Smokejacks Brickworks: Early/Lower Barremian, United Kingdom
collected by Rivett 1945, 1983-2003, 2007-2009

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Retriletes austroclavatidites
Nye et al. 2008 38 specimens
Schizoporis parvus
Nye et al. 2008 5 specimens
Tetranguladinium conspicuum
Nye et al. 2008 3 specimens
Flabellisporites sp.
Nye et al. 2008
"Clockhousea" sp.
Nye et al. 2008
Straitriletes sulcatus
Nye et al. 2008
Ricinospora sp.
Nye et al. 2008
Taxodiaceapollenites hiatus
Nye et al. 2008 32 specimens
Disaltriangulisporites perplexus
Nye et al. 2008 8 specimens
Iguanodontopus sp.
Austen 2010
cf. Bevhalstia pebja
Austen 2010
cf. Valdicosus chesteri
Austen 2010
Insecta - Odonata
Cretarchistigma greenwoodi Jarzembowski et al. 1998
Jarzembowski et al. 1998 1 specimen
MNEMG 1996.229
Insecta - Odonata - Aeschnidiidae
Aeschnidiidae indet. Handlirsch 1906
Fleck and Nel 2003 1 specimen
MNEMG S 1408 (1 measurement)
Lleidoaeschnidium maculatum n. sp. Fleck and Nel 2003
Fleck and Nel 2003 2 specimens
MNEMG S 1580, MNEMG S 612 (2 measurements)
Insecta - Odonata - Aktassiidae
Pseudocymatophlebia hennigi n. gen., n. sp. Nel et al. 1998
Nel et al. 1998 3 specimens
MNEMG 1996.224ab, MNEMG 1996.225a, MNEMG 1996.225b (2 measurements)
Insecta - Odonata - Cymatophlebiidae
Valdaeshna mikei n. sp. Zheng et al. 2018
Zheng et al. 2018 1 specimen
NHMUK PI II.3105
Insecta - Odonata - Mesochlorogomphidae
Mesochlorogomphus crabbi n. gen., n. sp. Fleck et al. 2008
Fleck et al. 2008 1 specimen
MNEMG 2008.27 (1 measurement)
Insecta - Odonata - Araripelibellulidae
Cretaneophya strevensi n. gen., n. sp. Jarzembowski and Nel 1996
Jarzembowski and Nel 1996 2 specimens
BMB 018810, MNEMG 1995.201
Insecta - Hymenoptera - Angarosphecidae
Archisphex boothi Jarzembowski 1991
Rasnitsyn et al. 1998 1 specimen
NHM In.64650/64656
Insecta - Hymenoptera - Baissidae
Manlaya ockleyensis n. sp. Rasnitsyn and Jarzembowski 1998
Rasnitsyn et al. 1998 1 specimen
BMB 016405/6 (coll. A.J. Ross) (1 measurement)
Insecta - Mecoptera - Englathaumatidae
Englathauma mellishae n. sp. Novokshonov et al. 2016
Novokshonov et al. 2016 6 specimens
NHMUK II.1396, 1394, 1395, 1397, 1399, 1400 (1 measurement)
Insecta - Neuroptera - Psychopsidae
Micropsychops parallelus n. gen., n. sp. Jepson et al. 2009
Jepson et al. 2009 1 specimen
BMNH Pal.PI II 1518a,b (1 measurement)
Cretapsychops corami n. gen., n. sp. Jepson et al. 2009
Jepson et al. 2009 1 specimen
BMNH Pal.PI II 1555a (1 measurement)
Valdipsychops logunovi n. gen., n. sp. Jepson et al. 2009
Jepson et al. 2009 1 specimen
recombined as Pterinoblattina logunovi
BMNH Pal.PI II 1551a,b (1 measurement)
Valdipsychops brigidae n. sp. Jepson et al. 2009
Jepson et al. 2009 1 specimen
recombined as Pterinoblattina brigidae
MNEMG 2009.36 (1 measurement)
Valdipsychops sp. Jepson et al. 2009
Jepson et al. 2009 1 specimen
synonym of Pterinoblattina
MNEMG 2009.39 (1 measurement)
Valdipsychops proudlovei n. sp. Jepson et al. 2009
Jepson et al. 2009 1 specimen
recombined as Pterinoblattina proudlovei
MNEMG 2009.37 (1 measurement)
Insecta - Neuroptera - Kalligrammatidae
Kalligrammatidae indet. (Handlirsch 1906)
Jepson et al. 2009 1 specimen
MNEMG 2009.40
Insecta - Coleoptera - Ommatidae
Brochocoleus keenani n. sp. Jarzembowski et al. 2013
Jarzembowski et al. 2013 1 specimen
BMNH II 3060.2 (coll. EBJ) (1 measurement)
Cionocoleus watsoni n. sp. Jarzembowski et al. 2013
Jarzembowski et al. 2013 1 specimen
BMB 018806 (coll. EBJ) (1 measurement)
