PBDB Collection

Blasi 2, Arén: Late/Upper Maastrichtian, Spain

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

Amphibia

	Amphibia indet. Linnaeus 1758
Amphibia - Temnospondyli - Palaeobatrachidae

	Palaeobatrachidae indet. Cope 1865	Blain et al. 2010	36 specimens
	3 incomplete maxillae [MPZ 2009/300-302], 14 ilia (1 left [MPZ 2009/303], 2 right [MPZ 2009/304-305] and 11 indet. [MPZ 2009/306-316]), 1 right scapula [MPZ 2009/317], 1 vertebra [MPZ 2009/318] and 4 radioulnae [MPZ 2009/319-322]
Amphibia - Temnospondyli

	aff. Paradiscoglossus sp. Estes and Sanchíz 1982	Blain et al. 2010	82 specimens
	1 mandible [MPZ 2009/229], 13 ilia (2 left [MPZ 2009/ 230-231], 7 right [MPZ 2009/232-238] and 4 indet. fragments [MPZ 2009/239-242]), 9 humeri [MPZ 2009/243-251], 1 urostyle [MPZ 2009/252], 4 vertebrae [MPZ 2009/253-256], 2 sacral vertebrae [MPZ 2009/257-258], 4 radioulnae [MPZ 2009/259-262], 1 femur [MPZ 2009/263] and 36 tibiofibulae [MPZ 2009/264-299]
Amphibia - Temnospondyli - Albanerpetontidae

	Albanerpeton aff. nexuosum (Estes 1981)	Blain et al. 2010	52 specimens
	original and current combination Albanerpeton nexuosus
	3 premaxillae [MPZ 2009/179-181], 4 maxillae (2 left [MPZ 2009/182-183] and 2 right [MPZ 2009/184-185]); 25 dentaries (14 left [MPZ 2009/186-199], 9 right [MPZ 2009/200-208] and 2 indet. [MPZ 2009/207-208]), 8 indeterminate partial jaws [MPZ 2009/211-218], 1 frontal [MPZ 2009/219], 5 humeri [MPZ 2009/221-225] and 2 trunk vertebrae [MPZ 2009/226-227]: misspelled "nexuosum"
Reptilia - Spheroolithidae

	Spheroolithus aff. europaeus Sellés et al. 2014	Pérez-Pueyo et al. 2021
Reptilia - Hadrosauridae

	Euhadrosauria indet. Weishampel et al. 1993	López-Martínez et al. 2001	4 specimens
	BLA 2b/2
Reptilia

	Theropoda indet. Marsh 1881		1 specimen
	MPZ2004/5, tooth
	= Neoceratosauria indet. Novas 1991	Canudo et al. 2005
	= Theropoda informal indet. 1 Marsh 1881	Torices et al. 2015

	Coelurosauria indet. Huene 1914	López-Martínez et al. 2001	50 specimens
	= Maniraptoriformes indet. Holtz and Padian 1995	Canudo et al. 2005
	MPZ 98/79, 80, 81, 82; 50 isolated teeth

	cf. Euronychodon sp. Antunes and Sigogneau-Russell 1991	López-Martínez et al. 2001	3 specimens
	MPZ 98/76, 77, 78, teeth
	= ? Paronychodon sp. Cope 1876	Torices et al. 2015
Reptilia - Dromaeosauridae

	? Dromaeosaurinae indet. Matthew and Brown 1922	López-Martínez et al. 2001	1 specimen
	"Dromaeosauridae type C"; MPZ 2004/6
	= cf. Dromaeosauridae informal indet. 6 Colbert and Russell 1969	Torices et al. 2004
	= ? Dromaeosauridae indet. Colbert and Russell 1969	Torices et al. 2015

	cf. Dromaeosauridae informal indet. 8 Colbert and Russell 1969	Torices et al. 2004	1 specimen
	MPZ 98/75
	= Dromaeosauridae informal indet. E Colbert and Russell 1969	Canudo et al. 2005

	cf. Dromaeosauridae informal indet. 4 Colbert and Russell 1969	Torices et al. 2004	2 specimens
	MPZ 98/72, 74, teeth
	= Dromaeosauridae informal indet. A Colbert and Russell 1969	Canudo et al. 2005
	= ? Richardoestesia sp. Currie et al. 1990	Torices et al. 2015

	cf. Dromaeosauridae informal indet. 7 Colbert and Russell 1969	Torices et al. 2004	2 specimens
	MPZ 98/73, 2004/7, teeth
	= Dromaeosauridae informal indet. D Colbert and Russell 1969	Canudo et al. 2005
	= ? Richardoestesia sp. Currie et al. 1990	Torices et al. 2015
Reptilia - Prismatoolithidae

