Macrofauna, Los Valles de Fuentiduenya (Segovia): Vallesian, Spain

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Mammalia - Proboscidea - Gomphotheriidae
Tetralophodon longirostris (Kaup 1832)
Mazo 1981 20 specimens
Upper teeth: 1 D2, 1 D3, 1 P4. Lower teeth: 2 D4, 1 M1, 1 mandible fragment, 3 molar fragments, 10 third molar fragments.
Mammalia - Proboscidea - Deinotheriidae
Deinotherium cf. giganteum Kaup 1829
Mazo 1981 1 specimen
1 molar fragment.
Mammalia - Carnivora - Amphicyonidae
Amphicyon castellanus n. sp. Fraile 1997
Ginsburg et al. 1981 8 specimens
recombined as Magericyon castellanus
upper teeth: 1 right maxialla with P4-M2, 1 canine, 1 2 M1, 2 M1 fragments. Lower teeth: 1 mandible with P4-M2, 1 M1.
Amphicyon major De Blainville 1841
Ginsburg et al. 1981 3 specimens
Amphicyon major eppelsheimensis. 1 lower M2, 1 partial lower M1 and 1 upper P3.
Thaumastocyon dirus n. sp. Ginsburg et al. 1981
Ginsburg et al. 1981 2 specimens
1 lower M1, ! lower M2.
Mammalia - Carnivora - Mustelidae
? Marcetia santigae
Ginsburg et al. 1981 1 specimen
1 upper P4.
Circamustela dechaseauxi
Ginsburg et al. 1981 1 specimen
1 upper M1 fragment.
Eomellivora liguritor Crusafont-Pairo and Ginsburg 1973
Ginsburg et al. 1981 2 specimens
synonym of Eomellivora piveteaui
1 upper P4, 1 ectocuneiform.
Mammalia - Carnivora - Mephitidae
Mephitinae indet. Bonaparte 1845
Ginsburg et al. 1981 3 specimens
1 mandible fragment, 1 upper canine, 1 lower canine.
Mammalia - Carnivora - Ursidae
Ursidae indet. Gray 1825
Ginsburg et al. 1981 3 specimens
2 right calcanea, 1 radius.
Mammalia - Carnivora - Percrocutidae
Pseudailurus quadridentatus
Ginsburg et al. 1981 32 specimens
Lower teeth: 1 mandible, 1 M1, 3 M1 fragments. Upper teeth: 8 canines, 1 P3, 1 P4. 1 humerus - distal epiphysis. Proximal epiphyses: 2 radial, 1 cubito. 5 metacarpals (II-IV), 4 Calcanea, 3 astragali, 1 3rd metatarsal.
Mammalia - Carnivora - Nimravidae
Sansanosmilus jourdani Filhol 1883
Ginsburg et al. 1981 13 specimens
1 mandbile with Ci-P4, 2 lower M1, 1 lower M1 fragment, 1 upper P3, 1 upper M1, 2 proximal radial epiphyses, 3 third metacarpals, 1 navicular, 1 cuneiform.
Mammalia - Carnivora - Felidae
Felidae indet. Gray 1821
Ginsburg et al. 1981 2 specimens
1 humerus distal epiphysis, 1 tibia distal epiphysis.
Machairodus alberdiae n. sp.
Ginsburg et al. 1981 8 specimens
1 mandible with P3-M1, 1 mandible fragment, 1 lower canine, 1 M1 fragment, 2 upper P3, 2 upper canines.
Machairodus aphanistus (Kaup 1833)
Ginsburg et al. 1981 10 specimens
4 lower canines, 1 lower P4, 1 lower P4 fragment, 2 lower M1 fragments, 1 upper P3, 1 upper P4 fragment.
Mammalia - Carnivora - Hyaenidae
Protictitherium crassum Deperet 1892
Ginsburg et al. 1981 32 specimens
all teeth.
Plioviverrops sp. Kretzoi 1938
Ginsburg et al. 1981 2 specimens
2 upper M1.
Lycyaena aff. chaeretis Gaudry 1861
Ginsburg et al. 1981 11 specimens