Cionocoleus elizabethae n. sp. Jarzembowski et al. 2013
Jarzembowski et al. 2013 1 specimen
BMNH II 3054 (coll. EBJ) (1 measurement)
Brochocoleus crowsonae n. sp. Jarzembowski et al. 2013
Jarzembowski et al. 2013 2 specimens
recombined as Diluticupes crowsonae
BMNH II 3060.1 (coll. EBJ), 3061 (coll. AR) (2 measurements)
Insecta - Coleoptera
Zygadenia tuberculata (Giebel 1856)
Jarzembowski et al. 2015 1 specimen
BMNH II 3063
Zygadenia angliae (Giebel 1856)
Ponomarenko 2006
Insecta - Hemiptera - Mimarachnidae
Saltissus fennahi n. sp. Luo et al. 2021
Luo et al. 2021 1 specimen
BMB 014919 (part)/NHM UK In 64637 (counterpart) (1 measurement)
Insecta - Orthoptera - Locustopsidae
Mesolocustopsis problematica n. sp. Gorochov et al. 2006
Gorochov et al. 2006
(1 measurement)
Insecta - Orthoptera - Baissogryllidae
Speculogryllus acutispeculum n. gen., n. sp. Gorochov et al. 2006
Gorochov et al. 2006
(1 measurement)
Insecta - Orthoptera - Elcanidae
Panorpidium bimaculatum Gorochov et al. 2006
Gorochov et al. 2006
Insecta - Diptera
Athericites gordoni n. gen., n. sp. Mostovski et al. 2003
Mostovski et al. 2003 1 specimen
recombined as Palaepangonius gordoni
BMB 0123835 (1 measurement)
Athericites kensmithi n. sp. Mostovski et al. 2003
Mostovski et al. 2003 1 specimen
recombined as Palaepangonius kensmithi
NHM In 64655 (1 measurement)
Ostracoda - Podocopida - Cyprididae
? Mantelliana sp. Anderson 1966
Nye et al. 2008 16 specimens
Cypridea clavata
Nye et al. 2008 237 specimens
Damonella cf. pygmaea
Nye et al. 2008 5 specimens
Ostracoda - Podocopida
Stenestroemia cf. cressida
Nye et al. 2008 10 specimens
Stenestroemia informal sp. A
Nye et al. 2008 3 specimens
Bivalvia
Bivalvia indet. Linnaeus 1758
Nye et al. 2008
Reptilia - Testudines
Testudines indet. Batsch 1788
Nye et al. 2008 1 specimen
plate
Reptilia - Crocodylia
Crocodylia indet. (Owen 1842)
Austen 2010
Reptilia - Goniopholididae
? Goniopholididae indet. Cope 1875
Benton and Spencer 1995 1 specimen
Goniopholis sp. Owen 1842
Austen 2005 1 specimen
Reptilia
Iguanodon atherfieldensis Hooley 1925
Benton and Spencer 1995 2 specimens
recombined as Mantellisaurus atherfieldensis
Iguanodon sp. Mantell 1825
1 individual
Iguanodon bernissartensis Boulenger 1881
1 specimen
vertebra
Reptilia - Spinosauridae
Baryonyx walkeri n. gen., n. sp. Charig and Milner 1986
Charig and Milner 1986 1 individual
(4 measurements)
Reptilia
cf. Titanosaurus sp. Lydekker 1877
Benton and Spencer 1995
nomen dubium belonging to Titanosauria
Pterosauria indet. (Kaup 1834)
Austen 2010
Actinopteri - Lepisosteiformes
Lepidotes sp. Agassiz 1832
Charig and Milner 1997
Chondrichthyes - Hybodontiformes - Hybodontidae
Hybodontidae indet. (Agassiz 1834)
Austen 2010
spine
Chlorophyceae - Scenedesmaceae
Scenedesmus bifidus
Nye et al. 2008 1 specimen
Scenedesmus novilunaris He 1992
Nye et al. 2008 191 specimens
Gymnospermopsida
Leioaletes calvatus
Nye et al. 2008 12 specimens
Equisetopsida - Cheirolepidiaceae
Classopollis torosus Burger 1965
Nye et al. 2008 44 specimens
Equisetopsida - Cupressales
Inaperturopollenites dubius Thomson and Pflug 1953
Nye et al. 2008 77 specimens
Angiospermae
Retimonocolpites informal sp. 1 Pierce 1961
Nye et al. 2008 9 specimens
Retimonocolpites informal sp. 2 Pierce 1961
Nye et al. 2008 10 specimens
Retimonocolpites informal sp. 4 Pierce 1961
Nye et al. 2008 1 specimen
Retimonocolpites informal sp. 3 Pierce 1961
Austen 2010
Coniferales - Podocarpaceae
Podocarpidites spp.
