Hambach mine horizon 6C: MN 5, Germany
List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Actinopteri
- Sciaenidae
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Atractoscion sp.
Gill 1862
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Actinopteri
- Spariformes
- Sparidae
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Sparidae indet.
(Cuvier 1817)
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Actinopteri
- Cypriniformes
- Cyprinidae
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Barbus sp.
Cuvier and Cloquet 1816
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Palaeotinca sp.
Obrhelová 1969
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Palaeoleuciscus sp.
Obrhelová 1969
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Amphibia
- Temnospondyli
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Latonia seyfriedi
von Meyer 1843
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Amphibia
- Temnospondyli
- Pelobatidae
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Pelobates sp.
Wagler 1830
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vel Eopelobates sp. | ||||||||||
Amphibia
- Temnospondyli
- Urodela
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Salamandridae indet.
Gray 1825
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various kinds | ||||||||||
Chelotriton paradoxus
Pomel 1853
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Proteidae indet.
Hogg 1838
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Euronecturus grogu n. sp.
Macaluso et al. 2021
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Macaluso et al. 2021 | |||||||||
Holotype: IPB-HaH 2121, an almost complete atlas; IPB-HaH 2120, atlas; IPB-HaH 2150, atlas; IPB-HaH 2119, atlas; IPB-HaH 2165, atlas | ||||||||||
Andrias scheuchzeri
Tschudi 1837
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cf. Salamandra sp.
Laurenti 1768
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Reptilia
- Chamaeleonidae
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Chameleontidae indet.
Gray 1825
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"Chamaeleontidae" | ||||||||||
Chamaeleonidae informal tooth morphotype 1
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Čerňanský et al. 2017 | 2 specimens | ||||||||
IPB-HaH 3901, right maxilla; IPB-HaH 3902, left maxilla | ||||||||||
Chamaeleonidae informal tooth morphotype 2
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Čerňanský et al. 2017 | 1 specimen | ||||||||
IPB-HaH 3903, right maxilla | ||||||||||
Chamaeleo aff. andrusovi
Čerňanský 2010
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Čerňanský et al. 2017 | 1 specimen | ||||||||
IPB-HaH 3900, right squamosal | ||||||||||
Reptilia
- Serpentes
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Serpentes indet.
Linnaeus 1758
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various kinds | ||||||||||
Reptilia
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Colubroidea indet.
Oppel 1811
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Čerňanský et al. 2017 | 6 specimens | ||||||||
IPB-HaH 3829; 3836; 3840; 3848; 3851; 3856, thirty-six caudal vertebrae | ||||||||||
Reptilia
- Elapidae
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cf. Naja sp.
Laurenti 1768
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Čerňanský et al. 2017 | 4 specimens | ||||||||
IPB-HaH 3833; 3845; 3850; 3859, six trunk vertebrae | ||||||||||
Reptilia
- Crotalidae
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Vipera informal "oriental viper" sp.
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Čerňanský et al. 2017 | 6 specimens | ||||||||
IPB-HaH 3846; 3809; 3811; 3812; 3819; 3860, eight trunk vertebrae | ||||||||||
Vipera informal "aspis complex" sp.
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Čerňanský et al. 2017 | 2 specimens | ||||||||
IPB-HaH 3839; 3847, three trunk vertebrae | ||||||||||
Reptilia
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Eoanilius sp.
Rage 1974
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Čerňanský et al. 2017 | 3 specimens | ||||||||
IPB-HaH 3801, 3802 and 3824, five trunk vertebrae (remainder prefixed by latter number) | ||||||||||
Reptilia
- Boidae
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Bavarioboa sp.
Szyndlar and Schleich 1993
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Čerňanský et al. 2017 | 5 specimens | ||||||||
IPB-HaH 3825, 3807, 3013, 3814, 3818, five trunk vertebrae | ||||||||||
cf. Falseryx sp.