lower teeth: 1 canine, 1 P2, 1 mandible fragment with P4, 1 M1, 2 M1 fragments. Upper teeth: 1 canine, 2 P3, 2 P4 fragments.
Mammalia - Suidae
Suidae indet. Gray 1821
Morales and Soria 1981 1 specimen
1 lower M2 fragment.
Mammalia - Moschidae
Micromeryx flourensianus Lartet 1851
Morales and Soria 1981 9 specimens
lower teeth: 1 P4, 1 M2, 1 M1, 2 M3. 1 upper P2. 1 humerus distal epiphysis, 1 calcaeum fragment, 1 very eroded astraglus.
Hispanomeryx duriensis n. gen., n. sp.
Morales et al. 1981 10 specimens
10 teeth, plus postcrania.
Mammalia - Giraffidae
Decennatherium pachecoi Crusafont 1952
Morales and Soria 1981 162 specimens
Upper teeth: 1 D3, 1 M3, 1 maxilla with M2-M3. Lower teeth: 1 mandible with D3-M1, 1 mandible with M1-M2, 1 mandible with P4-M1, 3 M3, 2 M2, 5 M1, 5 P4, 7 P3, 1 mandible with P2, 4 P2, 1 ulna, 1 radius, 5 humeri, 5 metacarpals, 4 tibiae, 6 metatarsals, 10 scaphoids, 8 semilunars, 10 pyramidals, 2 pisiforms, 6 magnotrapezoids, 7 unciforms, 10 malleolar, 9 astragali, 4 calcanea, 8 ectocuneiforms, 6 navicularcuboids. Phalanges: 13 first, 12 second, 2 third.
Mammalia - Cervidae
Euprox dicranoceros
Morales and Soria 1981 29 specimens
11 antler fragments, 1 cranial fragment with pedicle. Upper teeth 2 molars, 1 D4, 2 P3, 1 P2. Lower teeth: 1 M2, 2 M3 fragments. 4 astragali, 3 navicular cuboids, 1 tibia distal epiphysis.
Mammalia - Bovidae
Miotragocerus sp. Stromer 1928
Morales and Soria 1981 23 specimens
1 horncore fragment. Upper teeth: 1 M3, 1 M2, 1 M3 fragment, 3 fragmentary molars, 1 P1. 1 magnotrapezoid, 1 proximal tibia fragment, 1 incomplete calcaneum, 1 tuber calcis, . 1 navicularcuboid, 2 astragali, 1 incomplete astragalus, Phalanges: 1 first, 2 second, 2 third. 1 distal metapodial.
Mammalia - Tragulidae
Dorcatherium naui Kaup 1833
Morales and Soria 1981 72 specimens
1 mandible with M1-M2, 1 mandible with D3-M1. Lower teeth: 1 P2, 1 D2, 2 D4, 1 D4 fragment, 1 P3, 1 P4, 4 molars, 2 M3, 1 M3 fragment. Upper teeth: 1 D3 fragment, 1 P4, 6 molars, 2 D4. 2 scaphoids, 1 pyramidal, 1 magnotrapezoide, 2 metacarpals, 2 first phalanges, 5 second phalanges, 3 tibiae, 17 astragali, 4 calcanea, 8 navicularcuboids.
Mammalia - Perissodactyla - Equidae
Hipparion primigenium (von Meyer 1829)
Alberdi 1981 2108 specimens
recombined as Hippotherium primigenium
Hipparion primigenium melendezi. MNI 46 adults and 13 juveniles. Teeth and postcranial specimens.
Mammalia - Perissodactyla - Rhinocerotidae
Dicerorhinus sp. Gloger 1841
Alberdi et al. 1981
Crania and postcrania.
    = Aceratherium simorrense
Alberdi et al. 1981
Aceratherium simorrense duratonense n. ssp. Crania and postcrania.
Aceratherium cf. incisivum (Cuvier 1824)
Alberdi et al. 1981 6 specimens
1 upper P3. Lower teeth: 1 P3, 1 P4, 1 M3. 1 calcaneum, 1 cuboid.
Reptilia - Phasianidae
Palaeortyx gallica Milne-Edwards 1869
Sánchez‐Marco 2021
Reptilia - Suliformes - Phalacrocoracidae
Phalacrocorax ibericus (de Villalta 1963)
Mlikovsky 2002
nomen nudum belonging to Phalacrocorax
see common names

