PPP# 168 - Escudo de Veraguas [Escudo de Veraguas Fm] (Pliocene of Panama)

Also known as NMB Loc. 17621; 87-09-01

Where: Bocas, Panama (9.1° N, 81.5° W: paleocoordinates 9.8° N, 81.1° W)

• coordinate stated in text

• outcrop-level geographic resolution

When: Escudo de Veraguas Formation (Bocas del Toro Group), Piacenzian (3.6 - 2.6 Ma)

• STRATIGRAPHIC RELATIONS: The upper and lower contacts of the formation are not exposed. BIOSTRATIGRAPHY: from Collins et al. (1999), planktic foraminifera suggest 1.8 Ma near the top and 3.5 Ma at the base. PPP dataset indicates an age range between 3.6 and 3.5 Ma.

• bed-level stratigraphic resolution

Environment/lithology: deep-water; poorly lithified, burrowed, bioturbated, fine-grained, argillaceous sandstone and poorly lithified, burrowed, bioturbated, argillaceous siltstone

• ENVIRONMENT: from Collins et al. 1999 (p. 94), mixed middle neritic and outer nertic/upper bathyal, estimated depth of 125 m.
• GENERAL LITHOLOGY: from Collins et al. (1999), lower part of formation at the stratotype is moderately indurated, fine, silty sandstone and clayey siltstone, prevasively bioturbated and containing frequent, cemented, irregular burrow-concentrations and horizons with dense thalassinoid burrow systems.

Size class: macrofossils

Reposited in the BMNH, NMB, USNM

Collection methods: bulk, sieve,

• COVERAGE: Exhaustive for gastropoda, bivalvia and scaphopoda. COLLECTORS: unknown (PPP staff), 8/7/87. REPOSITORIES:collections presumably held at USNM, BMNH and Naturhistorisches Museum Basel (NMB),

Primary reference: L. S. Collins and A. G. Coates. 1999. A paleobiotic survey of Caribbean faunas from the Neogene of the Isthmus of Panama. Bulletins of American Paleontology (357) [A. Miller/A. Hendy/A. Hendy]more details

Purpose of describing collection: paleoecologic analysis

PaleoDB collection 61336: authorized by Austin Hendy, entered by Austin Hendy on 05.06.2006

Creative Commons license: CC BY (attribution)