Szyndlar and Rage 2003
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Čerňanský et al. 2017 | 1 specimen | ||||||||
IPB-HaH 3830, one trunk vertebra | ||||||||||
Reptilia
- Colubridae
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Natrix informal sp. 1
Laurenti 1768
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Čerňanský et al. 2017 | 2 specimens | ||||||||
IPB-HaH 3842 and 3843, two trunk vertebrae | ||||||||||
"Colubrinae" informal indet. 1
Oppel 1811
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Čerňanský et al. 2017 | 7 specimens | ||||||||
IPB-HaH 3828; 3835; 3838; 3844; 3805; 3806; 3855, fifty seven trunk vertebrae | ||||||||||
Texasophis sp.
Holman 1977
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Čerňanský et al. 2017 | 2 specimens | ||||||||
IPB-HaH 3832, 3841, two trunk vertebrae | ||||||||||
Telescopus sp.
Waner 1830
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Čerňanský et al. 2017 | 1 specimen | ||||||||
IPB-HaH 3853, one trunk vertebra | ||||||||||
"Coluber" sp.
Linnaeus 1758
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Čerňanský et al. 2017 | 14 specimens | ||||||||
IPB-HaH 3800; 3826; 3831; 3803; 3804; 3852; 3808; 3857; 3815; 3816; 3817; 3820; 3822; 3823, forty two trunk vertebrae | ||||||||||
"Natricinae" indet.
Bonaparte 1838
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Čerňanský et al. 2017 | 6 specimens | ||||||||
IPB-HaH 3827; 3834; 3837; 3849; 3854; 3821, nine trunk vertebrae | ||||||||||
Reptilia
- Lacertidae
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Lacertidae informal indet. 1
Bonaparte 1831
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Čerňanský et al. 2017 | 3 specimens | ||||||||
IPB-HaH 3701, left maxilla; IPB-HaH 3702 and 3703, two left dentaries | ||||||||||
Lacertidae informal indet. 2
Bonaparte 1831
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Čerňanský et al. 2017 | 2 specimens | ||||||||
IPB-HaH 3704, premaxilla; IPB-HaH 3705, left dentary | ||||||||||
Lacerta sp.
Linnaeus 1758
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Reptilia
- Anguidae
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Anguidae informal indet. 1
Gray 1825
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Čerňanský et al. 2017 | 2 specimens | ||||||||
IPB-HaH 3700, twenty osteoderms; IPB-HaH 3709, caudal vertebra | ||||||||||
Ophisaurus sp.
Daudin 1803
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Pseudopus informal sp. 1
Merrem 1820
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Čerňanský et al. 2017 | 1 specimen | ||||||||
IPB-HaH 3708, dorsal vertebra | ||||||||||
Pseudopus cf. ahnikoviensis
Klembara 2012
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Čerňanský et al. 2017 | 2 specimens | ||||||||
IPB-HaH 3706, right dentary; IPB-HaH 3707, left dentary | ||||||||||
Reptilia
- Testudines
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Trionychidae indet.
Gray 1825
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Klein and Mors 2003 | |||||||||
= Carettochelyinae indet.
Boulenger 1887
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Joyce et al. 2004 | |||||||||
Reptilia
- Testudines
- Emydidae
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Emydidae indet.
Rafinesque 1815
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2 types | ||||||||||
Reptilia
- Testudines
- Geoemydidae
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Clemmydopsis cf. turnauensis
(Meyer 1847)
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Klein and Mors 2003 | |||||||||
"Ocadia" sophiae
(Ammon 1911)
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Klein and Mors 2003 | |||||||||
recombined as Mauremys sophiae | ||||||||||
Reptilia
- Testudines
- Testudinidae
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Testudo sp.
Linnaeus 1758
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Klein and Mors 2003 | |||||||||
vel Geochelone sp. | ||||||||||
Reptilia
- Testudines
- Trionychidae
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Trionyx sp.
Geoffroy Saint-Hilaire 1809
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listed as "Trionyx s.l." | ||||||||||
Reptilia
- Testudines
- Chelydridae
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Chelydropsis sp.
Peters 1868
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= Chelydropsis murchisoni
Bell 1836
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Klein and Mors 2003 | |||||||||
Reptilia
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Diplocynodon sp.
Pomel 1847
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Aves indet.
Linnaeus 1758
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Mammalia
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Amphiperatherium frequens
(von Meyer 1846)
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Mammalia
- Proboscidea
- Proboscidea
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Proboscidea indet.