Nye et al. 2008 21 specimens
Callialasporites dampieri (Balme 1957)
Nye et al. 2008 48 specimens
original and current combination Zonallapollenites dampieri
Pinopsida - Pinales - Taxaceae
Abietineaepollenites microreticulatus
Nye et al. 2008 3 specimens
unclassified
Perinopollenites elatoides Couper 1958
Nye et al. 2008 15 specimens
Alisporites bilateralis Rouse 1959
Nye et al. 2008 18 specimens
Cerebropollenites mesozoicus Nilsson 1958
Nye et al. 2008 33 specimens
Lycopodiopsida - Lycopodiales - Lycopodiaceae
Coronatispora valdensis Dettman 1963
Nye et al. 2008 10 specimens
Lycopodiopsida - Lycopodiales
Retitriletes austroclavatidites Döring et al. 1963
Nye et al. 2008 38 specimens
Lycopodiopsida - Isoetales
Minerisporites sp. Potonie 1956
Nye et al. 2008
Lycopodiopsida - Isoetales - Isoetaceae
Isoetes sp. Linnaeus 1753
Nye et al. 2008
unclassified
Couperisporites complexus
Nye et al. 2008 15 specimens
Marchantiopsida - Sphaerocarpales
Aequitriradites verrucosus (Cookson and Dettmann 1958)
Nye et al. 2008 2 specimens
original and current combination Cirratriradites verrucosus
Marchantiopsida - Marchantiidae
Triporoletes reticulatus Playford 1971
Nye et al. 2008 88 specimens
unclassified
Coptospora striata Dettman 1963
Nye et al. 2008 11 specimens
Taurocusporites reduncus Stover 1962
Nye et al. 2008 5 specimens
Taurocusporites segmentatus Stover 1962
Nye et al. 2008 47 specimens
Microreticulatisporites crassiexinosus
Nye et al. 2008 7 specimens
Pilosisporites trichopapillosus Delcourt and Sprumont 1955
Nye et al. 2008 80 specimens
Lycopsida
Foveotriletes subtriangularis Brenner 1963
Nye et al. 2008 6 specimens
Kraeuselisporites hastilobatus
Nye et al. 2008 62 specimens
unclassified
Impardecispora apiverrucata Venkatachala et al. 1968
Nye et al. 2008 4 specimens
Pteridopsida
Echinatisporis varispinosus Srivastava 1975
Nye et al. 2008 204 specimens
Cyathidites minor Couper 1953
Nye et al. 2008 76 specimens
Cyathidites australis Couper 1953
Nye et al. 2008 42 specimens
Polypodiopsida - Blechnaceae
Leptolepidites verrucatus Couper 1953
Nye et al. 2008 1573 specimens
Polypodiopsida - Osmundaceae
Baculatisporites comaumensis Potonie 1953
Nye et al. 2008 91 specimens
Polypodiopsida - Gleicheniaceae
Gleicheniidites circinidites Dettmann 1963
Nye et al. 2008 4 specimens
Polypodiopsida
Verrucosisporites obscurilaesuratus Pocock 1962
Nye et al. 2008 12 specimens
Polypodiopsida - Polypodiales - Pteridaceae
Contignisporites dorsostriatus
Nye et al. 2008 3 specimens
Polypodiopsida - Schizaeales
Plicatella robusta
Nye et al. 2008 12 specimens
Plicatella concentrica
Nye et al. 2008 9 specimens
Plicatella tricornitata Potonié 1960
Nye et al. 2008 40 specimens
Polypodiopsida - Schizaeales - Schizaeaceae
Cicatricosisporites myrtellii
Nye et al. 2008 49 specimens
Cicatricosisporites mohriodes
Nye et al. 2008 48 specimens
Cicatricosisporites angicanalis
Nye et al. 2008 194 specimens
Cicatricosisporites spp.
