Åsen: Early/Lower Campanian, Sweden
collected 1990, 2001, 2010

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Chondrichthyes - Hybodontiformes - Hybodontidae
Hybodus sp. Agassiz 1834
Sørensen et al. 2013
Chondrichthyes - Hybodontiformes - Polyacrodontidae
Polyacrodus siversoni
Sørensen et al. 2013
Polyacrodus sp. Jaekel 1889
Sørensen et al. 2013
Chondrichthyes
Rhinobatoidea informal sp. 2
Sørensen et al. 2013
Chondrichthyes - Rajiformes - Rhinobatidae
Rhinobatos informal sp. 1 Link 1790
Sørensen et al. 2013
Rhinobatos informal sp. 2 Link 1790
Sørensen et al. 2013
Rhinobatos informal sp. 3 Link 1790
Sørensen et al. 2013
Rhinobatos casieri Herman 1977
Sørensen et al. 2013
Chondrichthyes - Rajiformes
Squatirhina sp. Casier 1947
Sørensen et al. 2013
Chondrichthyes - Selachii
Selachii indet.
Erikkson et al. 2011
coprolite morphotypes 3, 4
Chondrichthyes - Heterodontiformes - Heterodontidae
Heterodontus informal sp. 1 de Blainville 1816
Sørensen et al. 2013
Heterodontus informal sp. 2 de Blainville 1816
Sørensen et al. 2013
Chondrichthyes - Squatinidae
Squatina spp.
Sørensen et al. 2013
Squatina (Squatina) fortemordeo n. sp. Siversson et al. 2015
Siversson et al. 2015
Squatina (Squatina) lundegreni n. sp. Siversson et al. 2015
Siversson et al. 2015
Chondrichthyes - Squalidae
Squalidae spp.
Sørensen et al. 2013
Chondrichthyes - Carcharhiniformes - Triakidae
? Paratriakis sp.
Sørensen et al. 2013
Galeorhinus sp. de Blainville and Prévost 1816
Sørensen et al. 2013
Palaeogaleus sp. Gurr 1963
Sørensen et al. 2013
Chondrichthyes - Carcharhiniformes - Scyliorhinidae
Scyliorhinidae informal sp. 1 Gill 1862
Sørensen et al. 2013
Scyliorhinus germanicus Herman 1982
Sørensen et al. 2013
recombined as Crassescyliorhinus germanicus
Listed as ‘Scyliorhinus’ germanicus
Chondrichthyes - Orectolobiformes - Orectolobidae
Cretorectolobus sp. Case 1978
Sørensen et al. 2013
Cederstroemia nilsi n. sp. Siverson 1995
Siverson 1995
Chondrichthyes - Orectolobiformes - Parascylliidae
Pararhincodon spp.
Sørensen et al. 2013
Chondrichthyes - Orectolobiformes - Hemiscylliidae
Chiloscyllium sp. Müller and Henle 1837
Sørensen et al. 2013
Hemiscyllium hermani Müller 1989
Sørensen et al. 2013
Chondrichthyes - Lamniformes - Serratolamnidae
Serratolamna sp. Landemaine 1991
Sørensen et al. 2013
Chondrichthyes - Lamniformes - Pseudoscapanorhynchidae
Cretodus borodini (Cappetta and Case 1975)
Siverson 1992
recombined as Protolamna borodini
Chondrichthyes - Lamniformes - Carchariidae
Carcharias latus (Davis 1890)
Siverson 1992
Carcharias tenuis (Davis 1890)
Siverson 1992
Carcharias aasenensis Siverson 1992
Siverson 1992
Chondrichthyes - Lamniformes - Archaeolamnidae
Archaeolamna kopingensis (Davis 1890)
Siverson 1992
Chondrichthyes - Lamniformes - Mitsukurinidae
Scapanorhynchus perssoni Siverson 1992
Siverson 1992
Anomotodon hermani Siverson 1992
Siverson 1992
Chondrichthyes - Lamniformes - Alopiidae
Paranomotodon sp. Herman 1975
Siverson 1992
Chondrichthyes - Lamniformes - Anacoracidae
Squalicorax kaupi (Agassiz 1843)
Siverson 1992
