Bhagothoro Hill: Rupelian, Pakistan
collected by Vredenburg

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Pleurotoma (Hemipleurotoma) bonneti Cossmann 1900
SUBSPECIES: Pleurotoma (Hemipleurotoma) bonneti bhagothorensis
    = Gemmula (Gemmula) bonneti Cossmann 1900
Harzhauser et al. 2009
Bivalvia - Arcida - Arcidae
Arca semitorta Lamarck 1819
Vredenburg 1928
recombined as Trisidos semitorta
Arca submultiformis Vredenburg 1928
Vredenburg 1928
Bivalvia - Ostreida - Ostreidae
Ostrea protoimbricata Vredenburg 1928
Vredenburg 1928
Ostrea gajensis Vredenburg 1928
Vredenburg 1928
Ostrea fraasi Mayer-Eymar 1888
Vredenburg 1928
Ostrea cubitus Deshayes 1832
Vredenburg 1928
Bivalvia - Pectinida - Pectinidae
Pecten (Amussiopecten) labadyei (d'Archaic and Haime 1853)
Vredenburg 1928
Bivalvia - Pectinida - Anomiidae
Placuna (Indoplacuna) sindiensis Vredenburg 1923
Vredenburg 1928
recombined as Indoplacuna sindiensis
Bivalvia - Carditida - Crassatellidae
Crassatella carcarensis Michelotti 1847
Vredenburg 1928
Crassatella sulcata (Solander 1866)
Vredenburg 1928
Bivalvia - Carditida - Carditidae
Cardita (Venericardia) laurae (Brongniart 1823)
Vredenburg 1928
Cardita arduini Brongniart 1823
Vredenburg 1928
Bivalvia - Pholadida - Corbulidae
Corbula carinata Dujardin 1837
Vredenburg 1928
Bivalvia - Cardiida - Cardiidae
Cardium (Nemocardium) bhagothorense Vredenburg 1928
Vredenburg 1928
Cardium (Discors) naricum Vredenburg 1928
Vredenburg 1928
recombined as Nemocardium (Discors) naricum
Cardium (Trachycardium) sindiense Vredenburg 1928
Vredenburg 1928
Bivalvia - Cardiida - Solecurtidae
Solenocurtus sindiensis Vredenburg 1928
Vredenburg 1928
Bivalvia - Cardiida - Ungulinidae
Diplodonta incerta d'Archiac 1850
Vredenburg 1928
SUBSPECIES: Diplodonta incerta narica
Bivalvia - Cardiida - Veneridae
Cytherea (Callista) spendida
Vredenburg 1928
original and current combination Callista
Venus (Ventricola) multilamella (Lamarck 1818)
Vredenburg 1928
Venus (Omphaloclathrum) puerpera (Linnaeus 1758)
Vredenburg 1928
recombined as Periglypta puerpera
SUBSPECIES: Venus (Omphaloclathrum) puerpera aglaurae
Cytherea (Pitar) porrecta (Koenen 1894)
Vredenburg 1928
recombined as Pitar porrecta
Bivalvia - Nuculida - Nuculidae
Nucula narica Vredenburg 1928
Vredenburg 1928
Cephalopoda - Nautilida - Aturiidae
Aturia narica Vredenburg 1925
Gastropoda - Turbinidae
Turbo (Olearia) protocepoides Vredenburg 1928
Vredenburg 1928
recombined as Turbo protocepoides
Turbo (Senectus) radiatus (Gmelin 1791)
Vredenburg 1928
recombined as Turbo (Marmarostoma) radiatus
SUBSPECIES: Turbo (Senectus) radiatus naricus
Gastropoda - Opisthobranchia - Cylichnidae
Scaphander oligoturritus Sacco 1897
Gastropoda - Opisthobranchia - Akeridae
Acera narica Vredenburg 1925
recombined as Akera narica
Gastropoda - Heterostropha - Architectonicidae
Solarium naricum Vredenburg 1928
Vredenburg 1928
recombined as Architectonica narica
Gastropoda - Cerithiidae
Tympanotomus laevis Vredenburg 1928
Vredenburg 1928
Cerithium (Chondrocerithium) bhagothorense Vredenburg 1928
Vredenburg 1928
recombined as Cerithium bhagothorense
Cerithium (Ptychocerithium) tricingularum
Vredenburg 1928
Cerithium (Ptychocerithium) ighinai (Michelotti 1861)
Vredenburg 1928
Cerithium (Ptychocerithium) haimei Vredenburg 1928
Vredenburg 1928
Cerithium (Ptychocerithium) perlamellosum Vredenburg 1928
Vredenburg 1928
recombined as Ptychocerithium perlamellosum
Cerithium (Ptychocerithium) sindiense Vredenburg 1928
Vredenburg 1928
recombined as Cerithium sindiense
Cerithium naricum Vredenburg 1925
recombined as Bellatara narica
Gastropoda - Turritellidae
Turritella conofasciata (Sacco 1895)
Vredenburg 1928
recombined as Turritella (Haustator) conofaciata
Turritella asperula Brogniart 1823
Vredenburg 1928
recombined as Turritella (Haustator) asperula
Protoma deshayesi (d'Archiac 1850)
Vredenburg 1928