Nye et al. 2008 623 specimens
Cicatricosisporites australiensis Potonié 1956
Nye et al. 2008 36 specimens
Cicatricosisporites brevilaesuratus Kemp 1970
Nye et al. 2008 7 specimens
Cicatricosisporites hughesi Dettmann 1963
Nye et al. 2008 67 specimens
Cicatricosisporites hallei Delcourt and Sprumont 1955
Nye et al. 2008 112 specimens
Klukisporites foveolatus Pocock 1965
Nye et al. 2008 19 specimens
Undulatisporites undulapolus Brenner 1963
Nye et al. 2008 13 specimens
Polypodiopsida - Gleicheniales - Matoniaceae
Weichselia sp. Stiehler 1857
Austen 2010
Dictyophyllidites equiexinus Dettman 1963
Nye et al. 2008 9 specimens
unclassified
Foveoinaperturites forameniferus
Nye et al. 2008 5 specimens
Concavissimisporites sp. Delcourt and Sprumont 1955
Nye et al. 2008 18 specimens
Cycadopsida - Cycadales
Eucommiidites troedssonii Potonié 1958
Nye et al. 2008 2 specimens
unclassified
Coniferophyta indet.
Nye et al. 2008 209 specimens
other bisaccate pollen
Ginkgoopsida - Ephedrales
Ephedripites spp.
Nye et al. 2008 15 specimens
Ephedripites patapscoensis Brenner 1963
Nye et al. 2008 2 specimens
unclassified
Erlansonisporites sp. Potonié 1956
Nye et al. 2008
Chlorococcaceae
Tetraedron paraincus Batten 1988
Nye et al. 2008 1 specimen
see common names

Geography
Country:United Kingdom State/province:England County:Surrey
Coordinates: 51.1° North, 0.3° West (view map)
Paleocoordinates:41.4° North, 10.2° East
Basis of coordinate:based on nearby landmark
Geographic resolution:outcrop
Time
Period:Cretaceous Epoch:Early/Lower Cretaceous
Stage:Barremian 10 m.y. bin:Cretaceous 2
Key time interval:Early/Lower Barremian Small shelly zone: Cypridea clavata
Age range of interval:125.77000 - 121.40000 m.y. ago
Stratigraphy
Geological group:Wealden Formation:Weald Clay Member:Upper
Stratigraphic resolution:bed
Stratigraphy comments: about 7 m below the base of BGS Bed 5c, Cypridea clavata zone; Hughes's MCT phase 2 (1994)
Lithology and environment
Primary lithology:bioturbation,fine,concretionary,gray,green lithified sandy claystone
Secondary lithology:very fine,sideritic lithified siltstone
Includes fossils?Y
Includes fossils?Y
Lithology description: "most were encased in nodules of sideritic siltstone...but some lay unprotected in soft clay." "fine silty clay...light olive-grey in colour and containing large irregular nodules of bioturbated sideritic siltstone."
Environment:pond Tectonic setting:passive margin
Geology comments: "quiet, low-energy nature of...environment"; "non-marine origin." "fluvial and/or mudplain environment with areas of shallow water, lagoons and marsh"; "A temporary, shallow body of water on a marshy floodplain."
