CBI-14, Dzharakuduk II (Cretaceous to of Uzbekistan)

Also known as Dzarakuduk; Dzhara-Kuduk, Dzhirakuduk, Dzhyrakuduk, Kul’beke

Where: Navoi, Uzbekistan (42.1° N, 62.7° E: paleocoordinates 36.8° N, 57.0° E)

• coordinate based on nearby landmark

• small collection-level geographic resolution

When: Bissekty Formation, Middle Turonian to Middle Turonian (93.5 - 89.3 Ma)

• Approx. 57 m above the base of the formation

• bed-level stratigraphic resolution

Environment/lithology: fluvial; sandstone and conglomerate

• Formation comprises medium-grained, poorly lithified, cross-bedded fluvial sandstones and clast-supported, well-cemented infraformational conglomerates

Size classes: macrofossils, mesofossils

Collected by L. A. Nessov and Uzbek-Russian-British-American-Canadian (URBAC) expedition in 1974–1994, 1997–2006; reposited in the USNM

Collection methods: surface (float),

Primary reference: L. A. Nessov. 1985. Redkie kostnye ryby, nazemiye yashcheritsy i mlekopitayushchie zony limanov i primorskikh nizmennostey mela Kyzylkumov [Rare bony fishes, terrestrial lizards, and mammals in the estuarine and coastal lowland zone of the Kyzyl-Kum Cretaceous]. Yearbook of the All-Union Paleontological Association 28:199-219 [M. Carrano/M. Carrano]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 58168: authorized by Matthew Carrano, entered by Kaitlin Maguire on 25.01.2006, edited by Matthew Carrano, Roger Benson, Jonathan Tennant and Philip Mannion

Creative Commons license: CC BY (attribution)

