PBDB Collection

Paali Nala level C2 (DBC2): Early/Lower Oligocene, Pakistan
collected 2000 - 2004

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

Mammalia - Suidae

	Hyotherium sp. von Meyer 1834	Orliac et al. 2010
	UMC-DBC 2242 and 2244
Mammalia - Tayassuidae

	Pecarichoerus sp. Colbert 1933	Orliac et al. 2010
	UMC-DBC 2240 and 2241
Mammalia

	Ruminantia indet. Scopoli 1777	Métais et al. 2017
	DBC2 5037

	Sanitherium sp. von Meyer 1866	Orliac et al. 2010
	UMC-DBC 2239
Mammalia - Lophiomerycidae

	Lophiomeryx kargilensis Nanda and Sahni 1990	Métais et al. 2009
	DBC2 5009-5011, 5028, 5029, 5034, 5035, 5037, 5193-5200

	Nalameryx savagei (Nanda and Sahni 1990)	Métais et al. 2009
	DBC2 specimens

	Nalameryx sulaimani n. sp. Métais et al. 2009	Métais et al. 2009
	DBC2 5108 - type
Mammalia

	Paalitherium gurki n. gen., n. sp. Métais et al. 2017	Métais et al. 2017
	DBC2 5186 - holotype; referred material: DBC2 5036, 5042 and 5181

	Mosaicomeryx aff. quercyi (Jehenne 1987)	Métais et al. 2017
	DBC2 5040, 5043, 5158, 5160, 5172, 5173 and 5182
Mammalia - Palaeomerycidae

	aff. Amphitragulus sp. Pomel 1847	Métais et al. 2017
	DBC2 5038, 5041, 5157, 5161-5166, 5168 and 5185
Mammalia - Perissodactyla - Rhinocerotidae

	Epiaceratherium cf. magnum Uhlig 1999	Antoine et al. 2003
	DBC2-RHI-1-3
Mammalia - Rodentia

	Downsimys margolisi	Marivaux and Welcomme 2003
Mammalia - Rodentia - Diatomyidae

	cf. Fallomus sp. Flynn et al. 1986	Marivaux and Welcomme 2003
	DBC 295, 296, 476-494

	Fallomus quraishyi n. sp. Marivaux and Welcomme 2003	Marivaux and Welcomme 2003
	DBC 438 - type

	Fallomus razae Flynn et al. 1986	Marivaux and Welcomme 2003
	DBC 474, 475, 1001-1259, 1277

	Fallomus ginsburgi n. sp. Marivaux and Welcomme 2003	Marivaux and Welcomme 2003
	DBC 683 - type
Mammalia - Rodentia

	Bugtimys zafarullahi n. gen., n. sp. Marivaux et al. 2002	Marivaux et al. 2002
	DBC 1261 - holotype

	Hodsahibia gracilis n. sp. Marivaux and Welcomme 2003	Marivaux and Welcomme 2003
	DBC 302 - type

	Hodsahibia beamshaiensis n. sp. Marivaux and Welcomme 2003	Marivaux and Welcomme 2003
	DBC 1438 - type

	Hodsahibia azrae Flynn et al. 1986	Marivaux and Welcomme 2003
	DBC 308-373
Mammalia - Rodentia - Chapattimyidae

	Baluchimys chaudryi Flynn and Cheema 1994	Marivaux and Welcomme 2003

	Zindapiria quadricollis Flynn and Cheema 1994	Marivaux and Welcomme 2003
Mammalia - Rodentia - Cricetidae

	Atavocricetodon paaliense n. sp. Marivaux et al. 1999	Marivaux et al. 1999

	Pseudocricetodon nawabi n. sp. Marivaux et al. 1999	Marivaux et al. 1999
Mammalia - Rodentia - Dipodidae

	Allosminthus sp. Wang 1985	Marivaux and Welcomme 2003
Mammalia - Rodentia - Aplodontidae

	Oligopetes sp. Heissig 1979	Marivaux and Welcomme 2003
Mammalia - Primates - Ekgmowechashalidae

	Bugtilemur mathesoni n. gen., n. sp. Marivaux et al. 2001	Marivaux et al. 2001
Mammalia - Primates - Sivaladapidae

	Guangxilemur singsilai n. sp. Marivaux et al. 2002	Marivaux et al. 2002
Mammalia - Primates - Amphipithecidae

