RTMP BB 31, DPP: Late/Upper Campanian, Canada
collected by R. A. Mussieux & D. Tanke 1984

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Gastropoda
Prosobranchia indet. Milne-Edwards 1848
Eberth et al. 1988
rare
Bivalvia - Cardiida - Sphaeriidae
Sphaerium recticardinale
Eberth et al. 1988
Bivalvia - Unionida - Unionidae
Unionidae indet. Rafinesque 1820
Eberth et al. 1988
Chondrichthyes - Rajiformes - Rhinobatidae
Myledaphus bipartitus Cope 1876
54 elements
Osteichthyes
Osteichthyes informal indet. A Huxley 1880
Brinkman 1990 255 elements
Acipenseriformes - Acipenseridae
Acipenser albertensis Lambe 1902
Brinkman 1990 1 element
Amphibia - Temnospondyli
Anura indet. (Fischer von Waldheim 1813)
60 specimens
Anura informal indet. 3 (Fischer von Waldheim 1813)
Gardner and DeMar 2013 1 specimen
TMP 1987.029.0085*, maxilla
Anura informal Unnamed genus and species I
Gardner et al. 2016 1 specimen
TMP 1986.130.0047*, frontoparietal
Tyrrellbatrachus brinkmani Gardner 2015
Gardner 2015 1 specimen
TMP 1986.033.0033 (maxilla)
Hensonbatrachus kermiti Gardner and Brinkman 2015
Gardner and Brinkman 2015 1 specimen
TMP 1985.70.07 (maxilla)
Amphibia - Temnospondyli - Batrachosauroididae
Opisthotriton kayi Auffenberg 1961
Eberth et al. 1988 26 specimens
Amphibia - Temnospondyli - Urodela
Scapherpeton tectum Cope 1876
Eberth et al. 1988 152 specimens
Mammalia
Mammalia indet. Linnaeus 1758
Brinkman 1990 8 specimens
Mammalia - Multituberculata
Multituberculata indet. Cope 1884
Brinkman et al. 1998 3 specimens
Mammalia
Metatheria indet. (Huxley 1880)
Brinkman et al. 1998 1 specimen
Reptilia - Baenidae
Baenidae indet. Cope 1882
1 specimen
Reptilia - Testudines - Chelydridae
Chelydridae indet. Swainson 1839
1 specimen
Reptilia - Testudines - Trionychidae
Aspideretes sp. Hay 1904
Brinkman 1990 29 specimens
    = Trionychinae indet. Gray 1825
Alroy 2007
Reptilia - Lacertilia
Lacertilia indet. Owen 1842
Brinkman 1990 13 specimens
Reptilia
Leidyosuchus canadensis Lambe 1907
Brinkman 1990 13 specimens
Reptilia - Krokolithidae
Krokolithes sp. Hirsch 1985
Zelenitsky and Sloboda 2005
Reptilia
Ankylosauria indet. Osborn 1923
Brinkman 1990 11 specimens
Reptilia - Pachycephalosauridae
Pachycephalosauridae indet. Sternberg 1945
Brinkman et al. 1998 3 specimens
Reptilia - Ceratopsidae
Ceratopsidae indet. Marsh 1888
Brinkman 1990 7 specimens
Reptilia - Spheroolithidae
Spheroolithus albertensis Zelenitsky and Hills 1997
Zelenitsky and Sloboda 2005
Reptilia - Hadrosauridae
Hadrosauridae indet. Cope 1869
Brinkman 1990 104 specimens
Reptilia
cf. Thescelosaurus neglectus Gilmore 1913
Eberth et al. 1988 3 specimens
Reptilia - Theropoda
Theropoda indet. Marsh 1881
Brinkman 1990 5 specimens
Reptilia - Dromaeosauridae
cf. Dromaeosauridae indet. Colbert and Russell 1969
Brinkman 1990 1 specimen
Richardoestesia sp. Currie et al. 1990
Brinkman et al. 1998 2 specimens
Saurornitholestes langstoni Sues 1978
Brinkman 1990 21 specimens
Dromaeosaurus sp. Matthew and Brown 1922
Brinkman et al. 1998 2 specimens
Reptilia
Paronychodon sp. Cope 1876
Brinkman et al. 1998 2 specimens
? Aves indet. Linnaeus 1758
Brinkman et al. 1998 3 specimens
Reptilia - Elongatoolithidae
Dinosauria indet. Owen 1842
Eberth 1990
dinosaur egg shell fragments
    = Continuoolithus canadensis Zelenitsky et al. 1996
Zelenitsky and Sloboda 2005
Porituberoolithus warnerensis Zelenitsky et al. 1996
Zelenitsky and Sloboda 2005
Reptilia
Champsosaurus natator Parks 1933
Brinkman 1990 55 specimens
Actinopteri - Lepisosteiformes - Lepisosteidae
Lepisosteus sp. Agassiz 1843
19 elements
may be same as Atractosteus occidentalis in Table 1?
Actinopteri - Amiiformes - Amiidae
Kindleia sp. Jordan 1927
Eberth et al. 1988 11 elements
synonym of Cyclurus
Actinopteri
Teleostei informal indet. D Müller 1846
Brinkman 1990 51 elements
Teleostei indet. Müller 1846
Brinkman 1990 79 elements
Teleostei informal indet. A Müller 1846
Eberth and Brinkman 1997 3 elements
Teleostei informal indet. N Müller 1846
Eberth and Brinkman 1997 3 elements
Teleostei informal indet. L Müller 1846
Eberth and Brinkman 1997 2 elements
Actinopteri - Salmoniformes
Esocoidea indet. Bleeker 1859
Brinkman 1990 12 elements
Actinopteri - Elopiformes - Phyllodontidae
Paralbula casei Estes 1969
Brinkman 1990 3 elements
Actinopteri - Albuliformes - Albulidae
Coriops amnicolus Estes 1969
Eberth et al. 1988 26 elements
Actinopteri - Aspidorhynchiformes - Aspidorhynchidae
Belonostomus longirostris (Lambe 1902)
1 element
see common names