Taxonomic list

unclassified
  -
Actinopteri
 Acipenseriformes - Polyodontidae
Psephuroides kazakhorum n. gen. n. sp.14
Psephuroides kazakhorum n. gen. n. sp.14 Nessov 1997 paddlefish
including holotype
 Teleostei - Aspidorhynchidae
Belonostomus aciculiferus n. sp. Nessov 1985
TsNIGR 12/12000
 Teleostei -
Elopomorpha indet.13
TsNIGR 12/12000
 Albuliformes - Albulidae
Albula lapidosa Nessov 1985 bonefish
 Ichthyodectiformes - Saurodontidae
Asiamericana asiatica n. gen. n. sp.13
Asiamericana asiatica n. gen. n. sp.13 Nessov 1995
 Ichthyodectiformes - Ichthyodectidae
 Amiiformes - Amiidae
Amia limosa Nessov 1985 bowfin
 Lepisosteiformes - Lepisosteidae
Osteichthyes
 Salientia - Gobiatidae
Gobiates tatarinovi n. sp.17 Ročec and Nessov 1993 frog
LU-N6/405
Gobiates asiaticus n. sp.17 Ročec and Nessov 1993 frog
LU-N6/370 (holotype partial squamosal)
Gobiates dzhyrakudukensis n. sp.17 Ročec and Nessov 1993 frog
LU-N6/341 (holotype maxilla), 357
Gobiates kizylkumensis n. sp.17 Ročec and Nessov 1993 frog
LU-N6/363 (holotype partial squamosal)
Gobiatoides parvus n. gen. n. sp.17 Ročec and Nessov 1993 frog
LU-N6/344 (holotype partial right maxilla)
 Salientia - Discoglossidae
Aralobatrachus robustus17 Nessov 1981 painted frog
LU-N6/353, 367, 409, 435, 314 (maxilla), 299 (urostyle), 343, 419, 427, 296 (humeri),
Saevesoederberghia egredia n. gen. n. sp.17 Ročec and Nessov 1993 painted frog
LU-N6/375 (holotype partial maxilla)
Estesina elegans17 Ročec and Nessov 1993 painted frog
LU-N6/335 (maxilla)
cf. Kizylkuma sp.17 Nessov 1981 painted frog
LU-N6/319, 322, 423 (humeri)
Kizylkuma antiqua17 Nessov 1981 painted frog
LU-N6/271, 332, 398 (maxilla)
Procerobatrachus paulus n. gen. n. sp.17 Ročec and Nessov 1993 painted frog
LU-N6/412 (holotype partial maxilla)
 Caudata - Urodela
Nesovtriton mynbulakensis n. gen. n. sp.18
Nesovtriton mynbulakensis n. gen. n. sp.18 Skutschas 2009 salamander
ZIN PH 1/85, 5/49, 3/85, 4/85, 5/85, 8/85
Reptilia
 Testudines - Pantrionychidae
Khunnuchelys kizylkumensis5 Brinkman et al. 1993 turtle
CCMGE 303/12458, 8a/12458
 Saurischia -
Titanosauria indet.22 Bonaparte and Coria 1993 sauropod
 Theropoda -
Theropoda indet.13 theropod
 Avetheropoda - Dromaeosauridae
Dromaeosauridae indet.13 Colbert and Russell 1969 coelurosaur
Richardoestesia asiatica n. sp.20 Nessov 1995 coelurosaur
Multiple teeth
Itemirus medullaris21 Kurzanov 1976 coelurosaur
ZIN PH 2338/16, 2352/16, 2353/16, 2351/16, 90/16, 13/16, 2269/16, 182/16
 Avetheropoda - Troodontidae
Euronychodon asiaticus n. sp.13 Nessov 1995 coelurosaur
 Avetheropoda -
Paronychodon asiaticus n. sp.20 Nessov 1995 coelurosaur
Multiple teeth
Aves
 Ornithurae -
? Neornithes indet.6 bird
PO 3434a
Ichthyornis minusculus n. sp.12 Nessov 1990 bird
PO 3941
 Alexornithiformes - Alexornithidae
Sazavis prisca n. gen. n. sp.16, "Ichthyornis maltshevskyi n. sp." = Lenesornis maltshevskyi10
Sazavis prisca n. gen. n. sp.16 Nessov 1989 bird
PO 3472
"Ichthyornis maltshevskyi n. sp." = Lenesornis maltshevskyi10 Nessov 1986 bird
PO 3434
 Enantiornithes -
Enantiornis martini n. sp.8 Nessov and Panteleyev 1993 bird
PO 4609
Reptilia
 Avetheropoda -
Tyrannosauroidea indet.4 Walker 1964 coelurosaur
ZIN PH 1036/16, 1034/16, 1033/16, 1039/16, 684/16, 708/16, 756/16, 734/16, 120/16
? Alectrosaurus sp.13 Gilmore 1933 coelurosaur
"Segnosauria indet." = Therizinosauroidea13 Russell and Dong 1994 coelurosaur
 Ornithischia -
Ornithischia indet.13 ornithischian
 Neornithischia -
Turanoceratops tardabilis13 Nessov et al. 1989 ornithischian
One element - CCGME 628/12457 (partial braincase) - now referred to Titanosauria indet.
 Neornithischia - Ceratopsidae
? Ceratopsidae indet.13 Marsh 1888 ceratopsid
 Neornithischia -
