La Neuve: Middle Campanian, France
collected by Kerourio 1970

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Rognacia abbreviata
Garcia et al. 2000 1 specimen
Palaeostoa sp.
Garcia et al. 2000 1 specimen
Palaeostoa cf. dilatata
Garcia et al. 2000 3 specimens
Macrophysa sp.
Garcia et al. 2000 4 specimens
cf. Megalostoma julliani
Garcia et al. 2000
Palaeocyclophorus sp.
Garcia et al. 2000
Paracandona occitanica
Garcia et al. 2000
Amphibia - Temnospondyli
Anura indet. (Fischer von Waldheim 1813)
Garcia et al. 2000
Amphibia - Temnospondyli - Discoglossidae
Discoglossidae indet. Günther 1859
Garcia et al. 2000 170 specimens
Amphibia - Temnospondyli - Palaeobatrachidae
Palaeobatrachidae indet. Cope 1865
Garcia et al. 2000 16 specimens
Amphibia - Temnospondyli
Caudata indet. Scopoli 1777
Garcia et al. 2000 23 specimens
Amphibia - Temnospondyli - Albanerpetontidae
Albanerpetontidae indet. Fox and Naylor 1982
Garcia et al. 2000 20 specimens
Reptilia - Nodosauridae
? Struthiosaurus sp. Bunzel 1870
Garcia et al. 2000
Reptilia - Ankylosauridae
? Ankylosauridae indet. Brown 1908
Garcia et al. 2000
Reptilia - Rhabdodontidae
Rhabdodon aff. robustus (Nopcsa 1900)
Garcia et al. 2000 2 specimens
recombined as Zalmoxes robustus
Reptilia - Theropoda
Theropoda indet. Marsh 1881
Garcia et al. 2000 11 specimens
"eggshell with prismatic structural morphotype"
Reptilia - Laevisoolithidae
Laevisoolithidae indet. Mikhailov 1991
Garcia et al. 2000
"eggshell with ratite structural morphotype"
Reptilia
cf. Paronychodon sp. Cope 1876
Garcia et al. 2000 2 specimens
Reptilia - Dromaeosauridae
Deinonychosauria indet. Colbert and Russell 1969
1 specimen
NEV 12
    = Dromaeosauridae indet. Colbert and Russell 1969
Garcia et al. 2000
    = cf. Richardoestesia sp. Currie et al. 1990
Isasmendi et al. 2022
Reptilia - Krokolithidae
Krokolithidae indet. Kohring and Hirsch 1996
Garcia et al. 2000
eggshells
Reptilia
? Alligatoridae indet. Gray 1844
Garcia et al. 2000 1 specimen
    = Acynodon sp. Buscalioni et al. 1997
Puértolas-Pascual et al. 2016
Acynodon iberoccitanus Buscalioni et al. 1997
Garcia et al. 2000
Reptilia - Sauria
? Sauria indet. MacCartney 1802
Garcia et al. 2000 1 specimen
Reptilia - Lacertilia
Lacertilia indet. Owen 1842
Garcia et al. 2000 4 specimens
vertebrae, "eggshell with gekkonoid structural morphotype"
Reptilia - Testudines
Chelonia indet. (Latreille 1800)
Garcia et al. 2000
synonym of Testudines
Mammalia
? Mammalia indet. Linnaeus 1758
Garcia et al. 2000 1 specimen
Actinopteri - Lepisosteiformes - Lepisosteidae
Lepisosteus sp. Agassiz 1843
Garcia et al. 2000
Gastropoda - Heterostropha - Physidae
Physa cf. patula
Garcia et al. 2000 3 specimens
Physa aff. delecta
Garcia et al. 2000 3 specimens
Gastropoda - Heterostropha - Acteonidae
? Pupilla sp. Leach 1828
Garcia et al. 2000 1 specimen
spelled with current rank as Pupa (Pupilla)
Bivalvia - Cardiida - Sphaeriidae
? Pisidiidae indet. Gray 1857
Garcia et al. 2000 2 specimens
spelled with current rank as Pisidiinae
Ostracoda
Ostracoda indet. Latreille 1802
Garcia et al. 2000 3 specimens
Charales - Characea
Amblyochara begudiana Grambast 1962
Garcia et al. 2000
Peckichara pectinata
Garcia et al. 2000
"forme naine"
see common names

