Hangu Shales - Samana Range, Pakistan (Paleocene to of Pakistan)

Where: North-West Frontier, Pakistan (33.5° N, 71.1° E: paleocoordinates 0.1° N, 64.0° E)

• local area-level geographic resolution

When: Hangu Shale Member (Hangu Formation), Selandian to Selandian (61.6 - 56.0 Ma)

• The Hangu Shales are separated from underlying Albian rocks by about 500 ft of barren lithographic limestones and sandstones, the upper 140 ft of which constitute the Hangu Sandstones, which is conformable with the Hangu Shale, but unconformable with the Albian rocks. The shales are overlain conformably by 200 ft of limestone (the Lockhart Limestone). Refined age data obtained from Warwick et al. (2003) and references therein, which suggest a middle to late Paleocene age based on Pollen.

• formation-level stratigraphic resolution

Environment/lithology: marginal marine; lithified shale

• No specific environmental data presented in Cox (1932), but depositional environments in Warwick et al. (2003) suggest a marginal marine to shallow subtidal environment for the upper part of the Hangu Formation (terrestrial for the lower part).
• Shale. No more specific description than this

Size class: macrofossils

Collection methods: Collection made by Lt-Col. L.M. Davies during military service. Collection is derived from many sections along the continuous outcrop of Hangu Shale.

Primary reference: L. R. Cox. 1930. The fossil fauna of the Samana Range and some neighbouring areas: Part VIII. The Mollusca of the Hangu Shales. Memoirs of the Geological Suvery of India 14 [A. Miller/A. Hendy/A. Hendy]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 52132: authorized by Austin Hendy, entered by Austin Hendy on 26.07.2005

Creative Commons license: CC BY (attribution)

Taxonomic list

• Somewhat antiquated nomenclature, but many taxa carrying species level determinations and significant discussion of taxonomic relationships.
unclassified
  -
Gastropoda
 Ptenoglossa - Vasidae
Eovasum haimei d'Archiac and Haime 1854 snail
 Cerithioidea - Pachychilidae
Pirena torta n. sp.
Pirena torta n. sp. Cox 1930 snail
 Cerithioidea - Turritellidae
"Mesalia fasciata" = Sigmesalia fasciata, Mesalia pagoda n. sp., "Turritella ranikoti" = Kapalmerella ranikoti, Turritella lahirii n. sp., Turritella cf. hollandi, Turritella daviesi n. sp.
"Mesalia fasciata" = Sigmesalia fasciata Lamarck 1804 turret shell
Mesalia pagoda n. sp. Cox 1930 turret shell
"Turritella ranikoti" = Kapalmerella ranikoti Vredenburg 1928 turret shell
Turritella lahirii n. sp. Cox 1930 turret shell
Turritella cf. hollandi Cossmann and Pissarro 1909 turret shell
Turritella daviesi n. sp. Cox 1930 turret shell
 Cerithioidea - Batillariidae
Batillaria coronata n. sp. Cox 1930 snail
Pyrazus octogonus n. sp. Cox 1930 snail
 Architaenioglossa - Ampullinidae
Globularia brevispira n. sp. Cox 1930 snail
Globularia sulcata n. sp. Cox 1930 snail
Pachycrommium indicum n. sp. Cox 1930 snail
 Sorbeoconcha - Campanilidae
Campanile brookmani n. sp. Cox 1930 snail
 Sorbeoconcha - Strombidae
Rimella levis n. sp., Tibia samanaensis n. sp., "Terebelliopsis (Semiterebellum) cf. lanceolatum" = Tibia (Africoterebellum), Lambis goniophora
Rimella levis n. sp. Cox 1930 true conch
Tibia samanaensis n. sp. Cox 1930 true conch
"Terebelliopsis (Semiterebellum) cf. lanceolatum" = Tibia (Africoterebellum) true conch
Lambis goniophora Bellardi 1852 conch
 Neogastropoda - Buccinidae
Strepsidura tipperi n. sp. true whelk
Strepsidura scobina n. sp. true whelk
 Neogastropoda - Fasciolariidae
Streptochetus foxi n. sp. Cox 1930 snail
Dolicholatirus indicu n. sp. snail
 Neogastropoda - Melongenidae
Hemifusus montensis n. sp. Cox 1930 snail
 Neogastropoda - Clavatulidae
Turricula (Pleurofusia) hanguensis n. sp. Cox 1930 snail
 Neogastropoda - Muricidae
Murex (Phyllonotus) sandersoni n. sp. Cox 1930 murex snail
Murex (Phyllonotus) wadiai n. sp. Cox 1930 murex snail
 Neogastropoda - Volutidae
Voluta (Voluta) vredenburgi n. sp. Cox 1930 volute
Lyria samanaensis n. sp. Cox 1930 volute
Athleta (Volutocorbis) daviesi n. sp. Cox 1930 volute
"Athleta (Volutospina) wynnei n. sp." = Athleta (Volutocorbis) wynnei Cox 1930 volute
 Sorbeoconcha - Eocypraeidae
"Eocypraea cotteri n. sp." = Eocypraea (Eocypraea) cotteri
"Eocypraea cotteri n. sp." = Eocypraea (Eocypraea) cotteri Cox 1930 snail
 Sorbeoconcha - Cypraeidae
"Protocypraea kohatica n. sp." = Bernaya (Protocypraea) kohatica
"Protocypraea kohatica n. sp." = Bernaya (Protocypraea) kohatica Cox 1930 cowry
 Sorbeoconcha - Naticidae
Euspira roei n. sp. Cox 1930 moon snail
Euspirocrommium conicum n. sp. Cox 1930 moon snail
 Sorbeoconcha - Vermetidae
Vermetus hanguensis n. sp. Cox 1930 worm shell
 Heterostropha - Architectonicidae
Architectonica mainwaringi n. sp. Cox 1930 sundial
Bivalvia
 Ostreida - Ostreidae
Ostrea sp. Linnaeus 1758 oyster
 Pholadida - Corbulidae
"Corbula (Bicorbula) praeexarata n. sp." = Bicorbula, "Corbula samanaensis n. sp." = Corbula (Bicorbula) samanaensis
"Corbula (Bicorbula) praeexarata n. sp." = Bicorbula clam
"Corbula samanaensis n. sp." = Corbula (Bicorbula) samanaensis Cox 1930 clam
 Cardiida - Trapezidae
Trapezium daviesi n. sp. Cox 1930 clam
 Cardiida - Veneridae
Meretrix indica n. sp. Cox 1930 venus clam
 Cardiida - Cardiidae
Cardium inaequiconvexum Cossmann and Pissarro 1927 cockle
 Carditida - Carditidae
"Cardita hanguensis n. sp." = Venericardia jaquinoti
"Cardita hanguensis n. sp." = Venericardia jaquinoti d'Orbigny 1843 clam
numerous specimens
 Carditida - Crassatellidae
Crassatellites exiguus n. sp. Cox 1930 clam
Scaphopoda
 Dentaliida - Dentaliidae
Dentalium hanguense n. sp. Cox 1930 tusk shell