Illiger 1811
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"Elephantoidea" | ||||||||||
Mammalia
- Rodentia
- Castoridae
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Anchitheriomys sp.
Roger 1885
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Anchitheriomys suevicus
Schlosser 1884
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Steneofiber depereti
Mayet 1908
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Mammalia
- Rodentia
- Eomyidae
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Eomyidae indet.
Winge 1887
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Eomyops cf. catalaunicus
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Keramidomys carpathicus
(Schaub and Zapfe 1953)
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Mammalia
- Rodentia
- Anomalomyidae
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Anomalomys minor
Fejfar 1972
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Mammalia
- Rodentia
- Sciuridae
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Heteroxerus sp.
Stehlin and Schaub 1951
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Hylopetes sp.
Thomas 1908
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Miopetaurista lappi
Mein 1958
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Blackia miocaenica
Mein 1970
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Spermophilinus aff. bredai
(von Meyer 1848)
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? Ratufa sp.
Gray 1867
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Palaeosciurus sutteri
Ziegler and Fahlbusch 1986
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Mammalia
- Rodentia
- Cricetidae
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Karydomys wigharti n. sp.
Mörs and Kalthoff 2004
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Mörs and Kalthoff 2004 | |||||||||
Lartetomys cf. zapfei
Mein and Freudenthal 1971
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Eumyarion cf. bifidus
(Fahlbusch 1964)
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Eumyarion cf. weinfurteri
Schaub and Zapfe 1953
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Democricetodon mutilus
(Fahlbusch 1964)
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Megacricetodon collongensis
Mein 1958
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Fahlbuschia koenigswaldi
Freudenthal 1963
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Mammalia
- Rodentia
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Neocometes cf. similis
Fahlbusch 1966
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Mammalia
- Rodentia
- Gliridae
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Miodyromys aegercii
Baudelot 1972
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Bransatoglis cf. astaracensis
(Baudelot 1970)
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Glirudinus sp.
de Bruijn 1966
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Glirudinus undosus
Mayr 1979
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Eomuscardinus sp.
Hartenberger 1966
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synonym of Muscardinus | ||||||||||
Eomuscardinus sansaniensis
Lartet 1851
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Myoglis meini
de Bruijn 1966
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Paraglirulus werenfelsi
Engesser 1972
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Paraglirulus cf. agelakisi
Van der Meulen and De Bruijn 1982
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Microdyromys cf. legidensis
Daams 1981
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Mammalia
- Lagomorpha
- Ochotonidae
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Ochotonidae indet.
Thomas 1897
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Mammalia
- Primates
- Pliopithecidae
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Pliopithecidae indet.
Zapfe 1961
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Pliopithecus antiquus
(Blainville 1839)
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Mammalia
- Carnivora
- Hyaenidae
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? Protictitherium sp.
Kretzoi 1938
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Mammalia
- Carnivora
- Ursidae
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Ursavus elmensis
Stehlin 1917
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recombined as Ballusia elmensis | ||||||||||
Mammalia
- Carnivora
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Potamotherium miocenicum
Peters 1868
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Mammalia
- Carnivora
- Ailuridae
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Amphictis sp.
Pomel 1853
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Mammalia
- Carnivora
- Mustelidae
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Mustelidae indet.
Fischer 1817
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Trocharion albanense
Forsyth-Major 1903
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Mammalia
- Carnivora
- Amphicyonidae
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Amphicyonidae indet.
Haeckel 1886
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Pseudocyon steinheimensis
Fraas 1885
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Mammalia
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Taucanamo sansaniense
Lartet 1851
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Mammalia
- Suidae
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Hyotherium soemmeringi
von Meyer 1834
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Mammalia
- Cetacea
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Cetacea indet.
Brisson 1762
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cf. Acrodelphis sp.
Abel 1900
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= Odontoceti indet.
