1978, 1981–1982, 1985

Primary reference: A. N. Riabinin. 1931. Ostatki dinozavrov iz verkhnego mela nizov'ev r. Amu-Dar'i [On the dinosaurian remains from the Upper Cretaceous of the lower parts of the Amu-Daria River]. Zapiski Rossiyskogo Mineralogicheskogo Obshchestva 60(1):114-118 [M. Carrano/M. Carrano]more details

Purpose of describing collection: general faunal/floral analysis

PaleoDB collection 58208: authorized by Matthew Carrano, entered by Kaitlin Maguire on 27.01.2006, edited by Roger Benson, Matthew Carrano, John Alroy, Matthew Clapham and Philip Mannion

Creative Commons license: CC BY (attribution)

Taxonomic list

unclassified
  -
Equisetopsida
 Pinidae - Pinidae
Araucariaceae indet.11 Henckel and Hochstetter 1865
Angiospermae
  -
 Coniferales -
Cephalopoda
 Ammonoidea -
Gastropoda
  -
Gastropoda indet.14 Cuvier 1795 snail
Bivalvia
  -
Bivalvia indet.14 Linnaeus 1758 clam
Deuteropoda
  -
Malacostraca indet.14 Latreille 1802
Chondrichthyes
 Hybodontiformes - Distobatidae
Glickmanodus rarus n. gen. n. sp.11
Glickmanodus rarus n. gen. n. sp.11 Nessov et al. 1997 elasmobranch
including holotype: CCMGE 677/12457
 Hybodontiformes - Polyacrodontidae
Polyacrodus torosus11 elasmobranch
Polyacrodus ex gr. illingworthi11 Dixon 1850 elasmobranch
 Euselachii -
 Lamniformes - Anacoracidae
"Eoanacorax dalinkevichiusi" = Squalicorax dalinkevichiusi14
"Eoanacorax dalinkevichiusi" = Squalicorax dalinkevichiusi14 Glikman and Shvazhaite 1971 crow shark
 Lamniformes - Paraisuridae
Paraisurus sp.14 Glikman 1957 mackerel shark
 Holocephali -
 Chimaeriformes - Callorhynchidae
? Elasmodus sp.1 Egerton plownose chimaera
Osteichthyes
 Acipenseriformes - Polyodontidae
Polyodontidae indet.11 Bonaparte 1838 paddlefish
Hesperopsephurus kyatensis n. gen. n. sp.12 Nessov 1997 paddlefish
including holotype CCMGE 32/12177
Amphibia
 Salientia -
 Salientia - Pelobatidae
? Pelobatidae indet.6 Lataste 1879 spadefoot toad
 Caudata - Urodela
"Mynbulakia nongratis" = Eoscapherpeton6, Horezmia gracile n. gen. n. sp.5
"Mynbulakia nongratis" = Eoscapherpeton6 hellbender
Horezmia gracile n. gen. n. sp.5 Nessov 1981 salamander
ZIN PHA 77-2
Mammalia
 Theriamorpha - Bobolestidae
Bobolestes zenge n. gen. n. sp.8
Bobolestes zenge n. gen. n. sp.8 Nessov 1985 eutherian
from "Khodzhakul, SKH-20"; CCMGE 2/12176
 Theriamorpha - Zhelestidae
cf. Sheikhdzheilia rezvyii2 Averianov and Archibald 2005 eutherian
from "Khodzhakul, SKH-20"
 Theriamorpha - Pappotheriidae
Pappotheriidae sp.10 Slaughter 1965 metatherian
Teeth
Reptilia
 Testudines - Adocidae
Adocus kizylkumensis5 Nesov 1981 turtle
 Testudines - Lindholmemydidae
"Mongolemys cf. occidentalis" = Khodzhakulemys occidentalis7
"Mongolemys cf. occidentalis" = Khodzhakulemys occidentalis7 Nessov 1984 turtle