	Prismatoolithidae indet. Hirsch 1994	López-Martínez 2000
Reptilia

	Sauropoda indet. Marsh 1878	López-Martínez et al. 2001
Reptilia - Crocodyliformes

	"Trematochampsidae" indet. Buffetaut 1974	López-Martínez et al. 2001
	= Crocodyliformes indet. Hay 1930	Puértolas-Pascual et al. 2016
Reptilia - Krokolithidae

	Megaloolithidae indet. Zhao 1979	López-Martínez 2000
	dinosauroid tubospherulithic
	= Krokolithes sp. Hirsch 1985	Pérez-Pueyo et al. 2021
Reptilia

	Alligatoroidea indet. Gray 1844	López-Martínez et al. 2001	2 specimens
	= Eusuchia indet. Huxley 1875	Puértolas-Pascual et al. 2016

	Crocodylia indet. (Owen 1842)	López-Martínez et al. 2001
	= Arenysuchus gascabadiolorum Puértolas et al. 2011	Puértolas et al. 2011
	MPZ 2010/948-951; 4 teeth

	Acynodon sp. Buscalioni et al. 1997	López-Martínez et al. 2001	1 specimen
	MPZ-2017/1137, tooth
	= cf. Acynodon sp. Buscalioni et al. 1997	Pérez-Pueyo et al. 2021
Reptilia - Squamata

	Iguanidae indet. Oppel 1811	López-Martínez et al. 2001	3 specimens
	= Squamata indet. Oppel 1811	Blain et al. 2010
	Indeterminate lizard 1 (probably an iguanid, teiid or lacertid) - 1 partial unidentified bone with 3 teeth [MPZ 2009/323]
Reptilia

	Alethinophidia indet. (Nopcsa 1923)	Blain et al. 2010	1 specimen
	posterior caudal vertebra [MPZ 2009/331]

	Lacertilia indet. Owen 1842	López-Martínez et al. 2001	2 specimens
	= Scleroglossa indet. Estes et al. 1988	Blain et al. 2010
	Indeterminate lizard 2 - 1 left dentary [MPZ 2009/324] and 1 incomplete unidentified bone with 2 teeth [MPZ 2009/325]
Reptilia - Anguidae

	Anguidae indet. Gray 1825	Blain et al. 2010	5 specimens
	5 trunk vertebrae [MPZ 2009/326-330]
Reptilia - Testudines - Bothremydidae

	Bothremydidae indet. Baur 1891	Murelaga and Canudo 2005
Reptilia - Testudines

	cf. Polysternon sp. Portis 1882	López-Martínez et al. 2001
Reptilia

	Solemydidae indet. Lapparent de Broin and Murelaga 1996	Murelaga and Canudo 2005
	= Solemys sp. Lapparent de Broin and Murelaga 1996	Joyce 2017
Actinopteri

	Teleostei indet. Müller 1846	López-Martínez et al. 2001
Actinopteri - Lepisosteiformes - Lepisosteidae

	Lepisosteidae indet. Cuvier 1825	López-Martínez et al. 2001
Actinopteri - Pycnodontiformes

	Pycnodontiformes indet. Berg 1937	López-Martínez et al. 2001
Charales - Feistiellaceae

	Feistiella sp. Schudack 1986	López-Martínez et al. 2001
Charales - Characea

	Amblyochara concava	López-Martínez et al. 2001
	A. c. concava, A. c. mucronata, A. c. ssp.

	Amblyochara informal sp. A Grambast 1962	López-Martínez et al. 2001

	Peckichara sertulata	López-Martínez et al. 2001
see common names

Geography

Country:	Spain	State/province:	Aragón	County:	Huesca
Coordinates:	42.3° North, 0.7° East (view map)
Paleocoordinates:	34.3° North, 0.6° East
Basis of coordinate:	based on nearby landmark
Geographic resolution:	outcrop

Time

Period:	Cretaceous	Epoch:	Late/Upper Cretaceous
Stage:	Maastrichtian	10 m.y. bin:	Cretaceous 8
Key time interval:	Late/Upper Maastrichtian	Foram zone:	Abathomphalus mayaroensis
Age range of interval:	72.10000 - 66.00000 m.y. ago

Stratigraphy

Geological group:	Tremp	Formation:	La Posa
Local section:	Areny	Local bed:	8 m
Local order:	bottom to top
Stratigraphic resolution:	bed
Stratigraphy comments: lower part; 6.5 m interval; upper part of chron c30n, 65.6-68.7 mya The Conques Formation is equivalent to the "lower Red Garumen" facies. The Conques Formation was previously assigned a late maastrichtian age (Sellés and Villa 2015; Villa et al. 2015) Biostratigraphy and Sr stable isotope geochemistry suggest that the La Posa and Conques Formation are entirely early Maastrichtian, lying within chron C31r (see Caus et al. 2016; Oms et al. 2016; Fondevilla et al. 2016; Fondevilla et al. 2019). Actually the "Grey Garumnian" = La Posa Fm.