Turritella narica Vredenburg 1928
Vredenburg 1928
SUBSPECIES: Turritella narica baluchistanensis
Turritella bhagothorensis Vredenburg 1928
Vredenburg 1928
Turritella desmarestina Basterot 1825
Vredenburg 1928
recombined as Turritella (Peyrotia) desmarestina
Turritella pseudotethis Vredenburg 1928
Vredenburg 1928
Gastropoda - Naticidae
Sigaretus aquensis Recluz 1843
Vredenburg 1928
recombined as Sinum aquensis
SUBSPECIES: Sigaretus aquensis praecedens
Gastropoda - Hipponicidae
Hipponyx cornucopae (Lamarck 1802)
Vredenburg 1928
Gastropoda - Ficidae
Pirula condita Brongniart 1823
recombined as Ficus condita
Gastropoda - Cypraeidae
Cypraea (Erosaria) sindiensis Vredenburg 1925
Gastropoda - Eocypraeidae
Cypraea (Bernayia) subexcisa (Braun 1840)
recombined as Apiocypraea subexcisa
Gastropoda - Neogastropoda - Buccinidae
Cominella annandalei Vredenburg 1925
Gastropoda - Neogastropoda - Nassariidae
Hindsia nivea (Gmelin 1790)
recombined as Nassaria nivea
SUBSPECIES: Hindsia nivea narica and Hindsia nivea bhagothorensis
Gastropoda - Neogastropoda - Fasciolariidae
Fusus (Aptyxis) reticulatus Vredenburg 1925
recombined as Aptyxis reticulatus
Streptochetus pseudowaelii Vredenburg 1925
Euthriofusus subregularis (d'Archiac and Haime 1854)
INCLUDES: Euthriofusus subregularis narica
Lathyrus sindiensis Vredenburg 1921
recombined as Latirus sindiensis
Gastropoda - Neogastropoda - Cancellariidae
Trigonostoma indicum Vredenburg 1925
Uxia narica Vredenburg 1925
Gastropoda - Neogastropoda - Olividae
Olivella elegantula Rovereto 1900
Ancilla (Sparella) indica Vredenburg 1925
recombined as Ancilla indica
Gastropoda - Neogastropoda - Turridae
Pleurotoma (Hemipleurotoma) yenanensis (Noetling 1895)
SUBSPECIES: Pleurotoma (Hemipleurotoma) yenanensis narica
    = Hemipleurotoma narica Vredenburg 1925
Harzhauser et al. 2009
Gastropoda - Neogastropoda - Conidae
Conus (Lithoconus) ineditus (Michelotti 1861)
Conorbis dormitor (Solander 1766)
SUBSPECIES: Conorbis dormitor sindiensis and Conorbis dormitor bhagothorensis
    = Conorbis sindiensis Vredenburg 1925
Harzhauser et al. 2009
Conorbis bhagothorensis Vredenburg 1925
Harzhauser et al. 2009
Gastropoda - Neogastropoda - Terebridae
Terebra subtessellata d'Orbigny 1852
SUBSPECIES: Terebra subtessellata oligocenica
    = Terebra perturrita Sacco 1891
Harzhauser 2007
Gastropoda - Neogastropoda - Volutidae
Athleta (Volutospina) sindiensis Vredenburg 1925
Lyria anceps (Michelotti 1861)
Gastropoda - Neogastropoda - Harpidae
Harpa (Eocithara) narica Vredenburg 1925
recombined as Eocithara narica
Gastropoda - Neogastropoda - Marginellidae
Marginella (Glabella) narica Vredenburg 1925
Gastropoda - Neogastropoda - Muricidae
Murex (Haustellum) naricus Vredenburg 1925
Murex (Muricantha) ighinae (Bellardi 1872)
recombined as Muricantha ighinae
Gastropoda - Rostellariidae
Rostellaria sindiensis Vredenburg 1925
recombined as Varicospira sindiensis
Gastropoda - Seraphsidae
Terebellum (Seraphs) naricum Vredenburg 1925
recombined as Seraphs naricus
Gastropoda - Personidae
Persona reticulata (Linnaeus 1767)
SUBSPECIES: Persona reticulata subclathrata
    = Distorsio decipiens Reeve 1844
Harzhauser et al. 2009
Gastropoda - Cassidae
Cassidea (Semicassis) oligocatlantica
original and current combination Semicassis
Cassidea mamillaris (Grateloup 1847)
original and current combination Cassis mamillaris
SUBSPECIES: Cassidea mamillaris nummultiphila
Gastropoda - Ranellidae
Tritonium (Sassia) indicum Vredenburg 1925
recombined as Sassia indica
Gastropoda - Cymatiidae
Ranella tubercularis (Risso 1826)
Gastropoda - Xenophoridae
Tugurium subextensum (d'Orbigny 1852)
Vredenburg 1928
Gastropoda - Ampullinidae
Ampullospira (Euspirocrommium) oweni (d'Archiac and Haime 1854)
Vredenburg 1928
recombined as Pachycrommium oweni
Ampullina (Globularia) gibberosa (Grateloup 1847)
Vredenburg 1928
recombined as Globularia gibberosa
see common names