 Testudines - Carettochelyidae
Kizylkumemys sp.7 Nessov 1977 turtle
 Testudines - Nanhsiungchelyidae
Nanhsiungchelyidae indet.6 Yeh 1966 turtle
 Testudines - Macrobaenidae
cf. Kirgizemys sp.7, Oxemys gutta n. gen. n. sp.7
cf. Kirgizemys sp.7 Nessov and Khozatsky 1973 turtle
Oxemys gutta n. gen. n. sp.7 Nessov 1977 turtle
TsNIGR Museum, no. 2/11478
 Testudines -
"Chelonia indet." = Testudines14
"Chelonia indet." = Testudines14 Batsch 1788 turtle
 Squamata -
Iguania indet.4 Cuvier 1817 squamates
CMGE 40/12000 (frontal), 41/12000 (partial parietal)
Gekkota indet.11 Cuvier 1817 squamates
Oxia karakalpakensis n. gen. n. sp.9 Nessov 1985 lizard
Dentaries: CMGE 28/12000 (holotype; Nessov 1985); 29/12000, 43/12000, 44/12000
Hodzhakulia magna n. gen. n. sp.9 Nessov 1985 worm lizard
Maxillae: CMGE30/12000 (holotype, right), 31/12000 (left); right dentaries: 32/12000, 32a/12000
Varanoidea indet.6 Gray 1827 squamates
Scincomorpha indet.4 squamates
CMGE 42/12000 (partial right dentary)
 Loricata -
Crocodylia indet.14 crocodilian
 Ornithischia - Ankylosauridae
? Shamosaurinae indet.11 Tumanova 1983 ankylosaurid
also listed as Shamosauridae
 Ornithischia -
Kulceratops kulensis n. gen. n. sp.11, ? Gilmoreosaurus atavus n. sp.11
Kulceratops kulensis n. gen. n. sp.11 Nessov 1995 ceratopsian
CCMGE 495/12457
? Gilmoreosaurus atavus n. sp.11 Nessov 1995 ornithopod
CCMGE 576/12457
 Ornithischia - Hypsilophodontidae
? Hypsilophodontidae indet.11 hypsilophodontid
 Ornithischia - Hadrosauridae
Hadrosauridae indet. Cope 1869 hadrosaurid
 Theropoda - Coeluridae
Coeluridae indet.11 Marsh 1881 coelurid
 Theropoda - Dromaeosauridae
Richardoestesia asiatica15, ""Laelaps" cf. explanatus" = Dromaeosaurus explanatus11
Richardoestesia asiatica15 Nessov 1995 maniraptoran
ZIN PH 1222/16
""Laelaps" cf. explanatus" = Dromaeosaurus explanatus11 Cope 1876 maniraptoran
"possibly this is a peculiar species of Alectrosaurus"
 Theropoda -
Paronychodon asiaticus15 Nessov 1995 maniraptoran
ZIN PH 1071/16
Aves
  -
Aves indet.14 Linnaeus 1758 bird
 Gruiformes -
Horezmavis eocretacea n. gen. n. sp.13
Horezmavis eocretacea n. gen. n. sp.13 Nessov and Borkin 1983 bird
Reptilia
 Saurischia -
Titanosauria indet.3 Bonaparte and Coria 1993 sauropod
"Diplodocidae or Titanosauridae"; teeth: ZIN PH 2428/16, 2429/16, 2430/16
Actinopteri
 Holostei -
Holostei indet.6 Müller 1846
 Holostei - Dapediidae
"Dapedius sp." = Dapedium6
"Dapedius sp." = Dapedium6 Leach 1822
 Lepisosteiformes -
Lepidotes sp.6 Agassiz 1832 gar
 Aulopiformes - Enchodontidae
Enchodontidae indet.9 Woodward 1901
 Aulopiformes - Eurypholidae
Eurypholis sp.11 Pictet 1850
 Ichthyodectiformes -
Sultanuvaisia antiqua6 Nessov 1981
 Aspidorhynchiformes - Aspidorhynchidae