Geography
Country:France State/province:Provence-Alpes-Côte d'Azur County:Bouches-du-Rhône
Coordinates: 43.5° North, 5.7° East (view map)
Paleocoordinates:34.1° North, 6.5° East
Basis of coordinate:based on nearby landmark
Geographic resolution:small collection
Time
Period:Cretaceous Epoch:Late/Upper Cretaceous
Stage:Campanian 10 m.y. bin:Cretaceous 7
Key time interval:Middle Campanian
Age range of interval:83.60000 - 72.10000 m.y. ago
Stratigraphy
Stratigraphic resolution:bed
Stratigraphy comments: a subhorizontal layer in the Arc syncline; terminal/upper "Begudian"; no formation name given; middle Campanian following Fondevilla et al. (2019)
Lithology and environment
Primary lithology:gray sandstone
Secondary lithology:red,yellow argillaceous,carbonaceous marl
Includes fossils?Y
Lithology description: "an alternation of two meter-scale gray sandstone banks, and thicker ochre-red argillaceous marl beds. The sandstones are relatively thick; the clays contain silica grains and carbonate nodules. The fossiliferous layer is included high in the clay layer delimited by the sandstone banks."
Environment:lacustrine indet.
Geology comments: JA: presence of freshwater charophytes, gastropods, and ostracods in a claystone indicates a lacustrine setting
Taphonomy
Modes of preservation:body,mold/impression
Lagerst�tten type:concentrate
Degree of concentration:concentrated
Size of fossils:macrofossils,mesofossils,microfossils
Preservation of anatomical detail:good
Associated major elements:some
Disassociated major elements:many
Disassociated minor elements:many
Fragmentation:frequent
Collection methods and comments
Collection methods:field collection
Reason for describing collection:general faunal/floral analysis
Collectors:Kerourio Collection dates:1970
Metadata
Database number:59269
Authorizer:M. Carrano, P. Mannion Enterer:M. Carrano, P. Mannion
Modifier:M. Carrano Research group:vertebrate
Created:2006-03-22 06:50:11 Last modified:2021-05-07 09:52:42
Access level:the public Released:2006-03-22 06:50:11
Creative Commons license:CC BY
Reference information

Primary reference:
25139.ETE E. Buffetaut, B. Marandat, and B. Sigé. 1986. Découverte de dents de deinonychosaures (Saurischia, Theropoda) dans le Crétacé superieur du Sud de la France [Discovery of deinonychosaur teeth (Saurischia, Theropoda) in the Upper Cretaceous of southern France]. Comptes Rendus de l'Académie des Sciences à Paris, Série II 303:1393-1396 [M. Carrano/M. Carrano]

Secondary references:

45554 E. Buffetaut, G. Cuny, and J. Le Loeuff. 1991. French dinosaurs: the best record in Europe?. Modern Geology 16:17-42 [M. Carrano/M. Carrano]
16950ETE G. Garcia. 2000. Diversité des coquilles "minces" d'oeufs fossiles du Crétacé supérieur du sud de la France [Diversity of fossil eggshell fragments from the Upper Cretaceous of southern France]. Géobios 33(1):113-126 [M. Carrano/M. Carrano/J. Alroy]
16947ETE G. Garcia, S. Duffaud, M. Feist, B. Marandat, Y. Tambareau, J. Villatte, and B. Sigé. 2000. La Neuve, gisement à plantes, invertébrés et vertébrés du Bégudien (Sénonien supérieur continental) du bassin d'Aix-en-Provence [La Neuve, Aix-en-Provence Basin, SE France, a new fossil site bearing plants, invertebrates and vertebrates of Begudian age (continental Late Senonian)]. Geodiversitas 22(3):325-348 [M. Carrano/M. Carrano]
52039 J. D. Gardner and M. Böhme. 2008. Review of the Albanerpetontidae (Lissamphibia), with comments on the paleoecological preferences of European Tertiary albanerpetontids. In J. T. Sankey & S. Baszio (ed.), Vertebrate Microfossil Assemblages: Their Role in Paleoecology and Paleobiogeography 178-218 [M. Carrano/M. Carrano]
80599 E. Isasmendi, A. Torices, J. I. Canudo, P. J. Currie, and X. Pereda-Suberbiola. 2022. Upper Cretaceous European theropod palaeobiodiversity, palaeobiogeography and the intra‐Maastrichtian faunal turnover: new contributions from the Iberian fossil site of Laño. Papers in Palaeontology 8(1):e1419:1-38 [M. Carrano/M. Carrano]
76284 E. Puértolas-Pascual, A. Blanco, C. A. Brochu and J. I. Canudo. 2016. Review of the Late Cretaceous-early Paleogene crocodylomorphs of Europe: Extinction patterns across the K-PG boundary. Cretaceous Research 57:565-590 [P. Mannion/P. Mannion]
75349 A. G. Sellés, B. Vila, and A. Galobart. 2014. Diversity of theropod ootaxa and its implications for the latest Cretaceous dinosaur turnover in southwestern Europe. Cretaceous Research 49:45-54 [M. Carrano/M. Carrano/P. Mannion]