PBDB Collection

CDZh-17a, Dzharakuduk II, Kulbeke: Middle Turonian - Late/Upper Turonian, Uzbekistan
collected by L. A. Nessov; A. Averianov, H.-D. Sues 1977, 1991

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

Mammalia - Spalacotheriidae

	Shalbaatar bakht n. gen., n. sp. Nessov 1997	Nessov 1997
	including holotype manidble (N C.82622)
Mammalia

	Daulestes kulbeckensis n. gen., n. sp. Trofimov and Nessov 1979	Nessov 1982
Mammalia - Zhelestidae

	Sorlestes budan Nessov 1985	Nessov et al. 1998
	synonym of Zhelestes temirkazyk
	CCMGE 3/12953 (right M1)

	Aspanlestes aptap n. gen., n. sp. Nessov 1985	Nessov 1985	2 specimens
	CCMGE 4/12176
Mammalia - Zalambdalestidae

	Kulbeckia kulbecke Nessov 1993	Archibald and Averianov 2003	1 specimen
	CCMGE 5/12176, M2
Reptilia - Lacertilia

	Lacertilia indet. Owen 1842	Nessov 1995	1 specimen
Reptilia - Teiidae

	Teiidae indet. Gray 1827	Nessov 1995	1 specimen
Reptilia

	? Gekkota indet. Cuvier 1817	Nessov 1995	1 specimen
Reptilia - Testudines

	Anatolemys cf. maximus Khozatskii and Nessov 1979	Nessov 1984
Reptilia - Testudines - Trionychidae

	Trionyx informal species 2	Nessov 1984
	high-shelled, big Trionyx
Reptilia - Testudines - Lindholmemydidae

	Lindholmemys elegans n. sp. Riabinin 1935
Reptilia - Testudines - Adocidae

	Adocidae indet. Cope 1869	Danilov and Parham 2005	1 specimen
	a skull (ZISP PH 1/17) referred to Lindholmemys elegans by Nessov 1986 (fig. 11). Danilov & Parham (2005) "hypothesize that ZISP PH 1/17 is the skull of either Shachemys ancestralis or “Adocus” aksary"

	Adocus aksary Nessov and Krasovskaya 1984	Nessov 1984

	cf. Shachemydinae sp. Khosatzky 1977	Nessov 1984
Reptilia - Ornithischia

	Ornithischia indet. Seeley 1888	Nessov 1995	1 specimen
Reptilia

	Ornithopoda indet. Marsh 1881	Nessov 1995	1 specimen

	Gilmoreosaurus arkhangelskyi Nessov 1995	Nessov 1995	8 specimens
	CCMGE 566/12457
	= Levnesovia transoxiana Sues and Averianov 2009	Sues and Averianov 2009
Reptilia - Hadrosauridae

	Hadrosauridae indet. Cope 1869		2 specimens
	or ? Ceratopsidae
Reptilia - Theropoda

	Theropoda indet. Marsh 1881	Nessov 1995	3 specimens
Reptilia

	Paronychodon asiaticus (Nessov 1995)	Sues and Averianov 2013	3 specimens
	synonym of Paronychodon
	ZIN PH 1054/16, 1059/16, 1068/16
Reptilia - Dromaeosauridae

	Richardoestesia asiatica (Nessov 1995)	Sues and Averianov 2013	4 specimens
	original and current combination Asiamericana asiatica
	ZIN PH 1101/16, 1119/16, 1121/16, 1138/16

	Itemirus medullaris Kurzanov 1976	Sues and Averianov 2014	1 specimen
	ZIN PH 92/16
Reptilia - Caenagnathidae

	Caenagnathasia martinsoni n. gen., n. sp. Currie et al. 1994	Currie et al. 1994	7 specimens
	Holotype. CCMGE 401/12457, fused dentaries; referred material: CCMGE 402/12457, a partial right dentary; ZIN PO 4603, and ZIN PO 5234, cervical vertebrae; ZIN PH 935/16, ZIN PH 936/16, and ZIN PH 937/16, dorsal vertebrae
Reptilia

	Tyrannosauroidea indet. Walker 1964	Averianov and Sues 2012	1 specimen
	ZIN PH 2296/16

	? Alectrosaurus sp. Gilmore 1933	Nessov 1995	1 specimen
Reptilia - Ornithomimidae

	Ornithomimidae indet. Marsh 1890	Sues and Averianov 2016	1 specimen
	ZIN PH 435/16
Reptilia

	Segnosauria indet. Barsbold and Perle 1980	Nessov 1995	3 specimens
	synonym of Therizinosauroidea
Amphibia - Temnospondyli - Discoglossidae

	Aralobatrachus robustus n. gen., n. sp. Nessov 1981		2 specimens
	ZIN, PHA No 477-7 (holotype partial maxilla), LU-N4/54 (maxilla)
Amphibia - Temnospondyli - Urodela

	Mynbulakia surgayi Nessov 1981	Nessov 1995	1 specimen
	synonym of Eoscapherpeton asiaticum
Actinopteri - Amiiformes - Amiidae