Taphonomy
Modes of preservation:body,mold/impression,trace
Size of fossils:macrofossils,mesofossils
Spatial orientation:life position
Preservation of anatomical detail:good
Articulated whole bodies:many
Disassociated major elements:some
Disassociated minor elements:some
Fragmentation:occasional
Bioerosion:occasional
Encrustation:none
Feeding/predation traces:fractures,gastric dissolution
Temporal resolution:snapshot
Spatial resolution:autochthonous
Collection methods and comments
Collection methods:selective quarrying,surface (float),surface (in situ),mechanical,field collection
Rock censused:1000 cm2 (area)
Reason for describing collection:taxonomic analysis
Museum repositories:BMNH
Collectors:Rivett Collection dates:1945, 1983-2003, 2007-2009
Metadata
Also known as:Baryonyx type, Ockley, Smokejack's
Database number:45193
Authorizer:M. Carrano, M. Clapham Enterer:M. Carrano, K. Maguire, J. Karr, M. Clapham
Modifier:M. Carrano Research group:paleoentomology,vertebrate
Created:2004-10-13 14:25:23 Last modified:2021-12-13 13:56:37
Access level:the public Released:2004-10-13 14:25:23
Creative Commons license:CC BY
Reference information

Primary reference:
55264. W. H. E. Rivett. 1956. On some reptilian bones from the Weald Clay of Surrey. Proceedings of the Geologists' Association 67(1540):110-111 [M. Carrano/M. Carrano]

Secondary references:

63729 Anonymous. 1983. The beast that roamed Surrey 124 million years ago. New Scientist 99(1368):268 [M. Carrano/M. Carrano]
63731 Anonymous. 1983. Claw leads to discovery of ‘new’ dinosaur. Science News 124(5):70 [M. Carrano/M. Carrano]
17082ETE P. Austen. 2005. Smokejacks fieldwork 2001-2004. Wealden News 6:6-8 [M. Carrano/M. Carrano/M. Carrano]
32812 P. Austen. 2010. Smokejacks fieldwork 2007-2009. Wealden News (8)28-31 [M. Carrano/M. Carrano]
78161 D. J. Batten and P. A. Austen. 2011. The Wealden of south-east England. In D. J. Batten (ed.), English Wealden Fossils. The Palaeontological Association Field Guide to Fossils 14:15-51 [M. Carrano/M. Carrano]
15587ETE M. J. Benton and P. S. Spencer. 1995. Fossil Reptiles of Great Britain. Chapman & Hall, London 1-386 [M. Carrano/K. Maguire/J. Tennant]
11817ETE A. J. Charig and A. C. Milner. 1986. Baryonyx, a remarkable new theropod dinosaur. Nature 324(6095):359-361 [M. Carrano/M. Carrano/M. Carrano]
11816ETE A. J. Charig and A. C. Milner. 1990. The systematic position of Baryonyx walkeri, in the light of Gauthier's reclassification of the Theropoda. In K. Carpenter and P. J. Currie (eds.), Dinosaur Systematics: Perspectives and Approaches 127-140 [M. Carrano/M. Carrano/M. Carrano]
11766ETE A. J. Charig and A. C. Milner. 1997. Baryonyx walkeri, a fish-eating dinosaur from the Wealden of Surrey. Bulletin of the Natural History Museum, Geology Series 53(1):11-70 [M. Carrano/M. Carrano/M. Carrano]
35493 G. Fleck and A. Nel. 2003. Revision of the Mesozoic family Aeschnidiidae (Odonata: Anisoptera). Zoologica 153:1-170 [M. Clapham/J. Karr]
38360 G. Fleck, A. Nel, G. Bechly, X. Martínez-Delclòs, E. A. Jarzembowski and R. A. Coram. 2008. New lower cretaceous 'libelluloid' dragonflies (Insecta: Odonata: Cavilabiata) with notes about estimated divergences dates for this group. Palaeodiversity 1:19-36 [M. Clapham/J. Karr/J. Alroy]
35008 A. V. Gorochov, E. A. Jarzembowski, and R. A. Coram. 2006. Grasshoppers and crickets (Insecta: Orthoptera) from the Lower Cretaceous of southern England. Cretaceous Research 27:641-662 [M. Clapham/M. Clapham]
39642 E. A. Jarzembowski, X. Martínez-Delclòs, G. Bechly, A. Nel, R. Coram and F. Escuillié. 1998. The Mesozoic non-calopterygoid Zygoptera: description of new genera and species from the Lower Cretaceous of England and Brazil and their phylogenetic significance (Odonata, Zygoptera, Coenagrionoidea, Hemiphlebioidea, Lestoidea). Cretaceous Research 19:403-444 [M. Clapham/M. Clapham]