	Bugtipithecus inexpectans n. gen., n. sp. Marivaux et al. 2005
	UMC-DBC 2174 - holotype; referred material: UMC-DBC 2172, 2173, 2175-2178 and 2191
Mammalia - Primates - Eosimiidae

	Phileosimias brahuiorum n. sp. Marivaux et al. 2005
	UMC-DBC 2221 - type; referred material: UMC-DBC 2220 and 2222

	Phileosimias kamali n. gen., n. sp. Marivaux et al. 2005
	UMC-DBC 2199 - holotype; referred material: UMC-DBC 2192-2219
Mammalia

	Asiadidelphis akbarbugtii n. sp. Crochet et al. 2007	Crochet et al. 2007
	PMNH−DBC2−2180 - type; referred material: PMNH−DBC2−2179 and 2223
Chondrichthyes - Myliobatiformes - Gymnuridae

	Himantura sp. Müller and Henle 1837	Adnet et al. 2007
Chondrichthyes - Myliobatiformes - Aetobatidae

	Aetobatus sp. Blainville 1816	Adnet et al. 2007
Chondrichthyes - Myliobatiformes - Myliobatidae

	Myliobatis sp. Cuvier 1816	Adnet et al. 2007
Chondrichthyes - Myliobatiformes - Dasyatidae

	Dasyatis sp. Rafinesque 1810	Adnet et al. 2007
Chondrichthyes - Rhinopristiformes - Rhynchobatidae

	Rhynchobatus cf. pristinus Probst 1877	Adnet et al. 2007
Chondrichthyes - Rhinopristiformes - Pristidae

	? Pristis cf. lathami Galeotti 1837	Adnet et al. 2007
Chondrichthyes - Rajiformes - Rhinobatidae

	Rhinobatos sp. Link 1790	Adnet et al. 2007
Chondrichthyes - Carcharhiniformes - Sphyrnidae

	Sphyrna sp. Rafinesque 1810	Adnet et al. 2007
Chondrichthyes - Carcharhiniformes - Carcharhinidae

	Carcharhinus sp. de Blainville 1816	Adnet et al. 2007
	species 2

	Carcharhinus perseus n. sp. Adnet et al. 2007	Adnet et al. 2007

	Rhizoprionodon sp. Whitley 1929	Adnet et al. 2007
see common names

Geography

Country:	Pakistan	State/province:	Balochistan
Coordinates:	28.9° North, 69.1° East (view map)
Paleocoordinates:	19.0° North, 67.8° East
Basis of coordinate:	stated in text
Geographic resolution:	small collection

Time

Period:	Paleogene	Epoch:	Oligocene
Stage:	Rupelian	10 m.y. bin:	Cenozoic 4
Key time interval:	Early/Lower Oligocene
Age range of interval:	33.90000 - 27.82000 m.y. ago

Stratigraphy

Formation:	Chitarwata	Member:	Bugti
Stratigraphic resolution:	bed
Stratigraphy comments: "in the lowermost levels of the Lower Chitarwata Formation, which is early Oligocene in age"

Lithology and environment

Primary lithology:	sandstone
Includes fossils?	Y
Environment:	fluvial-deltaic indet.

Taphonomy

Modes of preservation:	body
Size of fossils:	macrofossils,mesofossils
Disassociated minor elements:	many

Collection methods and comments

Collection excludes:	some genera
Collection methods:	bulk,sieve,field collection
Reason for describing collection:	taxonomic analysis
Collection dates:	2000 - 2004
Collection method comments: material is in the collection of the Institut des Sciences d l'Evolution, Université de Montpellier II; collections were made by "intensive screen-washing"

Metadata

Also known as:	Bugti Hills; Gandoï syncline; DB-C2; Dera Bugti C2
Database number:	78520
Authorizer:	J. Alroy, M. Uhen, P. Coster, K. Beard, P. Mannion	Enterer:	J. Alroy, K. Beard, P. Mannion, M. Uhen, P. Coster
Modifier:	P. Mannion	Research group:	vertebrate
Created:	2008-02-07 14:26:51	Last modified:	2016-07-22 04:52:58
Access level:	the public	Released:	2008-02-07 14:26:51
Creative Commons license:	CC BY

Reference information

Primary reference:

26396.

