Level 2, Benguérir: Late/Upper Maastrichtian, Morocco

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Reptilia - Dyrosauridae
Dyrosauridae indet. Stefano 1903
Reptilia - Plesiosauria - Elasmosauridae
Elasmosauridae indet. Cope 1869
Reptilia - Mosasauridae
Eremiasaurus heterodontus LeBlanc et al. 2012
Mosasaurus beaugei Arambourg 1952
Prognathodon sp. Dollo 1889
Prognathodon currii Christiansen and Bonde 2002
Bardet et al. 2005 1 specimen
OCP.DEK/GE (one tooth)
Globidens phosphaticus Bardet et al. 2005
Bardet et al. 2005 2 specimens
OCP.DEK/GE 348 (posterior tooth); OCP.DEK/GE unnumbered (tooth)
Halisaurus arambourgi Bardet et al. 2005
"Platecarpus" ptychodon Arambourg 1952
nomen dubium belonging to Plioplatecarpinae
Reptilia - Pachyvaranidae
Pachyvaranus crassispondylus Arambourg 1952
Houssaye et al. 2011 1 specimen
OCP DEK/GE (vertebrae)
Reptilia - Testudines - Bothremydidae
Bothremydidae indet. Baur 1891
Reptilia - Testudines
Chelonioidea indet. (Baur 1893)
Actinopteri - Elopiformes - Phyllodontidae
Pseudoegertonia sp. Dartevelle and Casier 1949
Actinopteri - Aulopiformes - Dercetidae
Stratodus apicalis Cope 1872
Actinopteri - Aulopiformes - Enchodontidae
Enchodus libycus
Enchodus bursauxi
Actinopteri - Tetraodontiformes - Eotrigonodontidae
Stephanodus libycus
Actinopteri - Pycnodontiformes - Pycnodontidae
Pycnodontidae indet. (Agassiz 1833)
Chondrichthyes - Myliobatiformes
Coupatezia fallax (Arambourg 1952)
original and current combination Raja fallax
Coupatezia elevata Noubhani and Cappetta 1997
Ixobatis mucronata (Arambourg 1952)
Chondrichthyes - Myliobatiformes - Rhombodontidae
Rhombodus microdus
Rhombodus binkhorsti Dames 1881
Chondrichthyes - Myliobatiformes - Dasyatidae
Dasyatis sp. Rafinesque 1810
Chondrichthyes - Rajiformes - Sclerorhynchidae
Ganopristis leptodon Arambourg 1935
Dalpiazia stromeri Checchia-Rispoli 1933
Ctenopristis nougareti Arambourg 1940
Schizorhiza stromeri Weiler 1930
Chondrichthyes - Rajiformes - Rhinobatidae
"Rhinobatos" informal sp. 1 Link 1790
"Rhinobatos" informal sp. 2 Link 1790
"Rhinobatos" informal sp. 3 Link 1790
"Rhinobatos" informal sp. 4 Link 1790
Chondrichthyes - Carcharhiniformes - Scyliorhinidae
Pteroscyllium sp. Cappetta 1980
Scyliorhinus aff. elongatus (Davis 1887)
Chondrichthyes - Orectolobiformes - Ginglymostomatidae
Ganntouria variabilis Noubhani and Cappetta 1997
Plicatoscyllium pectinatum
Plicatoscyllium youssoufiaense Noubhani and Cappetta 1997
Plicatoscyllium lehneri (Leriche 1938)
Chondrichthyes - Lamniformes - Pseudocoracidae
Pseudocorax affinis (Agassiz 1843)
Chondrichthyes - Lamniformes - Mitsukurinidae
Anomotodon plicatus Arambourg 1952
Chondrichthyes - Lamniformes - Otodontidae
Cretolamna maroccana
Cretolamna appendiculata (Agassiz 1835)
recombined as Cretalamna appendiculata
Chondrichthyes - Lamniformes - Serratolamnidae
Serratolamna africana
Serratolamna serrata (Agassiz 1838)
Chondrichthyes - Lamniformes - Anacoracidae
Squalicorax bassanii
Squalicorax pristodontus (Agassiz 1835)
Chondrichthyes - Lamniformes - Carchariidae
"Carcharias" heathi Case and Cappetta 1997
synonym of Carcharias samhammeri
Chondrichthyes - Squalomorphii - Hexanchidae
Hexanchus microdon (Agassiz 1835)
Chondrichthyes - Squalomorphii - Pristiophoridae
Pristiophorus sp. Bleeker 1859
Chondrichthyes
Youssoubatis aff. ganntourensis
see common names

Geography
Country:Morocco
Coordinates: 32.3° North, 7.8° West (view map)
Paleocoordinates:24.4° North, 8.4° West
Basis of coordinate:based on nearby landmark
Geographic resolution:outcrop
Time
Period:Cretaceous Epoch:Late/Upper Cretaceous
Stage:Maastrichtian 10 m.y. bin:Cretaceous 8
Key time interval:Late/Upper Maastrichtian
Age range of interval:72.10000 - 66.00000 m.y. ago
Stratigraphy
Stratigraphic resolution:group of beds
Stratigraphy comments: The use of dissimilarity indices and agglomerative method underscores two distinct associations: a lower one including the levels L6 and L5, and an upper one comprising the levels L4 to L2. These two associations allow to separate a lower and an upper Maastrichtian level. Moreover, the special morphology of the teeth of S. pristodontus occurring in L2, compared to what is observed in lower levels, permits us to distinguish a lower Upper Maastrichtian and an upper Upper Maastrichtian faunal assemblage. L2 includes a 1.3 m lower part and 1.4 m upper part.
Lithology and environment
Primary lithology: poorly lithified phosphorite
Environment:marine indet.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils,mesofossils
Collection methods and comments
Collection methods:surface (in situ),field collection
Reason for describing collection:general faunal/floral analysis
Museum repositories:MNHN
Collection method comments: Repository: University of Montpellier 2; Office Chérifien des Phosphates, Khouribga, Morocco (OCP)
Metadata
Database number:99627
Authorizer:J. Sessa Enterer:L. Eccles
Modifier:M. Clapham Research group:vertebrate
Created:2010-11-17 07:10:21 Last modified:2020-09-07 22:20:51
Access level:the public Released:2010-11-17 07:10:21
Creative Commons license:CC BY
Reference information

Primary reference:
73776. H. Cappetta, N. Bardet, X. Pereda Suberbiola, S. Adnet, D. Akkrim, M. Amalik, and A. Benabdallah. 2014. Marine vertebrate faunas from the Maastrichtian phosphates of Benguérir (Ganntour Basin, Morocco): Biostratigraphy, palaeobiogeography and palaeoecology. Palaeogeography, Palaeoclimatology, Palaeoecology 409:217-238 [M. Clapham/M. Clapham]

Secondary references:

37608 N. Bardet, X. Pereda Suberbiola, M. Iarochène, M. Amalik, and B. Bouya. 2005. Durophagous Mosasauridae (Squamata) from the Upper Cretaceous phosphates of Morocco, with description of a new species of Globidens. Netherlands Journal of Geosciences — Geologie en Mijnbouw 84(3):167-175 [P. Mannion/P. Mannion]
38134 A. Houssaye, N. Bardet, J.-C. Rage, X. Pereda-Suberbiola, B. Bouya, M. Amaghzaz, and M. Amalik. 2011. A review of Pachyvaranus crassispondylus Arambourg, 1952, a pachyostotic marine squamate from the latest Cretaceous phosphates of Morocco and Syria. Geological Magazine 148(2):237-249 [P. Mannion/P. Mannion]