Geography
Country:Canada State/province:Alberta
Coordinates: 50.8° North, 111.5° West (view map)
Paleocoordinates:58.1° North, 76.0° West
Basis of coordinate:stated in text
Altitude:670 meters
Geographic resolution:outcrop
Time
Period:Cretaceous Epoch:Late/Upper Cretaceous
Stage:Campanian 10 m.y. bin:Cretaceous 7
Key time interval:Late/Upper Campanian
Age range of interval:83.50000 - 70.60000 m.y. ago
Stratigraphy
Geological group:Belly River Formation:Dinosaur Park
Local section:DPP-Brinkman Local bed:1.0 m
Stratigraphic resolution:bed
Stratigraphy comments: Microvertebrate sites of Brinkman (1990) are "...placed in stratigraphic sequence by reference to a disconformity located about sixty meters below the base of the Lethbridge Coal (Eberth, 1990). The stratigraphic position of each site is expressed in terms of meteres above or below the disconformity (Fig. 2, Brinkman, 1990)" where positive numbers are above and negative below.
Lithology and environment
Primary lithology:planar lamination,fine,medium,sideritic silty sandstone
Lithology description: BB 31 occurs within a 2.0 m thick fining-upward lithosome of locally contorted carbonaceous sediments that displays a faint horizontal lamination and locally developed ripple lamination. BB 31 includes locally developed iron-rich (sideritic) sediments associated with concentrated disarticulated pisidiid clam valves and rare unionids. At BB 31 the lower half of the lithosome is dominated by contorted lenses and laminae of silty, fine-to-medium-grained sandstone. The sandstones contain the densest accumulations of microfossils and pisidiid valves as well as a few disarticulated unionid valves and rare gastropod shells.
Environment:crevasse splay
Geology comments: Splay deposits typically comprise stacked sequences of interlaminated sands and siltstones separated by erosional surfaces, thus indicating repeated flooding and aggradation.
Taphonomy
Modes of preservation:body,replaced with siderite
Degree of concentration:concentrated
Size of fossils:macrofossils,mesofossils
Preservation of anatomical detail:good
Disassociated major elements:some
Disassociated minor elements:many
Size sorting:well
Fragmentation:occasional
Temporal resolution:time-averaged
Spatial resolution:autochthonous
Collection methods and comments
Collection excludes:some genera,some macrofossils
Collection methods:bulk,surface (float),sieve,field collection
Minimum sieve size:64.000 Maximum sieve size:81.000
Reason for describing collection:paleoecologic analysis
Museum repositories:TMP
Collectors:R. A. Mussieux & D. Tanke Collection dates:28 August 1984
Collection method comments: BB 31 was the first site in Dinosaur Park to yield egg shell fragments (fragments are <1 cm in maximum dimension).
Taxonomic list comments:"many invertebrate shells"
Metadata
Also known as:Bonebed 31
Database number:47816
Authorizer:R. Whatley, P. Mannion, P. Holroyd, M. Carrano, J. Alroy Enterer:R. Whatley, P. Mannion, P. Holroyd, M. Carrano, J. Alroy
Modifier:M. Carrano Research group:vertebrate
Created:2005-03-02 14:35:37 Last modified:2021-09-22 16:06:55
Access level:the public Released:2005-03-02 14:35:37
Creative Commons license:CC BY
Reference information