Flower 1867
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Fordyce and de Muizon 2001 | |||||||||
Mammalia
- Bovidae
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Orygotherium escheri
von Meyer 1838
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Mammalia
- Lagomerycidae
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Dicrocerus elegans
Lartet 1837
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Lagomeryx ruetimeyeri
Thenius 1949
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Mammalia
- Tragulidae
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Dorcatherium naui
Kaup 1833
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Dorcatherium guntianum
von Meyer 1846
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Mammalia
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Amphimoschus pontileviensis
Bourgeois 1873
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Mammalia
- Perissodactyla
- Rhinocerotidae
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cf. Prosantorhinus sp.
Heissig 1973
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Brachypotherium brachypus
Lartet 1848
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Mammalia
- Perissodactyla
- Equidae
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Anchitherium aurelianense
Cuvier 1825
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Mammalia
- Soricidae
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Soricidae indet.
Fischer 1814
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Dinosorex aff. zapfei
Engesser 1975
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Heterosorex sp.
Gaillard 1915
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Allosorex gracilidens
(Viret and Zapfe 1951)
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original and current combination Sorex gracilidens | ||||||||||
Mammalia
- Talpidae
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Talpini indet.
Fischer de Waldheim 1817
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spelled with current rank as Talpidae | ||||||||||
Talpidae indet.
(Fischer de Waldheim 1817)
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Ziegler and Mors 2000 | |||||||||
Desmaninae indet.
Thomas 1912
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Mygalea cf. antiqua
Pomel 1848
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Proscapanus sansaniensis
Lartet 1851
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Mammalia
- Plesiosoricidae
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Plesiosorex germanicus
(Seemann 1938)
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Mammalia
- Dimylidae
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Chainodus aff. intercedens
(Müller 1967)
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Plesiodimylus chantrei
Gaillard 1899
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Mammalia
- Erinaceidae
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Lanthanotherium aff. sansaniense
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original and current combination Lantanotherium | ||||||||||
"Mioechinus" sansaniensis
Lartet 1851
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Mammalia
- Chiroptera
- Rhinolophidae
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Rhinolophus cf. delphinensis
Gaillard 1899
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Mammalia
- Chiroptera
- Vespertilionidae
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Eptesicus cf. campanensis
Baudelot 1970
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Chondrichthyes
- Myliobatiformes
- Dasyatidae
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Dasyatidae indet.
Jordan 1888
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vel Myliobatidae | ||||||||||
Chondrichthyes
- Carcharhiniformes
- Scyliorhinidae
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Scyliorhinus distans
Probst 1879
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Chondrichthyes
- Carcharhiniformes
- Carcharhinidae
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Physogaleus sp.
Cappetta 1980
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Chondrichthyes
- Lamniformes
- Carchariidae
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Carcharias cf. reticulata
(Probst 1879)
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Odontaspis vorax
le Hon 1871
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synonym of Carcharias acutissimus | ||||||||||
see common names |
Geography
Country: | Germany |
Coordinates: | 50.9° North, 6.5° East (view map) |