Chondrichthyes - Lamniformes - Otodontidae
Cretalamna borealis (Priem 1897)
Siversson et al. 2015
recombined as Cretolamna borealis
Part of C. appendiculata in Siverson, 1992
Cretolamna appendiculata (Agassiz 1835)
Siverson 1992
    = Cretalamna sarcoportheta n. sp. Siversson et al. 2015
Siversson et al. 2015
Part of C. appendiculata
Chondrichthyes - Lamniformes - Pseudocoracidae
Pseudocorax laevis (Leriche 1906)
Sørensen et al. 2013
Chondrichthyes - Synechodontiformes - Paraorthacodontidae
Paraorthacodus andersoni (Case 1978)
Paraorthacodus conicus Davis 1890
Chondrichthyes - Synechodontiformes - Palaeospinacidae
Synechodus informal sp. 1 Woodward 1888
Sørensen et al. 2013
Synechodus informal sp. 2 Woodward 1888
Sørensen et al. 2013
Synechodus filipi n. sp. Siversson et al. 2015
Siversson et al. 2015
Osteichthyes
Osteichthyes indet. Huxley 1880
Erikkson et al. 2011
coprolite morphotype 5
Actinopteri - Lepisosteiformes - Lepisosteidae
Lepisosteidae indet. Cuvier 1825
Bazzi et al. 2015
Actinopteri
Teleostei indet. (Müller 1846)
Bazzi et al. 2015
Actinopteri - Aulopiformes - Enchodontidae
Enchodus sp. Agassiz 1835
Bazzi et al. 2015
Actinopteri - Crossognathiformes - Pachyrhizodontidae
cf. Pachyrhizodus sp. Agassiz 1850
Bazzi et al. 2015
Actinopteri - Ichthyodectiformes - Ichthyodectidae
Ichthyodectidae indet. Crook 1892
Bazzi et al. 2015
Actinopteri - Pachycormiformes - Pachycormidae
Protosphyraena sp. Leidy 1857
Bazzi et al. 2015
Actinopteri - Pycnodontiformes - Pycnodontidae
Pycnodontidae indet. (Agassiz 1833)
Bazzi et al. 2015
Anomoeodus subclavatus (Agassiz 1833)
Bazzi et al. 2015
Reptilia - Mosasauridae
? Platecarpus sp. Cope 1869
Sørensen et al. 2013
Tylosaurus ivoensis (Russell 1967)
Lindgren and Siverson 2002
recombined as Mosasaurus ivoensis
Hainosaurus sp. Dollo 1885
Lindgren 2005
Dollosaurus sp. Yakovlev 1901
Sørensen et al. 2013
synonym of Prognathodon
7 unnumbered teeth, Lund University
Clidastes propython Cope 1869
Lindgren and Siverson 2004
100 marginal teeth (LO 8231t, 8232t and 86 unnumbered, Lund University; RM PZ R1776, 1777, 1794–1803), 13 pterygoid teeth (RM PZ R1804, 1805; 11 unnumbered, Lund University), four cervical vertebrae (RM PZ R1778, 1779, 1782, 1789), six cervical-dorsal vertebrae (RM PZ R1783–1788), two dorsal vertebrae (RM PZ R1780, 1781) and 11 caudal vertebrae (RM PZ R1806–1816)
Eonatator sternbergi
Sørensen et al. 2013
Halisaurus sternbergi (Wiman 1920)
Lindgren and Siverson 2005
recombined as Eonatator sternbergii
58 teeth (RM PZ R1811-1819; 49 unnumbered) and one caudal vertebra (RM PZ R1810)
Reptilia - Leptoceratopsidae
Leptoceratopsidae indet. (Nopcsa 1923)
Lindgren et al. 2007 7 specimens
RM PZ R1827-R1834; LO 12044t, right manual phalanx
Reptilia
Ornithopoda indet. (Marsh 1881)
Poropat et al. 2016 2 specimens
LO 12045t, left pedal phalanx; LO 12046t, left pedal phalanx
Reptilia - Plesiosauria - Elasmosauridae
Scanisaurus sp. Persson 1960
Sørensen et al. 2013
Reptilia - Testudines
Chelonia indet. (Latreille 1800)
Sørensen et al. 2013
synonym of Testudines
Cephalopoda - Belemnitida - Belemnitellidae
