Vallipón, Castellote (Cretaceous of Spain)

Where: Aragón, Spain (40.8° N, 0.3° W: paleocoordinates 30.6° N, 9.7° E)

• coordinate estimated from map

• local area-level geographic resolution

When: Artoles Formation, Early/Lower Barremian (130.0 - 125.5 Ma)

• base of this formation (original name translates as "Artoles Marls and Limestones")

•Villanueva-Amadoz et al. 2015: The presence of cf. Arecipites microfoveolatus Ibrahim, Afropollis sp. and Stellatopollis barghoornii Doyle from the basal levels of Artoles Formation suggests a late Barremian age (Pons et al. 2013). The schizacean fern spore content supports a Barremian age.

•Bover-Arnal et al. 2016: Sr values from samples suggest a middle/late Early Barremian age for the lower-middle part of the Artoles Formation, and a Late Barremian age for the upper part. Previously the Barremian–Aptian boundary has been placed within the Artoles Formation, but late Barremian ammonites have been found in the overlying Morella Formation, ruling out an early Aptian age for the Artoles Formation.

• member-level stratigraphic resolution

Environment/lithology: estuary or bay; lithified, calcareous sandstone and conglomerate

• "deposited in an intertidal or subtidal environment very close [related[ to the coastline"
• Quartitic sandstones and conglomerates. "The rock is dissolved with 10% formic acid" before screenwashing.

Size classes: macrofossils, mesofossils

Preservation: trace, coprolite

Collected in 1995

Collection methods: bulk, chemical, sieve,

• Materials include bones, coprolites, and gastroliths. Some preparation using formic acid.

Primary reference: J. I. Ruiz-Omeñaca, J. I. Canudo, and G. Cuenca-Bescós. 1996. Dientes de dinosaurios ornitópodos en el Barremiense superior de Castellote y Mas de las Matas (Teruel) [Ornithopod dinosaur teeth in the Upper Barremian of Castellote and Mas de las Matas (Teruel)]. Tomo Estraordinario, 125 Aniversario de la RSEHN. Real Sociedad Española de Historia Natural, Madrid 315-319 [M. Carrano/M. Carrano/M. Carrano]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 28196: authorized by Matthew Carrano, entered by Matthew Carrano on 28.01.2003, edited by Kaitlin Maguire, Roger Benson, Philip Mannion and Evangelos Vlachos

Creative Commons license: CC BY (attribution)