Geography
Country:Pakistan
Coordinates: 26.4° North, 67.9° East (view map)
Paleocoordinates:16.9° North, 65.9° East
Basis of coordinate:stated in text
Geographic resolution:outcrop
Time
Period:Paleogene Epoch:Oligocene
Stage:Rupelian 10 m.y. bin:Cenozoic 4
Key time interval:Rupelian
Age range of interval:33.90000 - 27.82000 m.y. ago
Stratigraphy
Formation:Nari
Stratigraphic resolution:group of beds
Stratigraphy comments: STRATIGRAPHIC RELATIONS: Basal beds of regional succession, and is overlain by the Gaj beds (Early Miocene). From the Lower Nari. AGE: Early Oligocene, based on similarity with European faunas of Lattofian and Stampian age. STRATIGRAPHIC POSITION: Unknown position within unit.
Lithology and environment
Primary lithology: poorly lithified "limestone"
Includes fossils?Y
Lithology description: SPECIFIC LITHOLOGY: "Yellow and brown limestones" in Blanford (1879). Formation in the Kirthar Range to the north described as "cross-stratified, ripple-laminated, medium- to coarse-grained sandstone interbedded with minor amounts of shale. Limestone containing marine fossils and interbedded with shale and sandstone is commonly present in the lower part of the formation." LITHIFICATION: Poorly-lithified, based on figured specimens.
Environment:deep subtidal ramp
Geology comments: ENVIRONMENT: Fairly shallow, as larger foraminifera occur. In the Kirthar Range, the lithological description supports an deep subtidal environment, given the interbedding of cross-stratified or ripple-laminated sandstone with shale.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Collection methods and comments
Collection methods:surface (in situ),field collection
Reason for describing collection:taxonomic analysis
Collectors:Vredenburg
Collection method comments: COLLECTORS: Multiple collectors including W. Blanford, Wynne, Hira Lal, Ram Singh, and Noetling (staff of Geological Survey of India) during the early 1900's. REPOSITORY: Geological Survey of India, with voucher specimens possibly at the BMNH.
Taxonomic list comments:COVERAGE: Exhaustive for gastropoda, bivalvia, and scaphopoda. NOMENCLATURE: Antiquated, but from an authoritative publication, with use of species and subgenus resolution assignments. Nomenclature can be updated through reference to Harzhauser and Swarko publications.
Metadata
Database number:77159
Authorizer:A. Hendy Enterer:A. Hendy
Modifier:M. Clapham Research group:marine invertebrate
Created:2007-12-21 15:46:39 Last modified:2019-09-10 19:11:48
Access level:authorizer only Released:2008-12-21 15:46:39
Creative Commons license:CC BY
Reference information

Primary reference:
26063. 1925E. Vredenburg. 1925. Description of Mollusca from the post-Eocene Tertiary formation of north-western India: Cephalopoda, Opisthobranchiata, Siphonostomata. Memoirs of the Geological Survey of India 50(1):1-350 [A. Hendy/A. Hendy]

Secondary references:

25532 M. Harzhauser. 2007. Oligocene and Aquitanian gastropod faunas from the Sultanate of Oman and their biogeographic implications for the western Indo-Pacific. Palaeontographica Abteilung A 280:75-121 [A. Hendy/A. Hendy]
36478 M. Harzhauser, M. Euter, W. E. Piller, B. Berning, A. Kroh and O. Mandic. 2009. Oligocene and Early Miocene gastropods from Kutch (NW India) document an early biogeographic switch from Western Tethys to Indo-Pacific. Palaeontologische Zeitschrift 83:333-372 [A. Hendy/A. Hendy]
26112 E. Vredenburg. 1928. Description of Mollusca from the post-Eocene Tertiary formation of north-western India: Gastropoda (in part) and Lamellibranchiata. Memoirs of the Geological Survey of India 50(2):351-506 [A. Hendy/A. Hendy/A. Hendy]