35431 E. A. Jarzembowski and A. Nel. 1996. New fossil dragonflies from the Lower Cretaceous of SE England and the phylogeny of the superfamily Libelluloidea (Insecta: Odonata). Cretaceous Research 17:67-85 [M. Clapham/J. Karr/M. Clapham]
53708 E. A. Jarzembowski, B. Wang, H. C. Zhang and Y. Fang. 2015. Boring beetles are not necessarily dull: New notocupedins (Insecta: Coleoptera) from the Mesozoic of Eurasia and East Gondwana. Cretaceous Research 52:431-439 [M. Clapham/M. Clapham]
47319 E. A. Jarzembowski, E. V. Yan, B. Wang and H. C. Zhang. 2013. Brochocolein beetles (Insecta: Coleoptera) from the Lower Cretaceous of northeast China and southern England. Cretaceous Research 44:1-11 [M. Clapham/M. Clapham/M. Clapham]
50293 E. A. Jarzembowski, E. V. Yan, B. Wang and H. Zhang. 2013. Ommatin beetles (Insecta: Coleoptera) from the Lower Cretaceous of northeast China and southern England. Terrestrial Arthropod Reviews 6:135-161 [M. Clapham/M. Clapham]
39590 J. E. Jepson, V. N. Makarkin, and E. A. Jarzembowski. 2009. New lacewings (Insecta: Neuroptera) from the Lower Cretaceous Wealden supergroup of Southern England. Cretaceous Research 30:1325-1338 [M. Clapham/M. Clapham]
83058 C. H. Luo, X. J. Liu, T. Jiang, Z. S. Song, and E. A. Jarzembowski. 2021. The first mimarachnid planthopper (Hemiptera: Fulgoromorpha: Mimarachnidae: Saltissus) from the Wealden (Lower Cretaceous) of southern England. Historical Biology 34:2031-2036 [M. Clapham/M. Clapham]
38314 M. B. Mostovski, E. A. Jarzembowski, and R. A. Coram. 2003. Horseflies and Athericids (Diptera: Tabanidae, Athericidae) from the Lower Cretaceous of England and Transbaikalia. Paleontological Journal 37(2):162-169 [M. Clapham/J. Karr]
38253 A. Nel, G. Bechly, E. Jarzembowski and X. Martínez-Delclòs. 1998. A revision of the fossil petalurid dragonflies (Insecta: Odonata: Anisoptera: Petalurida). Paleontologia Lombarda 10:1-68 [M. Clapham/J. Karr]
58242 V. G. Novokshonov, A. J. Ross, E. Cook, W. Krzemiński, and A. Soszyńska-Maj. 2016. A new family of scorpionflies (Insecta; Mecoptera) from the Lower Cretaceous of England. Cretaceous Research 62:44-51 [M. Clapham/M. Clapham/M. Clapham]
29159ETE E. Nye, S. Feist-Burkhardt, D. J. Horne, A. J. Ross, and J. E. Whittaker. 2008. The palaeoenvironment associated with a partial Iguanodon skeleton from the Upper Weald Clay (Barremian, Early Cretaceous) at Smokejacks Brickworks (Ockley, Surrey, UK), based on palynomorphs and ostracods. Cretaceous Research 29(3):417-444 [M. Carrano/M. Carrano]
40715 A. G. Ponomarenko. 2006. On the Types of Mesozoic Archostematan Beetles (Insecta, Coleoptera, Archostemata) in the Natural History Museum, London. Paleontological Journal 40(1):90-99 [M. Clapham/L. Edwards]
78875 J. D. Radley and P. Allen. 2012. The Wealden (non-marine Lower Cretaceous) of the Weald Sub-basin, southern England. Proceedings of the Geologists' Association 123(2):245-318 [M. Carrano/M. Carrano]
39641 A. P. Rasnitsyn, E. A. Jarzembowski, and A. J. Ross. 1998. Wasps (Insecta: Vespida = Hymenoptera) from the Purbeck and Wealden (Lower Cretaceous) of southern England and their biostratigraphical and palaeoenvironmental significance. Cretaceous Research 19:329-391 [M. Clapham/M. Clapham]
66095 D. R. Zheng, E. A. Jarzembowski, S. C. Chang, B. Wang, and H. C. Zhang. 2018. New cymatophlebiid dragonflies from the Lower Cretaceous of China and England (Odonata: Anisoptera: Cymatophlebiinae, Valdaeshninae). Cretaceous Research 90:311-317 [M. Clapham/M. Clapham]