Taxonomic list

Gastropoda
  -
Gastropoda indet.12 Cuvier 1795 snail
Bivalvia
  -
Bivalvia indet.12 Linnaeus 1758 clam
Gnathostomata
  -
Chondrichthyes indet. Huxley 1880 cartilaginous fish
Chondrichthyes
 Hybodontiformes - Polyacrodontidae
Polyacrodus sp.9 Jaekel 1889 elasmobranch
Polyacrodus brevicostatus9 Patterson 1966 elasmobranch
"? Hybodus parvidens" = Polyacrodus parvidens9 Woodward 1916 elasmobranch
 Hybodontiformes - Lonchidiidae
"Lissodus aff. crenulatus" = Lonchidion crenulatum9
"Lissodus aff. crenulatus" = Lonchidion crenulatum9 Patterson 1966 elasmobranch
 Hybodontiformes -
Lissodus sp.9 Brough 1935 elasmobranch
Lissodus breve9 elasmobranch
 Synechodontiformes - Orthacodontidae
Orthacodontidae indet.9 de Beaumont 1960 elasmobranch
 Rajiformes -
Pseudohypolophus mcnultyi9 Thurmond 1971 guitarfish
misspelled "mcnultyui"
 Rajiformes - Rhinobatidae
"Rhinobatos cf. halteri" = Spathobatis halteri9
"Rhinobatos cf. halteri" = Spathobatis halteri9 Biddle and Landemaine 1988 guitarfish
 Lamniformes - Odontaspididae
Odontaspididae indet.9 Müller and Henle 1839 sand tiger shark
 Lamniformes - Eoptolamnidae
Eoptolamna eccentrolopha n. gen. n. sp.8
Eoptolamna eccentrolopha n. gen. n. sp.8 Kriwet et al. 2008 mackerel shark
Amphibia
 Salientia -
Mammalia
 Symmetrodonta -
Symmetrodonta indet.6 symmetrodont
MPZ 96/180
 Multituberculata - Pinheirodontidae
? Lavocatia sp.9 Canudo and Cuenca-Bescós 1996 multituberculate
MPZ 96/165, 96/175
 Multituberculata - Eobaataridae
"Eobaatarinae indet." = Eobaataridae6, Eobaatar sp.1, ? Eobaatar pajaronensis1
"Eobaatarinae indet." = Eobaataridae6 Kielan-Jaworowska et al. 1987 multituberculate
MPZ 96/166–173
Eobaatar sp.1 Kielan-Jaworowska et al. 1987 multituberculate
MPZ 96/156–159, MPZ2010/859–862; Three P1/3s... a ?P5 of Eobaatar may also belong to this species... other teeth such as the three upper incisors and upper and lower first and second molars may also belong to this species of Eobaatar
? Eobaatar pajaronensis1 Hahn and Hahn 2001 multituberculate
MPZ 96/160–164
 Multituberculata - Paulchoffatiidae
Paulchoffatiinae indet.6 Hahn 1971 multituberculate
MPZ 96/174, 96/176
 Theriamorpha -
? Theria indet.6 Parker and Haswell 1897 mammal
MPZ 96/177–179
 Eutriconodonta - Gobiconodontidae
Gobiconodon sp.9 Trofimov 1978 mammal
Osteichthyes
 Cynodontia -
Docodonta indet.5 Kretzoi 1946 dodocont
Reptilia
 Testudines -
Cryptodira indet.9 Cope 1868 turtle
 Testudines - Dortokidae
 Squamata - Paramacellodidae
Paramacellodidae indet.9 Estes 1983 squamates
 Loricata -
Bernissartia sp.9 Dollo 1883 crocodilian
Crocodylia indet. crocodilian
 Loricata - Goniopholididae
Goniopholis sp.9 Owen 1842 crocodilian
 Loricata - Atoposauridae
aff. Theriosuchus sp.13 Owen 1878 crocodilian
 Dinosauria -
 Ornithischia -
? Stegosauria indet.9 ornithischian
Ornithopoda indet.9 ornithopod
 Ornithischia - Hypsilophodontidae
Hypsilophodontidae indet. hypsilophodontid
MPZ 96/57
cf. Hypsilophodon sp.9 Huxley 1870 hypsilophodontid
 Ornithischia -
Iguanodontoidea indet.7 Hay 1902 ornithopod
MPZ 96/50–52
Iguanodontoidea indet.7 Hay 1902 ornithopod
 Theropoda -
Coelurosauria indet.9 coelurosaur
 Theropoda - Dromaeosauridae
Dromaeosauridae indet.4 Colbert and Russell 1969 maniraptoran
MPZ 96/100
 Theropoda - Spinosauridae
"Baryonychidae indet." = Baryonychinae9
"Baryonychidae indet." = Baryonychinae9 Charig and Milner 1986 tetanuran theropod
 Saurischia -
Pleurocoelus valdensis10 Lydekker 1889 sauropod
Titanosauriformes indet.2 Salgado et al. 1997 sauropod
MPZ 99/672
 Pterosauria -
Pterodactyloidea "indet. A"11 Plieninger 1901 pterosaur
MPZ 96/59, 98/39 to 98/48; 21 teeth
Pterodactyloidea "indet. B"11 Plieninger 1901 pterosaur
MPZ 96/58
 Plesiosauria -
Plesiosauria indet.9 de Blainville 1835 plesiosaur
Actinopteri
 Pycnodontiformes - Pycnodontidae
Pycnodontidae indet.9 Agassiz 1833
 Holostei -
Holostei indet.9 Müller 1846
 Lepisosteiformes -
Lepidotes sp.9 Agassiz 1832 gar
 Amiiformes - Amiidae
Amiidae indet.9 Huxley 1861 bowfin