Toogreeg Shire [PMPE]: Late/Upper Campanian, Mongolia
collected by PMPE 1968, 1971

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Reptilia - Dinosauria
Dinosauria indet. Owen 1842
Kielan-Jaworowska and Barsbold 1972
eggs
Reptilia - Theropoda
Theropoda indet. Marsh 1881
Kielan-Jaworowska and Barsbold 1972
"small theropod dinosaurs"
Reptilia - Elongatoolithidae
Elongatoolithidae indet. Zhao 1975
Sabath 1991 2 specimens
ZPAL MgOv-II/14a-b
Reptilia - Dromaeosauridae
Velociraptor mongoliensis Osborn 1924
Barsbold 1974 1 individual
GI SPS 100/25
Reptilia - Protoceratopsidae
Protoceratops andrewsi Granger and Gregory 1923
Barsbold 1974 3 individuals
GI SPS 100/512
Mammalia - Multituberculata - Djadochtatheriidae
Kryptobaatar dashzevegi Kielan-Jaworowska 1969
Kielan-Jaworowska and Hurum 1997 2 specimens
including GI SPS 8-2PST; PSS-MAE 113
Tugrigbaatar saichanensis n. gen., n. sp. Kielan-Jaworowska and Dashzeveg 1978
Kielan-Jaworowska and Dashzeveg 1978 1 individual
synonym of Kryptobaatar dashzevegi
Mammalia - Zalambdalestidae
Zalambdalestes lechei Gregory and Simpson 1926
Kielan-Jaworowska 1984 1 individual
see common names

Geography
Country:Mongolia State/province:Omnogov
Coordinates: 44.3° North, 103.3° East (view map)
Paleocoordinates:41.9° North, 92.3° East
Basis of coordinate:stated in text
Altitude:1150 meters
Geographic resolution:outcrop
Time
Period:Cretaceous Epoch:Late/Upper Cretaceous
Stage:Campanian 10 m.y. bin:Cretaceous 7
*Epoch:Senonian
*International age/stage:Campanian - Campanian
Key time interval:Late/Upper Campanian
Age range of interval:83.60000 - 72.10000 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Formation:Djadokhta Member:Tugrugyin
Stratigraphic resolution:group of beds
Stratigraphy comments: ETE dating method: time_unit, ETE age comment: . radiometric intage_min; several localities in the outcrop area with no specific stratigraphic information given
Lithology and environment
Primary lithology:tabular,"cross stratification",fine,ferruginous,white,yellow poorly lithified sandstone
Includes fossils?Y
Lithology description: ETE rock type adj: 0, ETE lithology comments: . "White-yellowish sand and sandstone"; "poorly cemented white, fine-grained sandstones. Megascopic sedimentary structures are usually absent, and only rarely medium- or large-scale cross-bedded units of tabular type can be seen. The sandstones contain very numerous oval or flat bodies of tough sandstone covered by ferruginous desert varnish and regular horizons of ferruginous cementation developed within sandstone beds. (10453)
Environment:dune
Geology comments: ETE sed env 1: eolian, ETE sed env 2: dune ETE event: storm, ETE env comment: "sndstrm,debris flow"
Taphonomy
Modes of preservation:body
Degree of concentration:dispersed
Size of fossils:macrofossils
Spatial orientation:life position
Preservation of anatomical detail:good
Articulated whole bodies:many
Associated major elements:some
Disassociated major elements:some
Disassociated minor elements:some
Fragmentation:occasional
Temporal resolution:snapshot
Spatial resolution:autochthonous
Collection methods and comments
Collection methods:selective quarrying,surface (float),surface (in situ),mechanical,field collection
Reason for describing collection:general faunal/floral analysis
Collectors:PMPE Collection dates:1968, 1971
Collection method comments: ETE reference list: 478
Metadata
Also known as:ETE Locality 1560, 1561, Fighting dinosaurs, Tugrikin Shire
Database number:22685
Authorizer:A. Behrensmeyer, R. Benson, M. Carrano Enterer:A. Behrensmeyer, M. Carrano, R. Benson
Modifier:M. Carrano Research group:taphonomy,vertebrate
Created:2001-04-23 01:00:00 Last modified:2023-11-21 12:09:54
Access level:the public Released:2001-04-23 01:00:00
Creative Commons license:CC BY
Reference information

Primary reference:
10453. R. Gradzinski and T. Jerzykiewicz. 1972. Additional geographical and geological data from the Polish-Mongolian Palaeontological Expeditions. Palaeontologia Polonica 27:17-306 [M. Carrano/M. Carrano/M. Carrano]

Secondary references:

63431 R. Barsbold. 1974. Poyedinok dinozavrov [Dueling dinosaurs]. Prioda 1974(2):81-83 [M. Carrano/M. Carrano]
36735ETE R. Barsbold. 1998. Fighting dinosaurs—they really fought. In A. M. G. de Carvalho, A. M. Andrade, V. F. dos Santos, J. Cascalho, R. Taborda, (eds.), I Encontro Internacional sobre Paleobiologia dos Dinossáurios. Museu Nacional de História Natural, Lisboa 74-78 [M. Carrano/M. Carrano]
86542 R. Barsbold. 2016. “The Fighting Dinosaurs”: the position of their bodies before and after death. Paleontological Journal 50(12):1412-1417 [M. Carrano/M. Carrano]
81410 K. Carpenter. 2000. Evidence of predatory behavior by carnivorous dinosaurs. Gaia 15:135-144 [M. Carrano/M. Carrano]
18347ETE P. J. Currie. 2000. Theropods from the Cretaceous of Mongolia. In M. J. Benton, M. A. Shishkin, D. M. Unwin, & E N. Kurichkin (eds.), The Age of Dinosaurs in Russia and Mongolia 434-455 [M. Carrano/M. Carrano/M. Carrano]
15459ETE D. Dashzeveg, L. Dingus, D.B. Loope, C.C. Swisher, III, T. Dulam and M.R. Sweeney. 2005. New stratigraphic subdivision, depositional environment, and age estimate for the Upper Cretaceous Djadokhta Formation, southern Ulan Nur Basin, Mongolia. American Museum Novitates (3498)1-31 [M. Carrano/K. Maguire/G. Varnham]
59136 R. Gradzinski, Z. Kielan-Jaworowska, and T. Maryanska. 1977. Upper Cretaceous Djadokhta, Barun Goyot and Nemegt formations of Mongolia, including remarks on previous subdivisions. Acta Geologica Polonica 27(3):281-318 [M. Carrano/M. Carrano]
58719 T. Jerzykiewicz. 1997. Djadokhta Formation. In P. J. Currie & K. Padian (ed.), Encyclopedia of Dinosaurs 188-191 [M. Carrano/M. Carrano]
51934 T. Jerzykiewicz. 1998. Okavango Oasis, Kalahari Desert: a contemporary analogue for the Late Cretaceous vertebrate habitat of the Gobi Basin, Mongolia. Geoscience Canada 25(1):15-26 [M. Carrano/A. Murch]
5680ETE,5% 16440T. Jerzykiewicz, P. J. Currie, D. A. Eberth, P. A. Johnston, E. H. Koster and J. J. Zheng. 1994. Djadokha Formation correlative strata in Chinese Inner Mongolia: an overview of the stratigraphy, sedimentary geology, and paleontology and comparisons with the type locality in the pre-Altai Gobi. Canadian Journal of Earth Sciences 39(10-11):2180-2195 [A. Behrensmeyer/A. Behrensmeyer/M. Carrano]
63572 Z. Kielan-Jaworowska. 1984. A caccia di dinosauri nel deserto di Gobi [Hunting dinosaurs in the Gobi Desert]. In G. M. Ronzoni, M. L. Greggio, & T. B. Guarinoni (eds.), Sulle Orme dei Dinosauri 87-109 [M. Carrano/M. Carrano]
10463 Z. Kielan-Jaworowska. 1984. Evolution of the therian mammals in the Late Cretaceous of Asia. Part V. Skull structure in Zalambdalestidae. Palaeontologia Polonica 46:107-117 [M. Carrano/M. Carrano/M. Carrano]
9899 Z. Kielan-Jaworowska and R. Barsbold. 1972. Narrative of the Polish-Mongolian Palaeontological Expeditions 1967-1971. Palaeontologia Polonica 27:5-136 [M. Carrano/M. Carrano/M. Carrano]
10465 Z. Kielan-Jaworowska and D. Dashzeveg. 1978. New Late Cretaceous mammal locality in Mongolia and a description of a new multituberculate. Acta Palaeontologica Polonica 23(2):115-128 [M. Carrano/M. Carrano/M. Carrano]
38300 Z. Kielan-Jaworowska and J. H. Hurum. 1997. Djadochtatheria - a new suborder of multituberculate mammals. Acta Palaeontologica Polonica 42(2):201-242 [R. Benson/R. Benson]
38909 Z. Kielan-Jaworowska, J. H. Hurum, and D. Badamgarav. 2003. An extended range of the multituberculate Kryptobaatar and the distribution of mammals in the Upper Cretaceous of the Gobi Desert. Acta Palaeontologica Polonica 48(2):273-278 [R. Benson/R. Benson]
10057 O. Lambert, P. Godefroit, H. Li, C.-Y. Shang, and Z.-M. Dong. 2001. A new species of Protoceratops (Dinosauria, Neoceratopsia) from the Late Cretaceous of Inner Mongolia (P. R. China). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Science de la Terre 71(supplement):5-28 [M. Carrano/M. Carrano/M. Carrano]
14217ETE K. Mikhailov, K. Sabath, and S. Kurzanov. 1994. Eggs and nests from the Cretaceous of Mongolia. In K. Carpenter, K. F. Hirsch, and J. R. Horner (eds.), Dinosaur Eggs and Babies, Cambridge University Press, Cambridge 88-115 [M. Carrano/M. Carrano/M. Carrano]
62057 H. Osmólska. 1997. Polish-Mongolian Paleontological Expeditions. In P. J. Currie & K. Padian (ed.), Encyclopedia of Dinosaurs 573-576 [M. Carrano/M. Carrano]
41004 K. Sabath. 1991. Upper Cretaceous amniotic eggs from the Gobi Desert. Acta Palaeontologica Polonica 36(2):151-192 [M. Carrano/M. Carrano]
17038ETE D. M. Unwin, A. Perle, and C. Trueman. 1995. Protoceratops and Velociraptor preserved in association: evidence for predatory behaviour in dromaeosaurid dinosaurs?. Journal of Vertebrate Paleontology 15(3, suppl.):57A-58A [M. Carrano/M. Carrano]