Kamenny Ravine (Ulema River), Isheevo, PIN 157, 2207 (Permian to of Russian Federation)

Also known as Kamennyi, Kamenny Valley, Yumrala Ravine, Sviaga River, Sviyaga River, Isheevo, Tartarstan, Kamenny Ovrag

Where: Tatarstan, Russian Federation (55.1° N, 48.5° E: paleocoordinates 29.3° N, 43.3° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Titanophoneus potens other zone, Urzhumian to Urzhumian (268.0 - 265.0 Ma)

• Isheevo Subassemblage of the Isheevo Assemblage; the later part of the Early Tatarian (=Urzhumian–early Severodvinian) (Golubev 2000; Benton 2012, Geology).

• group of beds-level stratigraphic resolution

Environment/lithology: fluvial; red sandstone and red mudstone

• "..depositing in a shallow branch or pool with slow current...Concentration of organic matters produced the brown tints of the sands.."
• fine reddish sands. "...complete absence of coarse sediments of pebbletype." Overlying red mudstones are unfossiliferous.

Size class: macrofossils

Collected by I. A. Efremov in 1934, 1935; reposited in the PIN

Collection methods: quarrying,

• "[...] we have employed the method of laying vast (up to 250 sq. mt.) grounds [...] for exposing the bone-bearing layer. This method proved to be quite satisfactory, as during the two seasons of excavating works (of 1934 and 1935) we have succeeded in obtaining of two practically complete skeletons of carnivorous Deinocephalians and of one skeleton of a herbivorous Deinocephalian. Unfortunately, this latter was embedded at the boundary of the layer of bone-bearing sands, just under the train of deluvial deposits, and was distorted by the landslip and subterranian waters to such an extent, that in spite of the extremely accurate preparation performed, we have succeeded to pick out from the matrix rock only the skull with a few vertebrae, and some ribs." (Efremov, 1940).

Primary reference: J. A. Efremov. 1940. Predvaritel'noye opisaniye novykh form permskoy i triasovoy fauny nazemnykh pozvonochnykh SSSR [Preliminary description of new forms from the Permian and Triassic terrestrial vertebrate fauna of the USSR]. Trudy Paleontologicheskogo Instituta Akademiy Nauk SSSR 10(2):1-140 [J. Alroy/R. Whatley/M. Carrano]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 27931: authorized by John Alroy, entered by Robin Whatley on 17.01.2003, edited by Richard Butler and Torsten Liebrecht

Creative Commons license: CC BY (attribution)

Taxonomic list

unclassified
  -
Chondrichthyes
 Polyspondyli - Onchidae
Onchus sp.5 Agassiz 1837 shark
 Hybodontiformes - Hybodontidae
Hybodus sp.5 Agassiz 1837 elasmobranch
Actinopteri
 Palaeonisciformes - Platysomidae
Platysomus sp.5 Agassiz 1843
 Palaeonisciformes - Acrolepidae
Acrolepis sp.5 Agassiz 1843
Osteichthyes
 Temnospondyli - Archegosauridae
"Melosaurus paucidens n. sp." = Tryphosuchus paucidens3
"Melosaurus paucidens n. sp." = Tryphosuchus paucidens3 Konzhukova 1955 tetrapod
PIN 157/107 (type)
Reptilia
 Procolophonia - Lanthanosuchidae
Lanthanosuchus watsoni n. gen. n. sp.2
Lanthanosuchus watsoni n. gen. n. sp.2 Efremov 1946 parareptile
PIN 271/1 (type), 157/91-98, 209
 Bolosauria - Bolosauridae
Permotriturus herrei n. gen. n. sp.3
Permotriturus herrei n. gen. n. sp.3 Tatarinov 1968 parareptile
PIN 157/500 (type)
 Gorgodontia - Ulemosauridae
Ulemosaurus svijagensis n. gen. n. sp.1
Ulemosaurus svijagensis n. gen. n. sp.1 Riabinin 1938 reptile
PIN 2207/2, skull
 Gorgodontia - Anteosauridae
"Titanophoneus potens n. gen. n. sp." = Titanophoneus potens, "Doliosaurus yanschinovi n. gen. n. sp." = Titanophoneus potens1, "Cliorhizodon efremovi n. sp." = Syodon biarmicum1
"Titanophoneus potens n. gen. n. sp." = Titanophoneus potens Efremov 1940 reptile
PIN 157/1 (type). PIN 157/180, a frontal. PIN 157/181, a frontal. PIN 157/182, a postorbital. PIN 157/186, transverse process of a pterygoid. PIN 157/187, a prefrontal. PIN 157/188, a postorbital. PIN 157/190, transverse process of a pterygoid. PIN 157/192, a postorbital. PIN 157/193, a pterygoid. PIN 157/194, a pterygoid. PIN 157/195, transverse process of a pterygoid. PIN 157/208, a partial squamosal. PIN 157/209, an isolated angular boss. PIN 157/272, an angular. PIN 157/275, a pterygoid. PIN 157/648, a palatine. PIN 157/972, a pterygoid.
"Doliosaurus yanschinovi n. gen. n. sp." = Titanophoneus potens1 Efremov 1940 reptile
PIN 157/3 (type)
"Cliorhizodon efremovi n. sp." = Syodon biarmicum1 Kutorga 1838 reptile
PIN 157/2, skull, PIN 157/635, palatine, PIN 157/670, quadrate, PIN 157/677, mandible, PIN 157/1045, partial periotic, PIN 157/1046, partial periotic
Osteichthyes
 Therapsida -
"Venjukovia invisa n. sp." = Ulemica invisa
"Venjukovia invisa n. sp." = Ulemica invisa Efremov 1940 therapsid
 Therocephalia - Lycosuchidae
? Porosteognathus sp.4 Vjuschkov 1952 therapsid
PIN 157/866, 957, 958, 959
Porosteognathus efremovi n. gen. n. sp.4 Vjuschkov 1952 therapsid
PIN 157/19 (type), parietals; 157/20, 21, 22, 24, 497, 498, 683, 684, 685, 686, 687, 688
 Seymouriamorpha -
Enosuchus breviceps n. gen. n. sp.3
Enosuchus breviceps n. gen. n. sp.3 Konzhukova 1955 tetrapod
PIN 271/99 (type)