Bellevue, locality C3 (C III): Early/Lower Maastrichtian, France
collected by Clottes, Raynaud 1986-

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Plantae indet. Haeckel 1866
Le Loeuff 1995
Reptilia
Titanosauria indet. Bonaparte and Coria 1993
Vila et al. 2012 1 specimen
left femur, MDE C3-1278
Hypselosaurus priscus Matheron 1869
500 specimens
    = Titanosauridae indet. Lydekker 1885
Le Loeuff 1993
"new species"
    = Ampelosaurus atacis n. gen., n. sp. Le Loeuff 1995
Le Loeuff 1995
incl. 6 femora, MDE C3-87, 1482, 78, 287, 182, 20
cf. Lirainosaurus astibiae Sanz et al. 1999
Vila et al. 2012 3 specimens
femora, MDE C3-61, C3-261, C3-203
Reptilia - Nodosauridae
Ankylosauria indet. Osborn 1923
Le Loeuff 2005
dermal plates
    = cf. Struthiosaurus austriacus Bunzel 1871
Laurent et al. 2001
Reptilia - Rhabdodontidae
Rhabdodon priscus (Matheron 1869)
Le Loeuff 1995 100 specimens
Reptilia - Dromaeosauridae
Theropoda indet. Marsh 1881
Le Loeuff 1995
    = Dromaeosauridae indet. Colbert and Russell 1969
Buffetaut and Le Loeuff 1997
    = cf. Variraptor mechinorum Le Loeuff and Buffetaut 1998
Laurent et al. 2001
Reptilia - Gargantuaviidae
Aves indet. Linnaeus 1758
Buffetaut and Le Loeuff 1997 1 specimen
    = Gargantuavis philoinos n. gen., n. sp. Buffetaut and Le Loeuff 1998
Buffetaut and Le Loeuff 1998
Reptilia - Pterosauria
Pterosauria indet. Kaup 1834
Buffetaut 1998
Several poorly preserved hollow bones
Reptilia
cf. Ischyrochampsa meridionalis Vasse 1995
Laurent et al. 2001
nomen dubium belonging to Neosuchia
Eusuchia indet. Huxley 1875
Le Loeuff 1995
    = Allodaposuchus precedens Nopcsa 1928
Buscalioni et al. 2001
    = Allodaposuchidae indet. Narváez et al. 2015
Puértolas-Pascual et al. 2016
Reptilia - Testudines
Testudines indet. Batsch 1788
Le Loeuff 1995
    = Foxemys mechinorum Tong et al. 1998
Gaffney et al. 2006
MDE C3 774, partial skull and shell fragments
Coelacanthimorpha - Coelacanthiformes - Mawsoniidae
Axelrodichthys megadromos Cavin et al. 2016
Cavin et al. 2020 1 specimen
MDE C3-02-57
Actinopteri - Lepisosteiformes - Lepisosteidae
Lepisosteus sp. Agassiz 1843
Le Loeuff 1995
see common names

Geography
Country:France State/province:Languedoc-Roussillon County:Aude
Coordinates: 42.9° North, 2.2° East (view map)
Paleocoordinates:34.8° North, 2.0° East
Basis of coordinate:based on nearby landmark
Altitude:356 meters
Geographic resolution:outcrop
Time
Period:Cretaceous Epoch:Late/Upper Cretaceous
Stage:Maastrichtian 10 m.y. bin:Cretaceous 8
Key time interval:Early/Lower Maastrichtian
Age range of interval:70.60000 - 66.00000 m.y. ago
Stratigraphy
Formation:Marnes Rouges Inférieures Member:Marnes de la Maurine
Stratigraphic resolution:bed
Stratigraphy comments: upper part of the Maurine Red Marls Formation
Lithology and environment
Primary lithology:red marl
Secondary lithology:lenticular,fine,coarse conglomerate
Includes fossils?Y
Lithology description: "fluvial marls interbedded with microconglomeratic lenses"
Environment:fluvial indet.
Geology comments: "meandering and braided fluvio-alluvial setting"
Taphonomy
Modes of preservation:body
Lagerst�tten type:concentrate
Degree of concentration:concentrated
Size of fossils:macrofossils
Preservation of anatomical detail:medium
Abundance in sediment:abundant
Associated major elements:some
Disassociated major elements:some
Disassociated minor elements:some
Fragmentation:frequent
Spatial resolution:parautochthonous
Collection methods and comments
Collection excludes:some macrofossils
Collection methods:selective quarrying,mechanical,field collection
Reason for describing collection:taxonomic analysis
Collectors:Clottes, Raynaud Collection dates:1986-
Collection method comments: also includes "remains of plants and invertebrates"
Metadata
Also known as:Campagne-sur-Aude
Database number:48850
Authorizer:M. Carrano, R. Benson, P. Mannion Enterer:M. Carrano, P. Mannion, R. Benson
Modifier:M. Carrano Research group:vertebrate
Created:2005-03-31 15:44:04 Last modified:2023-11-27 12:44:22
Access level:the public Released:2005-03-31 15:44:04
Creative Commons license:CC BY
Reference information