Levnesovia transoxiana19 Sues and Averianov 2009 bird foot dinosaur
CCMGE 565/12457, 10/12454, 595/12176
 Neornithischia - Hadrosauridae
Hadrosauridae indet.13 Cope 1869 hadrosaurid
Cionodon kyzylkumensis13 hadrosaurid
 Squamata -
Buckantaus crassidens n. gen. n. sp.
Buckantaus crassidens n. gen. n. sp. Nessov 1985 squamates
TsNIGR 26/12000
 Plesiosauria -
Mammalia
 Tribosphenida -
Sazlestes tis n. gen. n. sp.14
Sazlestes tis n. gen. n. sp.14 Nessov 1997 eutherian
including holotype (ZIN C.82553)
 Tribosphenida - Kennalestidae
Eoungulatum kudukensis n. gen. n. sp.15 Nessov et al. 1998 eutherian
CCMGE 2/12455 (holotype left M2)
cf. Eoungulatum kudukensis15 Nessov et al. 1998 eutherian
CCMGE 16/12953 (left M3), 17/12953 (right M2), 18/12953 (left M3)
 Ungulata - Periptychidae
Periptychidae indet.13 Cope 1882 condylarth
 Tribosphenida -
"Paranyctoides aralensis n. sp." = Paranyctoides quadrans2, "Paranyctoides aralensis" = Paranyctoides quadrans3
"Paranyctoides aralensis n. sp." = Paranyctoides quadrans2 Nessov 1982 eutherian
"Paranyctoides aralensis" = Paranyctoides quadrans3 Nessov 1982 eutherian
ZIN C.83131 (left dentary fragment)
 Tribosphenida - Zalambdalestidae
Kulbeckia kulbecke1 Nessov 1993 eutherian
 Tribosphenida - Zhelestidae
"Zhelestidae" indet.2, "Sorlestes budan n. gen. n. sp." = Zhelestes temirkazyk9, Parazhelestes robustus n. gen. n. sp.15, "Parazhelestes minor" = Parazhelestes mynbulakensis15, "Aspanlestes aptap" = Aspanlestes aptap2, "? Zhelestes bezelgen n. sp." = Aspanlestes aptap11
"Zhelestidae" indet.2 Nessov 1990 eutherian
incl. 1 gen. et sp. nov.
"Sorlestes budan n. gen. n. sp." = Zhelestes temirkazyk9 Nessov 1985 eutherian
CCMGE 3/12176 and extra specimens listed by Nessov et al 1998
Parazhelestes robustus n. gen. n. sp.15 Nessov 1993 eutherian
CCMGE 70/12455 (holotype left maxilla), CMGE 10/12953 (left M1), 10/12953 (right M2)
"Parazhelestes minor" = Parazhelestes mynbulakensis15 Nessov 1985 eutherian
sp. nov.
"Aspanlestes aptap" = Aspanlestes aptap2 Nessov 1985 eutherian
"? Zhelestes bezelgen n. sp." = Aspanlestes aptap11 Nessov 1985 eutherian
CCMGE 1/12455
 Theriiformes -
Theria indet.2 Parker and Haswell 1897 mammal
 Deltatheroida - Deltatheridiidae
"Sulestes karakshi" = Sulestes karakshi3, "Sulestes karakshi" = Sulestes karakshi3, "? Deltatheroides kizylkumensis" = Sulestes karakshi2
"Sulestes karakshi" = Sulestes karakshi3 Nessov 1985 metatherian
URBAC 00-001, left m2; URBAC 00-057, right dentary fragment; URBAC 03-009, left m2; URBAC 03-043, right dentary fragment; URBAC 03-090, left dentary fragment; URBAC 03-104, left dentary fragment; URBAC 04-149, left maxilla fragment; URBAC 04-164, left maxilla fragment; URBAC 04-169, right m3; URBAC 04-217, left m4; URBAC 04-291, right m2; URBAC 04-344, left m2; URBAC 06-006, right dentary; URBAC 06-019, right m3; URBAC 98–012, right m3; URBAC 99-017, left M2; ZIN 88467, right dentary fragment; URBAC03-222, right dentary; ZIN 89008, left dentary fragment; ZIN 90311, right dentary fragment; CCMGE 5/12455, left dentary fragment; ZIN 90312, left maxilla fragment; CCMGE 40/12000, right maxilla; ZIN 90309, left dentary fragment; ZIN 90310, right dentary fragment; URBAC 04-346, left m4
"Sulestes karakshi" = Sulestes karakshi3 Nessov 1985 metatherian
"? Deltatheroides kizylkumensis" = Sulestes karakshi2 Nessov 1985 metatherian
 Multituberculata -
Uzbekbaatar kizylkumensis n. gen. n. sp.7
Uzbekbaatar kizylkumensis n. gen. n. sp.7 Kielan-Jaworowska and Nessov 1992 multituberculate
including CCMGE 100/ 12455, right p4 (holotype) and specimens referred by Averianov & Archibald (2006); four other cimolodontan multituberculate specimens from this locality may or may not be referrable