Geography
Country:Spain State/province:Segovia
Coordinates: 41.4° North, 0.3° West (view map)
Paleocoordinates:41.6° North, 1.2° West
Basis of coordinate:stated in text
Time
Period:Neogene Epoch:Miocene
Stage:Tortonian 10 m.y. bin:Cenozoic 6
Key time interval:Vallesian
Age range of interval:11.63000 - 8.70000 m.y. ago
Stratigraphy
Regional section:Duero Basin
Stratigraphy comments: Miocene, MN9-10. There are 5 levels with fossil remains, level X is the highest in the section (approximately 100m above level Y3). However, the earlier large mammal collections are not associated with these divisions, so they have been entered separately.
Lithology and environment
Environment:alluvial fan
Geology comments: A pond system with a little alluvial fan in which most of the bones have been found.
Taphonomy
Size of fossils:macrofossils
Collection methods and comments
Reason for describing collection:taxonomic analysis
Collection method comments: Material from the 1978-1981 excavations is held in the Equipo de Vertebrados del Instituto de Geologia del CSIC, Madrid. Older material is kept at Sabadell Museum.
Metadata
Also known as:Macrofauna, Los Valles de Fuentidueña (Segovia)
Database number:40768
Authorizer:A. Turner, M. Uhen Enterer:H. O'Regan, M. Uhen
Modifier:H. O'Regan Research group:PACED
Created:2004-06-30 09:17:29 Last modified:2004-07-01 07:45:57
Access level:the public Released:2004-06-30 09:17:29
Creative Commons license:CC BY
Reference information

Primary reference:
11005.PACED M. Hoyos, M. A. Garcia del Cura, and S. Ordonyez. 1981. Caracteristicas geologicas de yacimiento de los Valles de Fuentiduenya (Segovia). Estudios Geologicos 37:345-351 [A. Turner/H. O'Regan/H. O'Regan]

Secondary references:

11013PACED M. T. Alberdi. 1981. El genero Hipparions en el yacimineto de Los Valles de Fuentidueña. Estudios Geologicos 37:425-437 [A. Turner/H. O'Regan/H. O'Regan]
11014PACED M. T. Alberdi, L. Ginsburg, and J. Morales. 1981. Rhinocerotidae del yacimiento de Los Valles de Fuentidueña (Segovia). Estudios Geologicos 37:439-465 [A. Turner/H. O'Regan/M. Uhen]
11018PACED M. T. Alberdi, N. Lopez, J. Morales, C. Sese, and D. Soria. 1981. Bioestratigraphia y biogeographia de la fauna de mamiferos de Los Valles de Fuentidueña (Segovia). Estudios geologicos 37:503-511 [A. Turner/H. O'Regan/H. O'Regan]
11011PACED L. Ginsburg, J. Morales, and D. Soria. 1981. Nuevos datos sobre los carnivoros de Los Valles de Fuentidueña (Segovia). Estudios Geologicos 37:383-415 [A. Turner/H. O'Regan/H. O'Regan]
11012PACED A. V. Mazo. 1981. Los proboscideos del yacimiento Neogeno de Los Valles de Fuentidueña (Segovia). Estudios Geologicos 37:417-423 [A. Turner/H. O'Regan/H. O'Regan]
58139 J. Mlikovsky. 2002. Cenozoic Birds of the World. 1-417 [M. Uhen/M. Uhen]
11015PACED J. Morales, S. Moya-Sola, and D. Soria. 1981. Presencia de la familia Moschidae (Artiodactyla, Mammalia) en el Vallesiense de España: Hispanomeryx duriensis novo gen. nova sp. Estudios Geologicos 37:467-475 [A. Turner/H. O'Regan/H. O'Regan]
11016PACED J. Morales and D. Soria. 1981. Los artiodactilos de Los Valles de Fuentidueña (Segovia). Estudios Geologicos 37:477-501 [A. Turner/H. O'Regan/H. O'Regan]
78374 A. Sánchez‐Marco. 2021. Old and new fossil birds from the Spanish Miocene. Journal of Iberian Geology [M. Uhen/M. Uhen]