Belemnitella mucronata (Schlotheim 1813)
Lindgren and Siverson 2002
Cephalopoda - Belemnitida
Belemnitida indet. Zittel 1895
Lindgren et al. 2007
    = Belemnellocamax mammillatus
Erikkson et al. 2011
Gonioteuthis quadrata
Lindgren and Siverson 2002
Bivalvia - Ostreida - Ostreidae
Ostreidae indet. Rafinesque 1815
Lindgren et al. 2007
see common names

Geography
Country:Sweden State/province:Skåne
Coordinates: 56.1° North, 14.5° East (view map)
Paleocoordinates:46.6° North, 14.5° East
Basis of coordinate:stated in text
Geographic resolution:outcrop
Time
Period:Cretaceous Epoch:Late/Upper Cretaceous
Stage:Campanian 10 m.y. bin:Cretaceous 7
Key time interval:Early/Lower Campanian Ammonoid zone: Belemnellocamax mammillatus
Age range of interval:83.60000 - 72.10000 m.y. ago
Stratigraphy
Stratigraphic resolution:group of beds
Stratigraphy comments: from two beds in lower part (1 m) of section; the "marine sand unit"; latest Early Campanian
Lithology and environment
Primary lithology:shelly/skeletal unlithified calcareous sandstone
Secondary lithology:coarse,shelly/skeletal,green unlithified sandstone
Includes fossils?Y
Includes fossils?Y
Lithology description: small tooth from "highly calcareous quartz sand with extremely abundant and well-preserved belemnite guards and articulated oysters" from uppermost 0.5 m of uppermost part of sequence; remainder from "a mottled, 10-cm thick storm deposit comprising greenish, coarse quartz sand mixed with fragmented oysters, belemnites and common vertebrate remains."
Environment:coastal indet.
Geology comments: shallow water marine storm deposit
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils,mesofossils
Preservation of anatomical detail:good
Disassociated major elements:many
Disassociated minor elements:some
Fragmentation:occasional
Spatial resolution:allochthonous
Collection methods and comments
Collection methods:bulk,selective quarrying,chemical,field collection
Minimum sieve size:2.500
Reason for describing collection:general faunal/floral analysis
Collection dates:1990s, 2001, 2010
Metadata
Also known as:Asen
Database number:74334
Authorizer:M. Carrano, P. Mannion, M. Clapham Enterer:M. Carrano, P. Mannion, M. Clapham, J. Tennant
Modifier:M. Carrano Research group:vertebrate
Created:2007-08-02 14:33:14 Last modified:2022-03-10 09:56:07
Access level:the public Released:2007-08-02 14:33:14
Creative Commons license:CC BY
Reference information

Primary reference:
73893. M. Siverson. 1992. Late Cretaceous Paraorthacodus (Palaeospinacidae, Neoselachii) from Sweden. Journal of Paleontology 66:994-1001 [M. Clapham/M. Clapham]

Secondary references:

77703 M. Bazzi, E. Einarsson, and B. P. Kear. 2015. Late Cretaceous (Campanian) actinopterygian fishes from the Kristianstad Basin of southern Sweden. In B. P. Kear, J. Lindgren, J. H. Hurum, J. Milàn, V. Vajda (eds.), Mesozoic Biotas of Scandinavia and its Arctic Territories. Geological Society of London Special Publications 434:277-292 [M. Clapham/M. Clapham]