Paleocoordinates: | 51.1° North, 5.0° East |
Basis of coordinate: | based on nearby landmark |
Geographic resolution: | outcrop |
Time
Period: | Neogene | Epoch: | Miocene |
Stage: | Langhian | 10 m.y. bin: | Cenozoic 5 |
Key time interval: | MN 5 | ||
Age range of interval: | 15.98000 - 13.82000 m.y. ago |
Stratigraphy
Formation: | Ville | Member: | Frimmersdorf | ||
Stratigraphic resolution: | bed | ||||
Stratigraphy comments: biozone MN 5, in the middle part of the main lignite seam, later authors list horizon as 6Ca/b
JA: "6C" entered as local bed, with no local section name Hambach 6C local fauna can be correlated with the upper part of mammalian Neogene unit MN 5, indicating an absolute age-range of 16.0–15.2 Ma for this late Orleanian, early middle Miocene fauna (Mörs et al. 2000; Mo€rs 2002; Mörs & Kalthoff 2004). |
Lithology and environment
Primary lithology: | lignite |
Secondary lithology: | lenticular,"cross stratification" sandy claystone |
Includes fossils? | Y |
Environment: | paralic indet. |
Geology comments: sediments are channel-fill, but environment interpreted as including brackish-estuarine, lacustrine-fluviatile and terrestrial environments
channel fill and floodplain deposits (horizon 6C according to the local lithostratigraphy) within the middle Miocene Frimmersdorf lignite seam (Mo€rs et al. 2000; Sch€afer et al. 2004). |
Taphonomy
Modes of preservation: | body |
Size of fossils: | macrofossils,mesofossils |
Disassociated major elements: | some |
Disassociated minor elements: | many |
Collection methods and comments
Collection methods: | field collection |
Reason for describing collection: | taxonomic analysis |
Collection method comments: this collection almost certainly comes from the same unit and same general area as Tiffney's collections Hambach 8251-13349
Macaluso et al. 2021: The new taxon is described based on material stored in the Steinmann Institute of the Rheinische Friedrich-Wilhelms-Universität Bonn, Germany. |
Metadata
Also known as: | Hambach 6C | ||
Database number: | 46955 | ||
Authorizer: | J. Alroy, M. Uhen, T. Cleary, P. Mannion | Enterer: | E. Leckey, G. Varnham, T. Cleary, M. Uhen |
Modifier: | G. Varnham | Research group: | vertebrate |
Created: | 2005-01-27 11:09:23 | Last modified: | 2022-03-28 07:01:55 |
Access level: | the public | Released: | 2005-01-27 11:09:23 |
Creative Commons license: | CC BY |
Reference information
Primary reference:
12472. | T. Mors, F. Hocht, and B. Wutzler. 2000. Die erst Wirbeltierfauna aus der miozanen Braunkohle der Niederrheinischen Bucht (Ville-Schichten, Tagebau Hambach) [The first vertebrate fauna from the Miocene Ville Series of the Lower Rhine Embayment (Hambach open cast mine, western Germany)]. Paläontologische Zeitschrift 74(1/2):145-170 [J. Alroy/E. Leckey/J. Alroy] |
Secondary references:
67910 | A. Čerňanský, Z. Szyndlar, and T. Mörs. 2017. Fossil squamate faunas from the Neogene of Hambach (northwestern Germany). Palaeobiodiversity and Palaeoenvironments 97:329-354 [T. Cleary/T. Cleary] | |
10151 | R. E. Fordyce and C. de Muizon. 2001. Evolutionary history of the cetaceans: a review. Secondary Adaptations of Tetrapods to Life in the Water 169-233 [M. Uhen/M. Uhen/M. Uhen] | |
12480 | W. G. Joyce, N. Klein, and T. Mors. 2004. Carettochelyine turtle from the Neogene of Europe. Copeia 2004(2):406-411 [J. Alroy/E. Leckey/J. Alroy] | |
12479 | N. Klein and T. Mors. 2003. Die Schildkroten (Reptilia: Testudines) aus dem Mittel-Miozan von Hambach (Niederrheinische Bucht, NW-Deutschland). Palaeontographica Abteilung A 268(1-3):1-48 [J. Alroy/E. Leckey/E. Leckey] | |
81028 | L. Macaluso, A. Villa, and T. Mörs. 2021. A new proteid salamander (Urodela, Proteidae) from the Middle Miocene of Hambach (Germany) and implications for the evolution of the family. Palaeontology 65(1):e12585 [P. Mannion/G. Varnham] | |
70399 | T. Mors. 2002. Biostratigraphy and paleoecology of continental Tertiary vertebrate faunas in the Lower Rhine Embayment (NW-Germany). Netherlands Journal of Geosciences 81(2):177-183 [T. Cleary/T. Cleary] | |
12482 | T. Mörs and D. C. Kalthoff. 2004. A new species of Karydomys (Rodentia, Mammalia) and a systematic re-evaluation of this rare Eurasian Miocene hamster. Palaeontology 47(6):1387-1405 [J. Alroy/E. Leckey/L. van den Hoek Ostende] | |
12476 | R. Ziegler and T. Mors. 2000. Marsupialia, Lipotyphla und Chiroptera (Mammalia) aus dem Miozan des Braunkohlentagebaus Hambach (Niederrheinische Bucht, NW-Deutschland). Palaeontographica Abteilung A 257(1-3):1-26 [J. Alroy/E. Leckey/E. Leckey] |