Primary reference:

16643.ETE D. B. Brinkman. 1986. Microvertebrate sites: progress and prospectus. In B. G. Naylor (ed.), Field Trip Guidebook to Dinosaur Provincial Park, 2 June 1986. Dinosaur Systematics Symposium, Tyrrell Museum of Palaeontology, Drumheller, Alberta 24-37 [M. Carrano/M. Carrano]

Secondary references:

19636 J. Alroy. 2007. Synonymies and reidentifications of North American fossil vertebrates and so forth. [J. Alroy/J. Alroy]
5929ETE 1710D. B. Brinkman. 1990. Paleontology of the Judith River Formation (Campanian) of Dinosaur National Park, Alberta, Canada: evidence from vertebrate microfossil locality. Palaeogeography, Palaeoclimatology, Palaeoecology 78:37-54 [A. Behrensmeyer/A. Behrensmeyer/M. Carrano]
19348ETE D. B. Brinkman, M. J. Ryan, and D. A. Eberth. 1998. The paleogeographic and stratigraphic distribution of ceratopsids (Ornithischia) in the Upper Judith River Group of western Canada. Palaios 13:160-169 [M. Carrano/M. Carrano]
5930ETE 1711D. A. Eberth. 1990. Stratigraphy and sedimentology of vertebrate microfossil sites in the uppermost Judith River Formation (Campanian), Dinosaur Provincial Park, Alberta, Canada. Palaeogeography, Palaeoclimatology, Palaeoecology 78:1-36 [A. Behrensmeyer/A. Behrensmeyer/M. Kosnik]
78157 D. A. Eberth. 2015. Origins of dinosaur bonebeds in the Cretaceous of Alberta, Canada. Canadian Journal of Earth Sciences 52(8):655-681 [M. Carrano/M. Carrano]
15455ETE D. A. Eberth, D. B. Brinkman, and P. A. Johnston. 1988. Bonebed 31. In D. A. Eberth (ed.), Palaeoecology of Upper Cretaceous Judith River Formation at Dinosaur Provincial Park, Alberta, Canada. Society of Vertebrate Paleontology 48th Annual Meeting, Field Trip "B" (October 12, 1988). Occasional Paper of theTyrrell Museum of Palaeontology 7:23-26 [M. Carrano/M. Carrano]
9017 D. A. Eberth and D. B. Brinkman. 1997. Paleoecology of an estuarine, incised-valley fill in the Dinosaur Park Formation (Judith River Group, Upper Cretaceous) of southern Alberta, Canada. Palaios 12:43-58 [J. Alroy/J. Alroy/J. Alroy]
56016 J. D. Gardner. 2015. An edentulous frog (Lissamphibia; Anura) from the Upper Cretaceous (Campanian) Dinosaur Park Formation of southeastern Alberta, Canada. Canadian Journal of Earth Sciences 52:569-580 [P. Mannion/P. Mannion]
56071 J. D. Gardner and D. B. Brinkman. 2015. A new frog (Lissamphibia, Anura) from the Late Cretaceous of Alberta, Canada. In O. R. P. Bininda-Emonds, G. L. Powell, H. A. Jamniczky, A. M. Bauer, J. Theodor (eds.), All Animals are Interesting: A Festschrift in Honour of Anthony P. Russell 35-105 [P. Mannion/P. Mannion/M. Carrano]
65027 J. D. Gardner and D. G. DeMar. 2013. Mesozoic and Palaeocene lissamphibian assemblages of North America: a comprehensive review. Palaeobiodiversity and Palaeoenvironments 93:459-515 [P. Holroyd/P. Holroyd/M. Carrano]
64015 J. D. Gardner, C. M. Redman, and R. L. Cifelli. 2016. The hopping dead: Late Cretaceous frogs from the middle-late Campanian (Judithian) of western North America. Fossil Imprint 72(1-2):78-107 [P. Holroyd/P. Holroyd/M. Carrano]
54476 D. H. Tanke and M. K. Brett-Surman. 2001. Evidence of hatchling and nestling-size hadrosaurs (Reptilia: Ornithischia) from Dinosaur Provincial Park (Dinosaur Park Formation: Campanian), Alberta. In D. H. Tanke & K. Carpenter (ed.), Mesozoic Vertebrate Life: New Research Inspired by the Paleontology of Philip J. Currie 206-218 [M. Carrano/M. Carrano]
18893ETE D. K. Zelenitsky and W. J. Sloboda. 2005. Eggshells. In P. J. Currie and E. B. Koppelhus (eds.), Dinosaur Provincial Park: A Spectacular Ancient Ecosystem Revealed. Indiana University Press, Bloomington 398-404 [M. Carrano/M. Carrano]