Primary reference:

30925.ETE P. Clottes and C. Raynaud. 1983. Le gisement à dinosauriens de Campagne-sur-Aude–Esperaza. Observations préliminaires. Premiers résultats [The dinosaur locality at Campagne-sur-Aude–Esperaza. Preliminary observations. First results]. Bulletin de la Société d'Études Scientifiques de l'Aude 83:5-14 [M. Carrano/M. Carrano]

Secondary references:

30137 E. Buffetaut. 1998. Ptérosaures et oisaux du Crétacé supérieur de l'Aude. Bulletin de la Société d'Etudes Scientifiques de l'Aude 98:11-17 [R. Butler/R. Butler]
30133 E. Buffetaut. 2008. Late Cretaceous pterosaurs from France: a review. Zitteliana B 28:249-255 [R. Butler/R. Butler]
43907 E. Buffetaut and J. Le Loeuff. 1989. Les dinosaures du Crétacé supérieur des Corbières, France [The Upper Cretaceous dinosaurs of Corbières, France]. Géochronique 32:4-5 [M. Carrano/M. Carrano]
13026ETE E. Buffetaut and J. Le Loeuff. 1997. Late Cretaceous dinosaurs from the foothills of the Pyrenees. Geology Today 1997(3-4):60-68 [M. Carrano/M. Carrano/M. Carrano]
13042ETE E. Buffetaut, J. Le Loeuff, L. Cavin, S. Duffaud, E. Gheerbrant, Y. Laurent, M. Martin, J.-C. Rage, H. Tong and D. Vasse. 1997. Late Cretaceous non-marine vertebrates from southern France: a review of recent finds. Géobios 20:101-108 [M. Carrano/M. Carrano/J. Alroy]
13031ETE E. Buffetaut and J. Le Loeuff. 1998. A new giant ground bird from the Upper Cretaceous of southern France. Journal of the Geological Society, London 155:1-4 [M. Carrano/M. Carrano/M. Carrano]
14444ETE A. D. Buscalioni, F. Ortega, D. B. Weishampel and C. M. Jianu. 2001. A revision of the crocodyliform Allodaposuchus precedens from the Upper Cretaceous of the Hateg Basin, Romania. Its relevance in the phylogeny of Eusuchia. Journal of Vertebrate Paleontology 21(1):74-86 [M. Carrano/M. Carrano/M. Carrano]
76814 L. Cavin, E. Buffetaut, Y. Dutour, G. Garcia, J. Le Loeuff, A. Méchin, P. Méchin, H. Tong, T. Tortosa, E. Turini, and X. Valentin. 2020. The last known freshwater coelacanths: new Late Cretaceous mawsoniid remains (Osteichthyes: Actinistia) from southern France. PLoS One 15(6):e0234183:1-19 [M. Carrano/M. Carrano]
74890 V. Díez Díaz, G. Garcia, X. Pereda Suberbiola, B. Jentgen-Ceschino, K. Stein, P. Godefroit, and X. Valentin. 2021. A new titanosaur (Dinosauria: Sauropoda) from the Upper Cretaceous of Velaux-La-Bastide Neuve (southern France). Historical Biology 33(11):2998-3017 [P. Mannion/P. Mannion/M. Carrano]
86641 V. Fondevilla, J. Dinarès-Turell, R. Edstrada, J. Le Loeuff, and O. Oms. 2014. Integrating the Pyrenean dinosaurs in a geological framework: the Aude section (Campanian-Maastrichtian, southern France). In G. d. J. A. Rodríquez, J. C. Lallena, E. D. Berenguer, J. G. García, A. L. Lisarri, J. P. Bel, E. P. Pascual, & R. S. Casal (eds.), New Insights on Ancient Life. XII EJIP, Boltaña, Sobrarbe 76-79 [M. Carrano/M. Carrano]
27976 E. S. Gaffney, H. Tong, and P. A. Meylan. 2006. Evolution of the side-necked turtles: The families Bothremydidae, Euraxemydidae, and Araripemydidae. Bulletin of the American Museum of Natural History 300:1-318 [M. Uhen/M. Uhen/M. Uhen]
80599 E. Isasmendi, A. Torices, J. I. Canudo, P. J. Currie, and X. Pereda-Suberbiola. 2022. Upper Cretaceous European theropod palaeobiodiversity, palaeobiogeography and the intra‐Maastrichtian faunal turnover: new contributions from the Iberian fossil site of Laño. Papers in Palaeontology 8(1):e1419:1-38 [M. Carrano/M. Carrano]
59044 Y. Laurent, J. Le Loeuff, M. Bilotte, E. Buffetaut, and G. S. Odin. 2001. Campanian-Maastrichtian continental-marine connection at the Aquitaine-Pyrenees-Provence area (S France). In G. S. Odin (ed.), The Campanian-Maastrichtian Boundary. Developments in Palaeontology and Stratigraphy 19:657-674 [M. Carrano/M. Carrano]
