Armuña, Segovia: Late/Upper Campanian, Spain
collected 1980

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Actinopteri - Lepisosteiformes - Lepisosteidae
Semionotiformes indet. Arambourg and Bertin 1958
Hernández et al. 2007
    = Lepisosteidae indet. Cuvier 1825
Pérez-García et al. 2016
UPUAM specimens
Teleostei indet. Müller 1846
Hernández et al. 2007
    = Lepisosteidae indet. Cuvier 1825
Pérez-García et al. 2016
UPUAM specimens
Reptilia - Theropoda
Theropoda indet. Marsh 1881
Sanz and Buscalioni 1987
UPUAM 14066, 14067 and 14069
Reptilia - Abelisauridae
cf. Arcovenator sp. Tortosa et al. 2014
Pérez-García et al. 2016
UPUAM 14044-14049
Reptilia - Rhabdodontidae
Ornithischia indet. Seeley 1888
2 specimens
    = Rhabdodon sp. Matheron 1869
Hernández et al. 2007
UP-AUM AR04, AR05
    = cf. Rhabdodon sp. Matheron 1869
Pérez-García et al. 2016
UPUAM AR04 and 05
Reptilia
Eusauropoda indet. Upchurch 1995
Pérez-García et al. 2016
UPUAM 14054 and 14055
Neosauropoda indet. Bonaparte 1986
Pérez-García et al. 2016
UPUAM 14057
Titanosauridae indet. Lydekker 1885
Le Loeuff 1993 3 specimens
    = Titanosauria indet. Bonaparte and Coria 1993
Hernández et al. 2007
UP-AUM AR03
    = Lithostrotia indet. Wilson and Upchurch 2003
Pérez-García et al. 2016
UPUAM 13951, 13952, 13956, 13957 and 14050-14053
Crocodylia indet. (Owen 1842)
Sanz and Buscalioni 1987 2 specimens
    = Allodaposuchus precedens Nopcsa 1928
Buscalioni et al. 2001
UPUAM A1 and A2
    = Eusuchia indet. Huxley 1875
Pérez-García et al. 2016
Musturzabalsuchus sp. Buscalioni et al. 1997
Hernández et al. 2007
UPUAM 502
    = Eusuchia indet. Huxley 1875
Pérez-García et al. 2016
Anguimorpha indet. (Fürbringer 1900)
Pérez-García et al. 2016
UPUAM 14056
Mosasauroidea indet. ()
Pérez-García et al. 2016
spelled with current rank as Mosasauria
STUS 14567
Reptilia - Testudines
Chelonia indet. (Latreille 1800)
Sanz and Buscalioni 1987
    = Dortoka sp. Lapparent de Broin and Murelaga 1996
Hernández et al. 2007
    = Dortoka vasconica Lapparent de Broin and Murelaga 1996
Pérez-García et al. 2016
UPUAM 14009-14011 and others
Iberoccitanemys convenarum (Laurent et al. 2002)
Pérez-García et al. 2016
synonym of Iberoccitanemys atlanticum
STUS 12111, STUS 12113-12115, STUS 12117-12119, STUS 14599-14608, STUS 14616, UPUAM 14012-14036
Reptilia
Hernández et al. 2007
STUS 12161, STUS 14588-14592, UPUAM 14005-14008
see common names

Geography
Country:Spain State/province:Castilla y Leon County:Segovia
Coordinates: 41.1° North, 4.3° West (view map)
Paleocoordinates:31.7° North, 1.9° West
Basis of coordinate:based on nearby landmark
Altitude:866 meters
Geographic resolution:small collection
Time
Period:Cretaceous Epoch:Late/Upper Cretaceous
Stage:Campanian 10 m.y. bin:Cretaceous 7
Key time interval:Late/Upper Campanian
Age range of interval:83.60000 - 72.10000 m.y. ago
Stratigraphy
Formation:Vegas de Matute
Stratigraphic resolution:bed
Lithology and environment
Primary lithology:"cross stratification",white,yellow argillaceous,sandy "siliciclastic"
Includes fossils?Y
Lithology description: "the fossiliferous section is composed of gravel, sand and clays of white and yellowish colors with cross-stratification"
Environment:delta plain
Geology comments: "a detrital unit...deltaic with freshwater influences."
Taphonomy
Modes of preservation:body
Degree of concentration:dispersed
Size of fossils:macrofossils,mesofossils
Preservation of anatomical detail:medium
Disassociated major elements:some
Disassociated minor elements:some
Collection methods and comments
Collection methods:field collection,survey of museum collection
Reason for describing collection:general faunal/floral analysis
Collection dates:1980s
Collection method comments: discovered during cartographic mapping for the National Geological Map of Spain (MAGNA)
Metadata
Database number:48861
Authorizer:M. Carrano, P. Mannion Enterer:M. Carrano, P. Mannion
Modifier:M. Carrano Research group:vertebrate
Created:2005-04-01 06:11:04 Last modified:2023-01-10 15:03:37
Access level:the public Released:2005-04-01 06:11:04
Creative Commons license:CC BY
Reference information

Primary reference:

30625.ETE J. L. Sanz. 1986. Nouveaux gisements de dinosaures dans le Crétacé espagnol [New dinosaur localities in the Spanish Cretaceous]. Les Dinosaures de La Chine à La France, Colloque International de Paléontologie, Toulouse, France, 2-6 Septembre 1985; Muséum d'Histoire Naturelle de Toulouse, Toulouse 81-88 [M. Carrano/M. Carrano/M. Carrano]

Secondary references:

83613 M. Aurell, D. Torromé, J. M. Gasca, P. Calvín, M. Pérez-Pueyo, J. Parrilla-Bel, E. Medrano-Aguado, C. Martín-Closas, A. Vicente, P. Sierra-Campos, and J. I. Canudo. 2022. Latest Cretaceous palaeogeographic evolution of northeast Iberia: Insights from the Campanian continental Montalbán subbasin (Spain). Earth-Science Reviews 235:10425 [M. Carrano/M. Carrano/P. Mannion]
14444ETE A. D. Buscalioni, F. Ortega, D. B. Weishampel and C. M. Jianu. 2001. A revision of the crocodyliform Allodaposuchus precedens from the Upper Cretaceous of the Hateg Basin, Romania. Its relevance in the phylogeny of Eusuchia. Journal of Vertebrate Paleontology 21(1):74-86 [M. Carrano/M. Carrano/M. Carrano]
17647ETE J. I. Canudo and J. I. Ruiz-Omeñaca. 2003. Los restos directos de dinosaurios teropódos (excluyendo Aves) en España [Direct remains of theropod dinosaurs (excluding Aves) in Spain]. Ciencias de la Tierra 26:347-373 [M. Carrano/M. Carrano]
26656ETE E. C. Hernández, J. L. Sanz, F. Ortega and F. Escaso. 2007. Restos de dinosaurios del Crátacico Superior de Armuña (Segovia) [Remains of dinosaurs from the Upper Cretaceous of Armuña (Segovia)]. Cantera Paleontológica. Diputación Provincial de Cuenca, Cuenca 133-142 [M. Carrano/M. Carrano]
80599 E. Isasmendi, A. Torices, J. I. Canudo, P. J. Currie, and X. Pereda-Suberbiola. 2022. Upper Cretaceous European theropod palaeobiodiversity, palaeobiogeography and the intra‐Maastrichtian faunal turnover: new contributions from the Iberian fossil site of Laño. Papers in Palaeontology 8(1):e1419:1-38 [M. Carrano/M. Carrano]
13022ETE J. Le Loeuff. 1993. European titanosaurids. Revue de Paléobiologie, Volume Spéciale 7:105-117 [M. Carrano/M. Carrano/M. Carrano]
7842 J. Le Loeuff and E. Buffetaut. 1991. Tarascosaurus salluvicus nov. gen., nov. sp., dinosaure théropode du Crétacé supérieur du sud de la France [Tarascosaurus salluvicus nov. gen., nov. sp., a theropod dinosaur from the Upper Cretaceous of southern France]. Géobios 25(5):585-594 [M. Carrano/M. Carrano/M. Carrano]
65863 A. Pérez-García, F. Ortega, A. Bolet, F. Escaso, A. Houssaye, J. Martínez-Salanova, C. Miguel Chaves, P. Mocho, I. Narváez, M. Segura, A. Torices, D. Vidal, and J. L. Sanz. 2016. A review of the upper Campanian vertebrate site of Armuña (Segovia Province, Spain). Cretaceous Research 57:591-623 [P. Mannion/P. Mannion/P. Mannion]
13712ETE J. E. Powell. 2003. Revision of South American titanosaurid dinosaurs: palaeobiological, palaeobiogeographical and phylogenetic aspects. Records of the Queen Victoria Museum Launceston 111:1-173 [M. Carrano/M. Carrano/M. Carrano]
70303 J. L. Sanz. 2000. Yacimientos ibéricos de dinosaurios [Iberian dinosaur localities]. Dinosaurios: Los Señores del Pasado [Dinosaurs: The Lords of the Land] 267-299 [M. Carrano/M. Carrano]
13504ETE J. L. Sanz and A. D. Buscalioni. 1987. New evidence of armoured titanosaurs in the Upper Cretaceous of Spain. P. J. Currie and E. H. Koster (eds.), Fourth Symposium on Mesozoic Terrestrial Ecosystems. Tyrrell Museum of Paleontology, Drumheller, Alberta 199-204 [M. Carrano/M. Carrano/M. Carrano]
19339ETE J. L. Sanz, A. D. Buscalioni, J. J. Moratalla, V. Francés, and M. Antón. 1990. Los reptiles Mesozoicos del registro español [The Mesozoic reptiles from the Spanish record]. Monografias del Museo Nacional de Ciencias Naturales 2:1-79 [M. Carrano/M. Carrano]
81943 J.-L. Sanz, Á D. Buscalioni, B. P. Pérez Moreno, J. Moratalla, and S. Jiménez García. 1992. Los dinosaurios de Castilla y León [The dinosaurs of Castilla y León]. In E. Jiménez-Fuentes (ed.), Vertebrados Fósiles de Castilla y León 47-57 [M. Carrano/M. Carrano]
19710ETE F. Torcida Fernández-Baldor. 2005. Los dinosaurios de Castilla y León [The dinosaurs of Castilla y León]. Patrimonio Histórico de Castilla y León 6(23):23-34 [M. Carrano/M. Carrano/M. Carrano]