Taxonomic list

• NOMENCLATURE: modern nomenclature, though only identified to subgenus resolution. IDENTIFIER: presumably identified by J. Jackson, J. Todd, H. Fortunato, and P. Jung.
Show authors, comments, and common names
unclassified
  -
? Turbonillidae indet., Turbonillidae indet.
Scaphopoda
 Gadilida - Entalinidae
Bathoxiphus sp.
Bathoxiphus sp. Pilsbry 1897 tusk shell
 Gadilida - Gadilidae
Gadilopsis sp.
Gadilopsis sp. Woodring 1925 tusk shell
 Dentaliida - Dentaliidae
Dentalium sp.
Dentalium sp. Linnaeus 1758 tusk shell
sensu-lato
Gastropoda
 Cocculinoidea - Cocculinidae
? Cocculina sp.
? Cocculina sp. Dall 1882 snail
 Epitonioidea - Epitoniidae
Epitonium (Asperiscala) sp.
Epitonium (Asperiscala) sp. de Boury 1909 wentletrap
 Vanikoroidea - Eulimidae
Eulimidae indet., Scalenostoma sp.
Eulimidae indet. Philippi 1853 snail
Scalenostoma sp. Deshayes 1863 snail
 Pseudomelanoidea - Pseudomelaniidae
? Crassispira (? Crassispira) sp., ? Lioglyphostoma sp., ? Compsodrillia sp., Miraclathurella sp.
? Lioglyphostoma sp. Woodring 1928 snail
? Compsodrillia sp. Woodring 1928 snail
Miraclathurella sp. Woodring 1928 snail
 Sorbeoconcha - Naticidae
Naticidae indet., "Natica (Naticarius) sp." = Naticarius
Naticidae indet. Guilding 1834 moon snail
"Natica (Naticarius) sp." = Naticarius Dumeril 1806 moon snail
 Sorbeoconcha - Atlantidae
Atlanta sp.
Atlanta sp. Lesueur 1817 snail
 Sorbeoconcha - Triphoridae
Triphoridae indet.
Triphoridae indet. Gray 1847 snail
 Sorbeoconcha - Rissoidae
Rissoidae indet., Alvania sp.
Rissoidae indet. Gray 1847 snail
Alvania sp. Risso 1826 snail
 Sorbeoconcha - Cassidae
Sconsia sp.
Sconsia sp. Gray 1847 snail
 Neogastropoda - Olividae
Oliva (Oliva) sp., Olivella (Minioliva) sp., Olivella (Macgintiella) sp.
Oliva (Oliva) sp. Bruguière 1789 olive snail
Olivella (Minioliva) sp. Olsson 1956 olive snail
Olivella (Macgintiella) sp. Olsson 1956 olive snail
 Neogastropoda - Fasciolariidae
Fusinus sp.
Fusinus sp. Rafinesque 1815 snail
 Neogastropoda - Buccinidae
Metula sp., Antillophos (Antillophos) sp.
Metula sp. Adams and Adams 1853 true whelk
Antillophos (Antillophos) sp. Woodring 1928 true whelk
 Neogastropoda - Nassariidae
Nassarius sp.
Nassarius sp. Duméril 1806 snail
sensu-lato
 Neogastropoda - Columbellidae
Cotonopsis (Cotonopsis) sp.
Cotonopsis (Cotonopsis) sp. Olsson 1942 snail
 Neogastropoda - Marginellidae
Marginellidae indet., Volvarina sp., ? Dentimargo sp.
Marginellidae indet. Fleming 1828 margin shell
Volvarina sp. Hinds 1844 margin shell
? Dentimargo sp. Cossmann 1899 margin shell
 Neogastropoda - Muricidae
Haustellum sp., Typhinae indet.
Haustellum sp. Schumacher 1817 murex snail
Typhinae indet. Cossmann 1903 murex snail
 Neogastropoda - Cystiscidae
? Persicula sp.
? Persicula sp. Schumacher 1817 snail
 Neogastropoda - Terebridae
Strioterebrum sp.
Strioterebrum sp. Sacco 1891 auger snail
 Neogastropoda - Conidae
Conus sp.
Conus sp. Linnaeus 1758 cone shell
 Neogastropoda - Mangeliidae
Ithycythara sp., Brachycythara sp., Vitricythara sp.
Ithycythara sp. Woodring 1928 snail
Brachycythara sp. Woodring 1928 snail
Vitricythara sp. Fargo 1953 snail
 Neogastropoda - Turridae
Polystira sp.
Polystira sp. Woodring 1928 turrid
 Neogastropoda - Drilliidae
Agladrillia (Agladrillia) sp., Agladrillia (Eumetadrillia) sp.
Agladrillia (Agladrillia) sp. Woodring 1928 snail
Agladrillia (Eumetadrillia) sp. Woodring 1928 snail
 Neogastropoda - Clathurellidae
? Glyphostoma sp.
? Glyphostoma sp. Gabb 1872 snail
 Heterostropha - Pyramidellidae
Pyramidellidae indet., Eulimella (Ebalina) sp.
Pyramidellidae indet. Gray 1840 snail
Eulimella (Ebalina) sp. Thiele 1929 snail
 Heterostropha - Architectonicidae
Architectonica sp., Pseudotorinia sp.
Architectonica sp. Röding 1798 sundial
Pseudotorinia sp. Sacco 1892 snail
 Heterostropha - Acteonidae
Acteon sp.
Acteon sp. Montfort 1810 snail
 Trochoidea - Trochidae
Calliostoma sp., Microgaza sp.
Calliostoma sp. Swainson 1840 top snail
Microgaza sp. Dall 1881 top snail
 Opisthobranchia - Retusidae
Sulcoretusa sp., Volvulella (Paravolvulella) sp., Volvulella (Volvulella) sp.
Sulcoretusa sp. Burch 1945 snail
Volvulella (Volvulella) sp. Newton 1891 snail
 Opisthobranchia - Cylichnidae
Acteocina sp.
Acteocina sp. Gray 1847 barrel-bubble
 Thecosomata - Cavoliniidae
Cavolinia sp., Styliola sp.
Cavolinia sp. Avilgaard 1791 snail
Styliola sp. Gray 1850 snail
Bivalvia
 Nuculanida - Nuculanidae
Saccella sp.
Saccella sp. Woodring 1925 pointed nut clam
 Nuculida - Nuculidae
Nucula (Lamellinucula) sp., Leionucula sp.
Nucula (Lamellinucula) sp. Schenck 1944 nut clam
Leionucula sp. Quenstedt 1930 nut clam
 Poromyida - Verticordiidae
Trigonulina sp., Haliris (Haliris) sp.
Trigonulina sp. d'Orbigny 1946 clam
Haliris (Haliris) sp. Dall 1886 clam
 Poromyida - Cuspidariidae
Cuspidaria (Cuspidaria) sp.
 Cardiida - Kelliellidae
Kelliella sp.
Kelliella sp. Sars 1865 clam
 Cardiida - Montacutidae
Mysella (Mysella) sp.
Mysella (Mysella) sp. Angas 1877 clam
 Cardiida - Tellinidae
"Eurytellina sp." = Tellina (Eurytellina)
"Eurytellina sp." = Tellina (Eurytellina) Fischer 1887 tellin clam
 Cardiida - Semelidae
Abra sp.
Abra sp. Lamarck 1818 abra
 Pectinida - Pectinoidae
Amusium sp.
Amusium sp. Röding 1798 scallop
 Arcida - Arcidae
Anadara (Rasia) sp.
Anadara (Rasia) sp. Gray 1857 ark
Anthozoa
 Scleractinia - Caryophylliidae
Asterosmilia profunda
Asterosmilia profunda Duncan 1864 stony coral