37143ETE M. E. Erikkson, J. Lindgren, K. Chin and U. Månsby. 2011. Coprolite morphotypes from the Upper Cretaceous of Sweden: novel views on an ancient ecosystem and implications for coprolite taphonomy. Lethaia [M. Carrano/M. Carrano]
34814 J. Lindgren. 2005. The first record of Hainosaurus (Reptilia: Mosasauridae) from Sweden. Journal of Paleontology 79(6):1157-1165 [M. Carrano/H. Street]
46318 J. Lindgren. 2005. Dental and vertebral morphology of the enigmatic mosasaur Dollosaurus (Reptilia, Mosasauridae) from the lower Campanian (Upper Cretaceous) of southern Sweden. Bulletin of the Geological Society of Denmakr 52:17-25 [P. Mannion/J. Tennant]
24962ETE J. Lindgren, P. J. Currie, M. Siverson, J. Rees, P. Cederström and F. Lindgren. 2007. The first neoceratopsian dinosaur remains from Europe. Palaeontology 50(4):929-937 [M. Carrano/M. Carrano]
44250 J. Lindgren and M. Siverson. 2002. Tylosaurus ivoensis: a giant mosasaur from the early Campanian of Sweden. Transactions of the Royal Society of Edinburgh: Earth Sciences 93:73-93 [P. Mannion/J. Tennant]
38113 J. Lindgren and M. Siverson. 2004. The first record of the mosasaur Clidastes from Europe and its palaeogeographical implications. Acta Palaeontologica Polonica 49(2):219-234 [P. Mannion/P. Mannion]
37934 J. Lindgren and M. Siverson. 2005. Halisaurus sternbergi, a small mosasaur with an intercontinental distribution. Journal of Paleontology 79(4):763-773 [P. Mannion/P. Mannion]
80639 S. F. Poropat, E. Einarsson, J. Lindgren, M. Bazzi, C. Lagerstam and B. P. Kear. 2016. Late Cretaceous dinosaurian remains from the Kristianstad Basin of southern Sweden. In B. P. Kear, J. Lindgren, J. H. Hurum, J. Milàn, V. Vajda (eds.), Mesozoic Biotas of Scandinavia and its Arctic Territories. Geological Society, London, Special Publications 434 [M. Carrano/M. Carrano]
73902 M. Siverson. 1992. Biology, dental morphology and taxonomy of lamniform sharks from the Campanian of the Kristianstad Basin, Sweden. Palaeontology 35:519-554 [M. Clapham/M. Clapham]
74444 M. Siverson. 1995. Revision of Cretorectolobus (Neoselachii) and description of Cederstroemia n. gen., a Cretaceous carpet shark (Orectolobiformes) with a cutting dentition. Journal of Paleontology 69:974-979 [M. Clapham/M. Clapham]
74344 M. Siversson, T. D. Cook, P. Cederström and H. E. Ryan. 2015. Early Campanian (Late Cretaceous) squatiniform and synechodontiform selachians from the Åsen locality, Kristianstad Basin, Sweden. Mesozoic Biotas of Scandinavia and its Arctic Territories. Geological Society, London, Special Publications 434:251-275 [M. Clapham/M. Clapham]
74347 M. Siversson, J. Lindgren, M. G. Newbrey, P. Cederström, and T. D. Cook. 2015. Cenomanian–Campanian (Late Cretaceous) mid-palaeolatitude sharks of Cretalamna appendiculata type. Acta Palaeontologica Polonica 60:339-384 [M. Clapham/M. Clapham]
45317 A. M. Sørensen, F. Surlyk, and J. Lindgren. 2013. Food resources and habitat selection of a diverse vertebrate fauna from the upper lower Campanian of the Kristianstad Basin, southern Sweden. Cretaceous Research 42(1):85-92 [P. Mannion/P. Mannion/M. Carrano]