Pasalar: MN 5, Turkey

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Sciuropteridae indet.
Mammalia - Talpidae
Talpidae indet. (Fischer de Waldheim 1817)
"Talpidae sp. 1"
    = Desmanodon cf. minor Engesser 1980
Engesser 1980
Mammalia - Soricidae
Soricidae indet. Fischer 1814
Soricidae sp. (small form)
Mammalia - Erinaceidae
Galerix cf. moedlingensis Rabeder 1973
    = Schizogalerix pasalarensis Engesser 1980
Engesser 1980
Erinaceinae indet. Fischer de Waldheim 1817
Mammalia - Carnivora - Felidae
Pseudaelurus larteti
"Pseudailurus (Schizailurus) larteti"
Pseudaelurus cf. quadridentatus (de Blainville 1843)
"Pseudailurus cr. quadridentatus"
Mammalia - Carnivora - Hyaenidae
Protictitherium intermedium Schmidt-Kittler 1976
recombined as Protoviverrops intermedium
Protictitherium sp. Kretzoi 1938
"Protictitherium sp. I"
Protictitherium aff. gaillardi Forsyth Major 1903
Mammalia - Carnivora - Ursidae
Ursavus cf. primaevus Gaillard 1899
Ursavus aff. intermedius von Koenigswald 1925
Hemicyon sansaniensis Lartet 1851
Plithocyon sp. Ginsburg 1955
Mammalia - Carnivora
Proputorius sp. Filhol 1890
Mammalia - Carnivora - Mustelidae
Plesiogulo sp. Zdansky 1934
said to be new
Trochictis sp. von Meyer 1842
Lutrinae indet. Baird 1857
"Lutrinae sp. I"
Mammalia - Carnivora - Amphicyonidae
Amphicyon cf. major De Blainville 1841
Mammalia - Perissodactyla - Equidae
Anchitherium sp. von Meyer 1844
Mammalia - Perissodactyla - Tapiridae
Tapirus sp. Brisson 1762
Mammalia - Perissodactyla - Rhinocerotidae
Beliajevina tekkayai Heissig 1974
recombined as Procoelodonta (Pasalarhinus) tekkayai
Mammalia - Suidae
Conohyus sp. Pilgrim 1926
Listriodon splendens
Mammalia
Schizochoerus sp. Crusafont-Pairo and Lavocat 1954
Mammalia - Bovidae
Bovidae indet. Gray 1821
Prostrepsiceros cf. rotundicornis
Protoryx cf. carolinae Major 1891
Mammalia - Cervidae
Cervidae indet. Gray 1821
Mammalia - Primates - Hominidae
Sivapithecus darwini
Ramapithecus wickeri
synonym of Sivapithecus
Mammalia - Lagomorpha - Ochotonidae
Alloptox sp. Dawson 1961
"Alloptox n.sp. 1"
Prolagus cf. oeningensis (König 1825)
Mammalia - Rodentia - Ctenodactylidae
Ctenodactylidae indet. Gervais 1853
Mammalia - Rodentia - Gliridae
Peridyromys lavocati n. sp. Pelaez-Campomanes and Daams 2002
Pelaez-Campomanes and Daams 2002
Pseudodryomys sp. de Bruijn 1966
Mammalia - Rodentia - Sciuridae
cf. Spermophilinus bredai (von Meyer 1848)
Mammalia - Rodentia - Spalacidae
cf. Rhizomys sp. Gray 1831
Mammalia - Rodentia - Cricetidae
Democricetodon sp. Fahlbusch 1964
Megacricetodon sp. (Fahlbusch 1964)
    = Megacricetodon andrewsi n. sp. Pelaez-Campomanes and Daams 2002
Pelaez-Campomanes and Daams 2002
Cricetodon sp. Lartet 1851
"Cricedoton ('Palaeocricetus') sp."
Mammalia - Rodentia - Dipodidae
Dipodidae indet. Waterhouse 1842
Mammalia - Tubulidentata - Orycteropodidae
Orycteropus sp. Geoffroy 1791
Mammalia - Proboscidea - Gomphotheriidae
Gomphotherium angustidens Burmeister 1837
Mammalia - Proboscidea - Deinotheriidae
Deinotherium giganteum Kaup 1829
Mammalia - Chiroptera
Chiroptera indet. Blumenbach 1779
Reptilia - Chamaeleonidae
Chamaeleonidae informal indet. 3 Gray 1825
Georgalis et al. 2022
A fragment of a left dentary (BP 2019-181)
Reptilia - Testudines - Geoemydidae
Mauremys cf. caspica (Gmelin 1774)
Staesche et al. 2007
see common names