L. Marivaux, P.-O. Antoine, S. R. H. Baqri, M. Benammi, and Y. Chaimanee. 2005. Anthropoid primates from the Oligocene of Pakistan (Bugti Hills): Data on early anthropoid evolution and biogeography. Proceedings of the National Academy of Sciences 102(24):8436-8441 [J. Alroy/J. Alroy]

Secondary references:

48852		S. Adnet, P. -O. Antoine, S. R. Hassan Baqri, J. Crochet, L. Marivaux, J. Welcomme, and G. Métais. 2007. New tropical carcharhinids (chondrichthyes, Carcharhiniformes) from the late Eocene–early Oligocene of Balochistan, Pakistan: Paleoenvironmental and paleogeographic implications. Journal of Asian Earth Sciences 30(2):303-323 [M. Uhen/S. Alam/M. Uhen]
59765		P.-O. Antoine, S. Ducrocq, L. Marivaux, Y. Chaimanee, J.-Y. Crochet, J.-J. Jaeger, and J.-L. Welcomme. 2003. Early rhinocerotids (Mammalia: Perissodactyla) from South Asia and a review of the Holarctic Paleogene rhinocerotid record. Canadian Journal of Earth Sciences 40:365-374 [P. Mannion/P. Mannion]
57997		J. Y. Crochet, P. O. Antoine, M. Benammi, N. Iqbal, L. Marivaux, G. Metais, and J. L. Welcomme. 2007. A herpetotheriid marsupial from the Oligocene of Bugti Hills, Balochistan, Pakistan. Acta Palaeontologica Polonica. Acta Palaeontologica Polonica 52(3):633-637 [P. Coster/P. Coster/P. Mannion]
59639		L. Marivaux. 2006. The eosimiid and amphipithecid primates (Anthropoidea) from the Oligocene of the Bugti Hills (Balochistan, Pakistan): New insight into early higher primate evolution in south Asia. Palaeovertebrata 34(1-2):29-109 [P. Mannion/P. Mannion]
58070		L. Marivaux, M. Vianey-Liaud, and J.-L. Welcomme. 1999. Première découverte de Cricetidae (Rodentia, Mammalia) oligocènes dans le synclinal sud de Gandoï (Bugti Hills, Balouchistan, Pakistan). Comptes Rendus de l'Académie des Sciences-Series IIA-Earth and Planetary Science 329(11):839-844 [P. Coster/P. Coster/P. Mannion]
59704		L. Marivaux, M. Vianey-Liaud, J.-L. Welcomme and J.-J. Jaegar. 2002. The role of Asia in the origin and diversification of hystricognathous rodents. Zoologica Scripta 31(3):225-239 [P. Mannion/P. Mannion]
28025		L. Marivaux, J.-L. Welcomme, P.-O. Antoine, G. Metais, I. M. Baloch, M. Benammi, Y. Chaimanee, S. Ducrocq, and J.-J. Jaeger. 2001. A fossil lemur from the Oligocene of Pakistan. Science 294:587-591 [K. Beard/K. Beard]
28023		L. Marivaux, J.-L. Welcomme, S. Ducrocq and J.-J. Jaeger. 2002. Oligocene sivaladapid primate from the Bugti Hills (Balochistan, Pakistan) bridges the gap between Eocene and Miocene adapiform communities in southern Asia. Journal of Human Evolution 42:379-388 [K. Beard/K. Beard]
59702		L. Marivaux and J.-L. Welcomme. 2003. New diatomyid and baluchimyine rodents from the Oligocene of Pakistan (Bugti Hills, Balochistan): systematic and paleobiogeographic implications. Journal of Vertebrate Paleontology 23(2):420-434 [P. Mannion/P. Mannion]
61475		G. Métais, B. Mennecart, and G. Roohi. 2017. A new assemblage of stem pecoran ruminants from the Oligocene Chitarwata Formation, Bugti Hills, Baluchistan, Pakistan: Paleoenvironmental and paleobiogeographic implications. Journal of Asian Earth Sciences 136:40-49 [P. Mannion/P. Mannion]
59705		G. Métais, J.-L. Welcomme, and S. Ducrocq. 2009. New lophiomerycid ruminants from the Oligocene of the Bugti Hills (Balochistan, Pakistan). Journal of Vertebrate Paleontology 29(1):231-241 [P. Mannion/P. Mannion]
59923		M. J. Orliac, P.-O. Antoine, G. Roohi and J.-L. Welcomme. 2010. Suoidea (Mammalia, Cetartiodactyla) from the early Oligocene of the Bugti Hills, Balochistan, Pakistan. Journal of Vertebrate Paleontology 30(4):1300-1305 [P. Mannion/P. Mannion]