13022ETE J. Le Loeuff. 1993. European titanosaurids. Revue de Paléobiologie, Volume Spéciale 7:105-117 [M. Carrano/M. Carrano/M. Carrano]
13019ETE J. Le Loeuff. 1995. Ampelosaurus atacis (nov, gen., nov. sp.), un nouveau Titanosauridae (Dinosauria, Sauropoda) du Crétacé supérieur de la Haute Vallée de l'Aude (France) [Ampelosaurus atacis (nov, gen., nov. sp.), a new titanosaurid (Dinosauria, Sauropoda) from the Upper Cretaceous of the Upper Aude Valley (France)]. Comptes Rendus de l'Académie des Sciences à Paris, Série IIa 321:693-699 [M. Carrano/M. Carrano/M. Carrano]
36728ETE J. Le Loeuff. 1998. Les dinosaures du Crétacé supérieur de l'Île Ibéro-Armoricaine [The dinosaurs of the Upper Cretaceous of the Ibero-Armorican Island]. In A. M. G. de Carvalho, A. M. Andrade, V. F. dos Santos, J. Cascalho, R. Taborda, (eds.), I Encontro Internacional sobre Paleobiologia dos Dinossáurios. Museu Nacional de História Natural, Lisboa 49-64 [M. Carrano/M. Carrano]
19511ETE J. Le Loeuff. 2005. Osteology of Ampelosaurus atacis (Titanosauria) from southern France. In K. Carpenter and V. Tidwell (eds.), Thunder-Lizards: The Sauropodomorph Dinosaurs. Indiana University Press, Bloomington 115-137 [M. Carrano/M. Carrano]
76797 J. Le Loeuff. 2010. Late Cretaceous sauropods of southwestern Europe: Quid novi?. 8th Meeting of the European Association of Vertebrate Palaeontologists. Abstract Volume 50-51 [M. Carrano/M. Carrano/M. Carrano]
67934 J. Le Loeuff and E. Buffetaut. 1995. The evolution of Late Cretaceous non-marine vertebrate fauna in Europe. Sixth Symposium on Mesozoic Terrestrial Ecosystems and Biota, Short Papers 181-184 [M. Carrano/M. Carrano]
36767ETE J. Le Loeuff, E. Buffetaut, L. Cavin, M. Martin, V. Martin and H. Tong. 1995. An armored titanosaurid sauropod from the Late Cretaceous of southern France and the occurrence of osteoderms in the Titanosauridae. In M. G. Lockley, V. F. dos Santos, C. A. Meyer, & A. P. Hunt (eds.), Aspects of Sauropod Paleobiology. GAIA 10:155-159 [M. Carrano/M. Carrano]
14749ETE J. Le Loeuff and E. Buffetaut. 1996. Late Cretaceous dromaeosaurids from southern France. Journal of Vertebrate Paleontology 16(3, suppl.):48A [M. Carrano/M. Carrano]
36369ETE J. Le Loeuff and E. Buffetaut. 1996. Ampelosaurus atacis, the dinosaur of the wineyard. The Dinosaur Society UK Quarterly 1(4):1-2 [M. Carrano/M. Carrano]
76450 M. Pincemaille-Quillevere. 2000. Rhabdodon priscus (Ornithopoda, Dinosauria) sites from southern France. In M.-C. Buchy (ed.), 5th European Workshop on Vertebrate Palaeontology, Program. Abstracts. Excursion GuidesGiganotosaurus carolinii 61-62 [M. Carrano/M. Carrano]
13712ETE J. E. Powell. 2003. Revision of South American titanosaurid dinosaurs: palaeobiological, palaeobiogeographical and phylogenetic aspects. Records of the Queen Victoria Museum Launceston 111:1-173 [M. Carrano/M. Carrano/M. Carrano]
76284 E. Puértolas-Pascual, A. Blanco, C. A. Brochu and J. I. Canudo. 2016. Review of the Late Cretaceous-early Paleogene crocodylomorphs of Europe: Extinction patterns across the K-PG boundary. Cretaceous Research 57:565-590 [P. Mannion/P. Mannion]
55429 B. Vila, A. Galobart, J. I. Canudo, J. Le Loeuff, J. Dinarès-Turell, V. Riera, O. Oms, T. Tortosa, and R. Gaete. 2012. The diversity of sauropod dinosaurs and their first taxonomic succession from the latest Cretaceous of southwestern Europe: Clues to demise and extinction. Palaeogeography, Palaeoclimatology, Palaeoecology 350–352:19-38 [M. Carrano/M. Carrano]