Lithology and environment

Primary lithology:	lenticular,gray marl
Includes fossils?	Y
Lithology description: "grey marls", 2 m thick
Environment:	lagoonal/restricted shallow subtidal
Geology comments: "lagoonal/marsh marls"

Taphonomy

Modes of preservation:	body
Degree of concentration:	concentrated
Size of fossils:	macrofossils,mesofossils
Preservation of anatomical detail:	medium
Disassociated major elements:	many
Disassociated minor elements:	many
Fragmentation:	frequent

Collection methods and comments

Collection methods:	bulk,sieve,field collection
Rock censused:	5000 kg
Reason for describing collection:	biostratigraphic analysis
Collection method comments: includes seven types of eggshell

Metadata

Also known as:	Blasi 2a, 2b
Database number:	45437
Authorizer:	M. Carrano, P. Holroyd, P. Mannion	Enterer:	M. Carrano, P. Mannion, J. Tennant, P. Holroyd
Modifier:	M. Carrano	Research group:	vertebrate
Created:	2004-10-22 13:35:56	Last modified:	2023-01-10 15:34:34
Access level:	the public	Released:	2004-10-22 13:35:56
Creative Commons license:	CC BY

Reference information

Primary reference:

17660.

ETE

J. I. Canudo, L. Ardévol, G. Cuenca-Bescós, N. López-Martínez, X. Murelaga, X. Pereda-Suberbiola, J. I. Ruiz-Omeñaca and X. Orue-Etxebarria. 1999. New dinosaur localities near the Cretaceous/Tertiary boundary from Arén (Huesca, Spain). In J. I. Canudo & G. Cuenca-Bescós (ed.), IV European Workshop on Vertebrate Paleontology. Programme and Abstracts. Field Guide 40-41 [M. Carrano/M. Carrano/M. Carrano]

Secondary references:

37367		H.-A. Blain, J.-I. Canudo, G. Cuenca-Bescós and N. López-Martínez. 2010. Amphibians and squamate reptiles from the latest Maastrichtian (Upper Cretaceous) of Blasi 2 (Huesca, Spain). Cretaceous Research 31(4):433-446 [P. Mannion/P. Mannion]
14882	ETE	J. I. Canudo. 2004. El caso del dinosaurio hadrosaurio de Arén (Huesca): muerte accidental o asesinato? [The case of the hadrosaur dinosaur of Arén (Huesca): accidental death or assassination?]. Naturaleza Aragonesa 13:4-14 [M. Carrano/K. Maguire/M. Carrano]
53564		J. I. Canudo, J. L. Barco, R. Royo-Torres and J. I. Ruiz-Omeñaca. 2001. Los saurópodos (Dinosauria) del Tithónico (Jurásico Superior) y del Cretácico de Aragón [The sauropoda (Dinosauria) from the Tithonian (Upper Jurassic) and Cretaceous of Aragón]. I Jornadas Internacionales sobre Paleontología de Dinosaurios y Su Entorno Actas:309-318 [M. Carrano/M. Carrano/M. Carrano]
17600	ETE	J. I. Canudo, J. L. Barco, P. Cruzado-Caballero, G. Cuenca-Bescós, J. I. Ruiz-Omeñaca and R. Royo-Torres. 2005. Evidencias de predación de dinosaurios terópodos en el Maastrichtiense superior, Cretácico superior de Arén (Huesca) [Evidence of theropod dinosaur predation in the upper Maastrichtian, Upper Cretaceous of Arén (Huesca)]. Lucas Mallada 12:29-58 [M. Carrano/M. Carrano]
28738	ETE	J. I. Canudo, G. Cuenca-Bescós, and J. I. Ruiz-Omeñaca. 2004. Los últimos dinosaurios europeos: debates recientes sobre la extinción de los dinosaurios [The last European dinosaurs: recent debates on the extinction of the dinosaurs]. Historia Natural 7:42-51 [M. Carrano/M. Carrano]
17647	ETE	J. I. Canudo and J. I. Ruiz-Omeñaca. 2003. Los restos directos de dinosaurios teropódos (excluyendo Aves) en España [Direct remains of theropod dinosaurs (excluding Aves) in Spain]. Ciencias de la Tierra 26:347-373 [M. Carrano/M. Carrano]
81303		J. Company, P. Cruzado-Caballero, and J. I. Canudo. 2013. A diminutive hadrosaur from the Late Maastrichtian of the south-central Pyrenees (Spain). In F. Torcida Fernández-Baldor & P. Huerta (ed.), VI Jornadas Internacionales sobre Paleontología de Dinosaurios y su Entorno. Libro de Resúmenes 53-55 [M. Carrano/M. Carrano]
70541		J. Company, P. Cruzado-Caballero, and J. I. Canudo. 2015. Presence of diminutive hadrosaurids (Dinosauria: Ornithopoda) in the Maastrichtian of the south-central Pyrenees (Spain). Journal of Iberian Geology 41(1):71-81 [M. Carrano/M. Carrano]
42686		J. Company and Z. Szentesi. 2012. Amphibians from the Late Cretaceous Sierra Perenchiza Formation of the Chera Basin, Valencia Province, Spain. Cretaceous Research 37:240-245 [P. Mannion/P. Mannion/M. Carrano]
17658	ETE	P. Cruzado-Caballero and J. I. Canudo. 2006. Nuevas evidencias de ornitópodos "enanos" en el Maastrichtiense superior Europeo (Blasi 5, Arén, Huesca) [New evidences of "dwarf" ornithopods in the European upper Maastrichtian (Blasi 5, Arén, Huesca)]. IV Encuentro de Jóvenes Investigadores en Paleontología, Salamanca, Spain 75-76 [M. Carrano/M. Carrano]
28219	ETE	P. Cruzado-Caballero, J. I. Canudo, and J. I. Ruiz-Omeñaca. 2007. Los dinosaurios hadrosaurios (Ornithischia: Ornithopoda) del Maastrichtiense superior (Cretácico Superior) de Arén (Huesca, España) [The hadrosaurian dinosaurs (Ornithischia: Ornithopoda) from the upper Maastrichtian (Upper Cretaceous) of Arén (Huesca, Spain)]. IV Jornadas Internacionales sobre Paleontología de Dinosaurios y su Entorno, Salas de los Infantes, Spain 55-56 [M. Carrano/M. Carrano/M. Carrano]
19147	ETE	F. M. Dalla Vecchia. 2006. Telmatosaurus and the other hadrosaurids of the Cretaceous European Archipelago. An overview. Natura Nascosta 32:1-55 [M. Carrano/M. Carrano]
41120		F. M. Dalla Vecchia. 2009. Telmatosaurus and the other hadrosauroids of the Cretaceous European Archipelago. An update. Natura Nascosta 39:1-18 [M. Carrano/M. Carrano]
56054		F. M. Dalla Vecchia. 2014. An overview of the latest Cretaceous hadrosauroid record in Europe. In D. A. Eberth & D. C. Evans (ed.), Hadrosaurs 268-297 [M. Carrano/M. Carrano]
55204		A. Galobart, A. Santos-Cubedo, M. Suñer and J. I. Canudo. 2011. Paleontological discoveries and studies. In À. Galobart, M. Suñer, & B. Poza (eds.), Dinosaurs of Eastern Iberia 1-23 [M. Carrano/M. Carrano/M. Carrano]
80599		E. Isasmendi, A. Torices, J. I. Canudo, P. J. Currie, and X. Pereda-Suberbiola. 2022. Upper Cretaceous European theropod palaeobiodiversity, palaeobiogeography and the intra‐Maastrichtian faunal turnover: new contributions from the Iberian fossil site of Laño. Papers in Palaeontology 8(1):e1419:1-38 [M. Carrano/M. Carrano]
