Layna, Cerro Pelado: Ruscinian, Spain

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Amphibia - Temnospondyli - Bufonidae
cf. Bufo sp. Laurenti 1768
Mammalia - Soricidae
Paenelimnoecus pannonicus
Crochet 1986
Episoriculus gibberodon Petenyi 1864
Mammalia - Erinaceidae
Erinaceus sp. Linnaeus 1758
Mammalia - Carnivora - Ursidae
Ursus ruscinensis
Soria and Morales 1976 2 specimens
1 mandible fragment with M1, 1 lower M2 fragment,
Mammalia - Carnivora - Canidae
Nyctereutes donnezani Depérer 1890
Soria and Aguirre 1976
Mammalia - Carnivora - Felidae
Felinae indet. Trouessart 1885
Felini indet.
Lynx aff. issiodorensis (Croizet and Jobert 1828)
Machairodontinae indet. Gill 1872
Machairodontini indet.
Mammalia - Carnivora - Hyaenidae
Hyaena pyrenaica
    = Hyaena donnezani Viret 1954
Aguirre and Morales 1990
Euryboas lunensis (Del Campana 1914)
    = Chasmaporthetes lunensis Del Campana 1914
Aguirre and Morales 1990
Mammalia - Bovidae
Gazella borbonica
Leptobos sp. Rutimeyer 1878
Mammalia - Perissodactyla - Equidae
Hipparion fissurae Crusafont and Sondaar 1971
Mammalia - Perissodactyla - Rhinocerotidae
Dicerorhinus miquelcrusafonti
Mammalia - Lagomorpha - Ochotonidae
Prolagus aff. michauxi
Prolagus ibericus
Mammalia - Lagomorpha - Leporidae
Lepus sp. Linnaeus 1758
Oryctolagus laynensis
Mammalia - Rodentia - Cricetidae
Blancomys neglectus
Mimomys stehlini
Trilophomys pirenaicus
Ruscinomys europaeus Depéret 1890
Mammalia - Rodentia - Muridae
Stephanomys donnezani
Occitanomys brailloni Michaux 1969
Rhagapodemus frequens
Castillomys crusafonti Michaux 1969
Apodemus cf. dominans
Apodemus sp. cf. dominans
Apodemus jeanteti
Anthracomys meini
Mammalia - Rodentia - Gliridae
Hypnomys sp. Bate 1918
Eliomys intermedius Priant 1953
Mammalia - Rodentia - Sciuridae
Sciurus sp. Linnaeus 1758
Mammalia - Rodentia - Hystricidae
Hystrix sp. Linnaeus 1758
Mammalia - Primates - Cercopithecidae
Dolichopithecus ruscinensis Depéret 1889
Reptilia - Varanidae
Varanus sp. Merrem 1820
Reptilia - Serpentes
Serpentes indet. Linnaeus 1758
Reptilia - Colubridae
Malpolon mlynarskii n. sp. Szyndlar 1988
Szyndlar 1988 26 specimens
1 anterior braincase (MNCN 10000 = holotype), 27 precaudal vertebrae (MNCN 10001 to 10025)
cf. Macroprotodon sp. Guichenot 1850
Szyndlar 1988 1 element
1 "tiny" vertebra (MNCN 10026), a re-identification of Coronella sp. as listed by Jaén & Sanchiz (1985)
Reptilia - Elapidae
cf. Naja sp. Laurenti 1768
Szyndlar 1988
no number of specimens given
Reptilia - Crotalidae
Vipera sp. Laurenti 1768
Szyndlar 1988 1 element
1 "tiny" vertebra (MNCN 10031)
Vipera maxima n. sp. Szyndlar 1988
Szyndlar 1988 17 specimens
1 trunk vertebra (MNCN 10032 = holotype), 19 precaudal vertebrae (MNCN 10033 to 10048)
Reptilia - Testudines
Chelonia indet. (Latreille 1800)
synonym of Testudines
see common names

Geography
Country:Spain State/province:Soria
Coordinates: 41.1° North, 2.3° West (view map)
Paleocoordinates:41.2° North, 2.7° West
Basis of coordinate:based on nearby landmark
Time
Period:Neogene Epoch:Pliocene
10 m.y. bin:Cenozoic 6
Key time interval:Ruscinian
Age range of interval:5.33300 - 3.10000 m.y. ago
Stratigraphy
Stratigraphy comments: Ruscinian = Alfambrian, MN15, Middle Pliocene.
Site was found during iron mine prospecting in 1966. It is a karst deposit in Jurassic magnesiferous limestones.
Lithology and environment
Environment:karst indet.
Geology comments: red karst deposit.
Taphonomy
Size of fossils:macrofossils,mesofossils
Collection methods and comments
Collection methods:bulk,field collection
Reason for describing collection:general faunal/floral analysis
Collection method comments: The mine prospcetors hole was enlarged to 5x4x4 metres and some bones were recovered during this excavation in 1967, Later excavations from 1972 onwards were conducted by dividing the site into 2m squares and sieved.
Site discoverd by Aguirre and excavated by Crusafont, Aguirre and Michaux.
Metadata
Database number:40548
Authorizer:A. Turner, J. Mueller, L. van den Hoek Ostende Enterer:H. O'Regan, T. Liebrecht, L. van den Hoek Ostende
Modifier:H. O'Regan Research group:PACED
Created:2004-06-25 04:28:07 Last modified:2004-06-25 07:28:07
Access level:the public Released:2004-06-25 04:28:07
Creative Commons license:CC BY
Reference information

Primary reference:
10947.PACED E. Aguirre and D. Soria. 1981. Accumulated bones in a Pliocene cave in Cerro Pelado, Spain. National Geographic Society Research Reports 13:69-81 [A. Turner/H. O'Regan/H. O'Regan]

Secondary references:

10948PACED E. Aguirre and J. Morales. 1990. Villafranchian faunal record of Spain. Quartärpaläontologie 8:7-11 [A. Turner/H. O'Regan/T. Liebrecht]
19698 J.-Y. Crochet. 1986. Insectivores pliocènes du sud de la France (Languedoc-Roussillon) et du nord-est de l'Espagne. Palaeovertebrata 16(3):145-171 [L. van den Hoek Ostende/L. van den Hoek Ostende]
11046PACED D. Soria and E. Aguirre. 1976. El canido de Layna: revision de los Nyctereutes fosiles. Trabajos sobre Neogeno-Cuaternario 5:83-116 [A. Turner/H. O'Regan/H. O'Regan]
11048PACED D. Soria and J. Morales. 1976. Hallazgo de un Ursido en el yacimiento de Layna (Soria). Trabajos sobre Neogeno-Cuaternario 5:129-140 [A. Turner/H. O'Regan/H. O'Regan]
43604 Z. Szyndlar. 1988. Two new extinct species of the Genera Malpolon and Vipera (Reptilia, Serpentes) from the Pliocene of Layna (Spain). Acta Zoologica Cracoviensia 31:607-706 [J. Mueller/T. Liebrecht]