	Amia limosa Nessov 1985	Nessov 1985
	nomen dubium belonging to Amiinae
Actinopteri - Lepisosteiformes - Lepisosteidae

	Lepisosteidae indet. Cuvier 1825	Nessov 1995	1 specimen
Actinopteri - Ichthyodectiformes

	? Aidachar sp. Nessov 1981	Nessov 1995	1 specimen
Chondrichthyes - Rajiformes - Ptychotrygonidae

	Ptychotrygon sp. Jaekel 1894	Nessov 1985	1 specimen
Chondrichthyes - Rajiformes - Rhinobatidae

	Myledaphus tritus	Nessov 1985	2 specimens
see common names

Geography

Country:	Uzbekistan	State/province:	Navoi
Coordinates:	42.1° North, 62.7° East (view map)
Paleocoordinates:	36.8° North, 57.0° East
Basis of coordinate:	based on nearby landmark
Geographic resolution:	small collection

Time

Period:	Cretaceous	Epoch:	Late/Upper Cretaceous
Stage:	Turonian	10 m.y. bin:	Cretaceous 6
Key time interval:	Middle Turonian - Late/Upper Turonian
Age range of interval:	93.90000 - 89.80000 m.y. ago

Stratigraphy

Formation:	Bissekty
Local section:	Dzharakuduk	Local bed:	26 m
Local order:	bottom to top
Stratigraphic resolution:	bed
Stratigraphy comments: lower part of formation, 26 m above base Also described as Taykarshinskaya beds.

Lithology and environment

Primary lithology:	not reported
Environment:	terrestrial indet.

Taphonomy

Modes of preservation:	body
Size of fossils:	macrofossils
Preservation of anatomical detail:	good
Associated major elements:	all
Fragmentation:	frequent

Collection methods and comments

Collection methods:	selective quarrying,mechanical,field collection
Reason for describing collection:	taxonomic analysis
Collectors:	L. A. Nessov; A. Averianov, H.-D. Sues	Collection dates:	1977, 1991
Taxonomic list comments:Nessov (1984) listed turtles from "Dzharakuduk (= Kulbecke, =Bissekty) locality, central Kizylkum Desert and unamed beds adajcent to Tjulkili Top", but only gave details for CDZh-17a

Metadata

Also known as:	Dzarakuduk, Dzhyrakuduk
Database number:	46959
Authorizer:	M. Carrano, R. Benson	Enterer:	M. Carrano, K. Maguire, R. Benson
Modifier:	M. Carrano	Research group:	vertebrate
Created:	2005-01-27 11:26:13	Last modified:	2022-01-11 12:56:04
Access level:	the public	Released:	2005-01-27 11:26:13
Creative Commons license:	CC BY

Reference information

Primary reference:

64195.

L. A. Nessov. 1981. Amfibii i reptilii v ekosistemakh Mela sredney Azii [Amphibia and reptiles in Cretaceous ecosystems of central Asia]. In I. V. Darevsky (ed.), The Problems of Herpetology. Fifth Herpetological Conference. Abstracts 91-92 [M. Carrano/M. Carrano/M. Carrano]

Secondary references:

39419		J. D. Archibald and A. O. Averianov. 2003. The Late Cretaceous placental mammal Kulbeckia. Journal of Vertebrate Paleontology 23(2):404-419 [R. Benson/R. Benson]
17405	ETE	J. D. Archibald and A. O. Averianov. 2006. Late Cretaceous asioryctitherian eutherian mammals from Uzbekistan and phylogenetic analysis of Asioryctitheria. Acta Palaeontologica Polonica 51(2):351-376 [M. Carrano/M. Carrano]
71016		A. O. Averianov and H.-D. Sues. 2012. Skeletal remains of Tyrannosauroidea (Dinosauria: Theropoda) from the Bissekty Formation (Upper Cretaceous: Turonian) of Uzbekistan. Cretaceous Research 34:284-297 [M. Carrano/M. Carrano]
18347	ETE	P. J. Currie. 2000. Theropods from the Cretaceous of Mongolia. In M. J. Benton, M. A. Shishkin, D. M. Unwin, & E N. Kurichkin (eds.), The Age of Dinosaurs in Russia and Mongolia 434-455 [M. Carrano/M. Carrano/M. Carrano]
9243		P. J. Currie, S. J. Godfrey, and L. Nessov. 1994. New caenagnathid (Dinosauria, Theropoda) specimens from the Upper Cretaceous of North America and Asia. Canadian Journal of Earth Sciences 30(10-11):2255-2272 [M. Carrano/M. Carrano/M. Carrano]
41746		I. G. Danilov and J. F. Parham. 2005. A reassessment of the referral of an isolated skull from the Late Cretaceous of Uzbekistan to the stem-testudinoid turtle genus Lindholmemys. Journal of Vertebrate Paleontology 25(4):784-791 [R. Benson/R. Benson/P. Holroyd]
38127		L. A. Nessov. 1982. Drevneishie mlekopitaiushchie SSSR [Ancient mammals of the USSR]. Palyentologicheskogo Obshchyestva 25:228-243 [M. Carrano/M. Carrano]
41769		L. A. Nessov. 1984. Data on late Mesozoic turtles from the USSR. Studia Geologica Salamanticensia, vol. especial 1 (Studia Palaeocheloniologica I) 1:215-223 [R. Benson/R. Benson]
38131		L. A. Nessov. 1985. Redkie kostnye ryby, nazemiye yashcheritsy i mlekopitayushchie zony limanov i primorskikh nizmennostey mela Kyzylkumov [Rare bony fishes, terrestrial lizards, and mammals in the estuarine and coastal lowland zone of the Kyzyl-Kum Cretaceous]. Yearbook of the All-Union Paleontological Association 28:199-219 [M. Carrano/M. Carrano]
38153		L. A. Nessov. 1985. Novye mlekopitayushchie Mela Kyzylkumov [New mammals from the Cretaceous of Kyzyl-Kum]. Vestnik Leningradskogo Universiteta, Geologiia, Geografiia, series 7 17:8-18 [M. Carrano/M. Carrano]
16510	ETE	L. A. Nessov. 1995. Dinozavri severnoi Yevrazii: Novye dannye o sostave kompleksov, ekologii i paleobiogeografii [Dinosaurs of northern Eurasia: new data about assemblages, ecology, and paleobiogeography]. Institute for Scientific Research on the Earth's Crust, St. Petersburg State University, St. Petersburg 1-156 [M. Carrano/K. Maguire/M. Carrano]
39261		L. A. Nessov. 1997. In L. B. Goloneva & A. O. Averianov (ed.), Nemorskiye Pozvonochnyye Melovogo Perioda Severnoy Yevrazii [Cretaceous Non-Marine Vertebrates of Northern Eurasia] 1-218 [R. Benson/R. Benson/M. Carrano]
2649		L. A. Nessov, J. D. Archibald, and Z. Kielan-Jaworowska. 1998. Ungulate-like mammals from the Late Cretaceous of Uzbekistan and a phylogenetic analysis of Ungulatomorpha. Bulletin of Carnegie Museum of Natural History 34:40-88 [J. Alroy/J. Alroy/R. Benson]
39415		L. A. Nessov, D. Sigogneau-Russell, and D. E. Russell. 1994. A survey of Cretaceous tribosphenic mammals from middle Asia (Uzbekistan, Kazakhstan and Tajikistan), of their geological setting, age and faunal environment. Palaeovertebrata 23:51-92 [R. Benson/R. Benson]
18862		Z. Ročec and L. A. Nessov. 1993. Cretaceous anurans from Central Asia. Palaeontographica Abteilung A 226:1-54 [E. Fara/E. Fara/M. Uhen]
61991		M. J. Ryan. 1997. Middle Asian dinosaurs. In P. J. Currie & K. Padian (ed.), Encyclopedia of Dinosaurs 442-444 [M. Carrano/M. Carrano]
69215		M. A. Shishkin. 2000. Mesozoic amphibians from Mongolia and the Central Asiatic republics. In M. J. Benton, M. A. Shishkin, D. M. Unwin, & E N. Kurichkin (eds.), The Age of Dinosaurs in Russia and Mongolia 297-308 [M. Carrano/M. Carrano]
30213	ETE	H.-D. Sues and A. Averianov. 2009. A new basal hadrosauroid dinosaur from the Late Cretaceous of Uzbekistan and the early radiation of duck-billed dinosaurs. Proceedings of the Royal Society B: Biological Sciences 276(1667):2549-2555 [M. Carrano/M. Carrano]
46400		H.-D. Sues and A. Averianov. 2013. Enigmatic teeth of small theropod dinosaurs from the Upper Cretaceous (Cenomanian–Turonian) of Uzbekistan. Canadian Journal of Earth Sciences 50(3):306-314 [P. Mannion/J. Tennant/M. Carrano]
53745		H.-D. Sues and A. Averianov. 2014. Dromaeosauridae (Dinosauria: Theropoda) from the Bissekty Formation (Upper Cretaceous: Turonian) of Uzbekistan and the phylogenetic position of Itemirus medullaris Kurzanov, 1976. Cretaceous Research 51:225-240 [M. Carrano/M. Carrano]
53929		H.-D. Sues and A. O. Averianov. 2015. New material of Caenagnathasia martinsoni (Dinosauria: Theropoda: Oviraptorosauria) from the Bissekty Formation (Upper Cretaceous: Turonian) of Uzbekistan. Cretaceous Research 54:50-59 [P. Mannion/J. Tennant/M. Carrano]
79047		H.-D. Sues and A. Averianov. 2016. Ornithomimidae (Dinosauria: Theropoda) from the Bissekty Formation (Upper Cretaceous: Turonian) of Uzbekistan. Cretaceous Research 57:90-110 [M. Carrano/M. Carrano]
41695		V. B. Sukhanov. 2000. Mesozoic turtles of Middle and Central Asia. In M. J. Benton, M. A. Shishkin, D. M. Unwin, & E N. Kurichkin (eds.), The Age of Dinosaurs in Russia and Mongolia 309-367 [R. Benson/R. Benson/M. Carrano]