62124		W. G. Joyce. 2017. A Review of the Fossil Record of Basal Mesozoic Turtles. Bulletin of the Peabody Museum of Natural History 58(1):65-113 [P. Holroyd/P. Holroyd/P. Mannion]
56237		Y. Laurent. 2003. Les faunes de vertébrés continentaux du Maastrichtien supérieur d'Europe: systématique et biodiversité [The continental vertebrate faunas of the upper Maastrichtian of Europe: systematics and biodiversity]. Strata, Série 2 41:1-81 [M. Carrano/M. Carrano/M. Carrano]
61140		N. López-Martínez. 2000. Eggshell sites from the Cretaceous-Tertiary transition in south-central Pyrenees (Spain). In A. M. Bravo & T. Reyes (ed.), First International Symposium on Dinosaur Eggs and Babies, Extended Abstracts 95-115 [M. Carrano/M. Carrano]
17699	ETE	N. López-Martínez, J. I. Canudo, and G. Cuenca-Bescós. 1999. Latest Cretaceous eggshells from Arén (southern Pyrénées, Spain). First International Symposium on Dinosaur Eggs and Babies, Isona i Conda Dellà, Spain 35-36 [M. Carrano/M. Carrano]
11838	ETE	N. López-Martínez, J. I. Canudo, L. Ardèvol, X. Pereda Suberbiola, X. Orue-Etxebarria, G. Cuenca-Bescós, J. I. Ruiz-Omeñaca, X. Murelaga, and M. Feist. 2001. New dinosaur sites correlated with Upper Maastrichtian pelagic deposits in the Spanish Pyrenees: implications for the dinosaur extinction pattern in Europe. Cretaceous Research 22(1):41-61 [M. Carrano/M. Carrano/M. Carrano]
83615		M. Moreno-Azanza, L. Ezquerro, M. Pérez-Pueyo and J. M. Gasca. 2021. Huevos de dinosaurio en la Sierra Exteriores de Huesca [Dinosaur eggs in the Sierras Exteriores of Huesca]. Lucas Mallada 23:61-81 [M. Carrano/M. Carrano]
59760		X. Murelaga and J. I. Canudo. 2005. Descripción de los restos de quelonios del Maastrichtiense superior de Aren y Serraduy (Huesca) [Description of chelonian remains from the Upper Maastrichtian of Aren and Seraduy (Huesca)]. Geogaceta 38:51-54 [P. Holroyd/P. Holroyd/M. Uhen]
17698	ETE	O. Oms and J. I. Canudo. 2004. Datación magnetoestratigrafica de los dinosaurios del Cretácico terminal (Maastrichtiense superior) de Arén (Huesca, Unidad Surpirenaica Central) [Magnetostratigraphic dating of the terminal Cretaceous dinosaurs (upper Maastrichtian) of Arén (Huesca, central Pyrénées)]. Geo-Temas 6(5):51-54 [M. Carrano/M. Carrano]
81262		M. Pérez-Pueyo, P. Cruzado-Caballero, M. Moreno-Azanza, B. Vila, D. Castanera, J. M. Gasca, E. Puértolas-Pascual, B. Bádenas, and J. I. Canudo. 2021. The tetrapod fossil record from the uppermost Maastrichtian of the Ibero-Armorican Island: an integrative review based on the outcrops of the western Tremp Syncline (Aragón, Huesca Province, NE Spain). Geosciences 11:162:1-38 [M. Carrano/M. Carrano]
47749		A. Prieto-Marquez, F. M. D. Vecchia, R. Gaete and A. Galobart. 2013. Diversity, relationships, and biogeography of the lambeosaurine dinosaurs from the European archipelago, with description of the new aralosaurin Canardia garonnensis. PLoS One 8(7):e69835:1-44 [P. Mannion/J. Tennant/M. Carrano]
36564		E. Puértolas, J. I. Canudo, and P. Cruzado-Caballero. 2011. A new crocodylian from the late Maastrichtian of Spain: Implications for the initial radiation of crocodyloids. PLoS One 6(6):e20011:1-12 [P. Mannion/P. Mannion/M. Carrano]
76284		E. Puértolas-Pascual, A. Blanco, C. A. Brochu and J. I. Canudo. 2016. Review of the Late Cretaceous-early Paleogene crocodylomorphs of Europe: Extinction patterns across the K-PG boundary. Cretaceous Research 57:565-590 [P. Mannion/P. Mannion]
56091		A. Torices, P. J. Currie, J. I. Canudo and X. Pereda-Suberbiola. 2015. Theropod dinosaurs from the Upper Cretaceous of the South Pyrenees Basin of Spain. Acta Palaeontologica Polonica 60(3):611-626 [P. Mannion/J. Tennant/M. Carrano]
17618	ETE	A. Torices, J. I. Ruiz-Omeñaca, J. I. Canudo and N. López-Martínez. 2004. Nuevos datos sobre los dinosaurios terópodos (Saurischia: Theropoda) del Cretácico superior de los Pirineos Sur-Centrales (Huesca y Lleida) [New data on the theropod dinosaurs (Saurischia: Theropoda) from the Upper Cretaceous of the south-central Pyrenées (Huesca and Lleida)]. Geo-Temas 6(5):71-74 [M. Carrano/M. Carrano]
55315		B. Vila, M. Suñer, A. Santos-Cubedo, J. I. Canudo, B. Poza and A. Galobart. 2011. Saurischians through time. In A. Galobart, M. Suñer, & B. Poza (eds.), Dinosaurs of Eastern Iberia 130-168 [M. Carrano/M. Carrano]