Geography
Country:Turkey State/province:Bursa
Coordinates: 40.0° North, 28.3° East (view map)
Paleocoordinates:40.1° North, 28.7° East
Basis of coordinate:based on nearby landmark
Geographic resolution:outcrop
Time
Period:Neogene Epoch:Miocene
Stage:Langhian 10 m.y. bin:Cenozoic 5
Key time interval:MN 5
Age range of interval:15.98000 - 13.82000 m.y. ago
Stratigraphy
Stratigraphic resolution:bed
Lithology and environment
Primary lithology:concretionary,gray,green calcareous siltstone
Secondary lithology:fine,pebbly,gray,white sandy sandstone
Includes fossils?Y
Lithology description: "The fossiliferous sediments consist of 2.2 m of silts and sandstones, often quite coarse"; there are two fossiliferous levels, the upper a "0.8 m grey-green silt, unbedded, with many small calcareous concretions" and the lower a "1.0 m whitish to light grey sand, fine grained, unbedded, with frequent lenses of coarser sands and gravels... badly sorted, non calcified"; the list pertains only to the lower bed
Environment:fluvial indet.
Geology comments: "fluviatile, with rapid sedimentation in high energy conditions in the main fossiliferous bed followed by quieter conditions, but with occasional high energy episodes, later on" that probably represents "initial molasse-like fill of a tectonic basin near the lower slopes of a mountain range"
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils,mesofossils
Disassociated minor elements:many
Fragmentation:frequent
Collection methods and comments
Reason for describing collection:general faunal/floral analysis
Collection method comments: "discovered in 1969 during a lignite exploration... Some of these localities [including Pasalar] were subsequently investigated by a German research team"
Metadata
Also known as:Paşalar
Database number:68547
Authorizer:L. van den Hoek Ostende, J. Alroy, E. Vlachos, J. Head Enterer:L. van den Hoek Ostende, J. Alroy, G. Varnham, E. Vlachos
Modifier:G. Varnham Research group:vertebrate
Created:2007-01-25 16:28:07 Last modified:2023-06-29 04:33:38
Access level:the public Released:2007-01-25 16:28:07
Creative Commons license:CC BY
Reference information

Primary reference:

24292. P. Andrews and H. Tobien. 1977. New Miocene locality in Turkey with evidence of the origin of Ramapithecus and Sivapithecus. Nature 268:699-701 [J. Alroy/J. Alroy]

Secondary references:

19450 B. Engesser. 1980. Insectivora und Chiroptera (Mammalia) aus dem Neogen der Türkei. Schweizerischen Paläontologischen Abhandlungen 102:46-149 [L. van den Hoek Ostende/L. van den Hoek Ostende/L. van den Hoek Ostende]
85133 G. L. Georgalis, A. Čerňanský, F. Göktaş, B. Alpagut, A. and S. Mayda. 2022. The antiquity of asian chameleons - first potential Chameleonidae and associated squamate fauna from the lower and middle Miocene of Anatolia. Journal of Vertebrate Paleontology 42(2):e2160644 [J. Head/G. Varnham/G. Varnham]
26760 P. Pelaez-Campomanes and R. Daams. 2002. Middle Miocene rodents from Paşalar, Anatolia, Turkey. Acta Palaeontologica Polonica 47(1):125-132 [L. van den Hoek Ostende/L. van den Hoek Ostende]
61765 K. Staesche, H.V. Karl, and U. Staesche. 2007. Fossile Schildkroten aus der Turkei. In U. Staesche (ed.), Fossile Schildkroten aus drei Kontinenten 98:91-149 [E. Vlachos/E. Vlachos]