Cerrada Roya (CRH): Early/Lower Barremian, Spain
collected by Herrero

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Actinopteri - Pycnodontiformes
Pycnodontidae indet. (Agassiz 1833)
Canudo et al. 1997
    = Pycnodontiformes indet. Berg 1937
Ruiz-Omeñaca et al. 2004
Actinopteri
"Teleostei" indet. Müller 1846
Canudo et al. 1997
    = Teleostei indet. Müller 1846
Ruiz-Omeñaca et al. 2004
"Holostei" indet. (Müller 1846)
Canudo et al. 1997
Actinopteri - Lepisosteiformes
Lepidotes sp. Agassiz 1832
Canudo et al. 1997
    = "Lepidotes" sp. Agassiz 1832
Ruiz-Omeñaca et al. 2004
Actinopteri - Amiiformes
Amiidae indet. Huxley 1861
Canudo et al. 1997
    = Amiiformes indet. Hay 1929
Ruiz-Omeñaca et al. 2004
Reptilia - Spinosauridae
Spinosaurinae indet. Sereno et al. 1998
Sánchez-Hernández et al. 2007 4 specimens
MPG CR(m)-1, MPG CR(m)-2, MPG CR(m)-4, MPG CR(m)-6
Reptilia - Nodosauridae
Nodosauridae indet. Marsh 1890
Canudo et al. 1997
Reptilia - Dryosauridae
cf. Valdosaurus sp. Galton 1977
Sanz et al. 1987 1 specimen
CRH-1-1, femur
    = ? Valdosaurus sp. Galton 1977
Ruiz-Omeñaca and Canudo 2003
    = Dryosauridae indet. Milner and Norman 1984
Galton 2009
Reptilia - Camarasauridae
Titanosauridae indet. Lydekker 1885
Barco et al. 1999 1 specimen
    = Titanosauriformes indet. Salgado et al. 1997
Canudo et al. 2001
    = ? Camarasauridae indet. Cope 1877
Ruiz-Omeñaca et al. 2004
Reptilia
Pterodactyloidea indet. (Plieninger 1901)
Sánchez-Hernández et al. 2007
Teeth, morphotype 2
Pterosauria indet. Kaup 1834
Ruiz-Omeñaca et al. 2004
    = Ornithocheiridae indet. Seeley 1870
Sánchez-Hernández et al. 2007
Teeth, morphotypes 1 and 3
Reptilia - Istiodactylidae
Istiodactylidae indet. Howse et al. 2001
Sánchez-Hernández et al. 2007
Teeth, morphotype 4
Reptilia
Neosuchia sp. Clark 1988
Ruiz-Omeñaca et al. 2004
Theriosuchus sp. Owen 1878
Ruiz-Omeñaca et al. 2004
    = aff. Theriosuchus sp. Owen 1878
Tennant et al. 2016
Bernissartia sp. Dollo 1883
1 individual
    = Bernissartia fagesii Dollo 1883
Buscalioni et al. 1984
CR 82 - housed in the Zoology Department, University Autonoma de Madrid - immature specimen including a complete skull and postcranial skeleton
Reptilia - Squamata
Lacertilia indet. Owen 1842
Canudo et al. 1997
    = Squamata indet. Oppel 1811
Ruiz-Omeñaca et al. 2004
Reptilia - Paramacellodidae
Paramacellodus sp. Hoffstetter 1967
Richter 1994 5 specimens
P2-H 1-5, five tooth or crown fragments
Reptilia - Teiidae
Meyasaurus sp. Vidal 1915
Ruiz-Omeñaca et al. 2004
Ilerdaesaurus sp. Hoffstetter 1966
Canudo et al. 1997
synonym of Meyasaurus
Reptilia - Testudines
Galvechelone lopezmartinezae n. sp. Pérez-García and Murelaga 2012
Pérez-García and Murelaga 2012 1 specimen
Holotype: MPG/CR(m3), articulated antero-medial area of a caparace, retaining the complete first three dorsal vertebrae and the anterior half of the fourth and the dorsal half of the eighth cervical vertebra.
Chelonia indet. (Latreille 1800)
Canudo et al. 1997
synonym of Testudines
Mammalia
Mammalia indet. (Linnaeus 1758)
Ruiz-Omeñaca et al. 2004
Mammalia - Multituberculata - Eobaataridae
? Parendotherium sp. Crusafont-Pairó and Adrover 1966
Hahn and Hahn 2001 1 specimen
P-2.H-3 (m2) as 'Parentotherium vel Eobaatar'
    = Eobaatar sp. Kielan-Jaworowska et al. 1987
Kielan-Jaworowska et al. 2004
Eobaatar hispanicus n. sp. Hahn and Hahn 1992
Hahn and Hahn 1992 1 specimen
Y.H.-1 (holotype right ?p5)
Parendotherium herreroi Crusafont-Pairo and Adrover 1965
Hahn and Hahn 2001 5 specimens
Mammalia - Multituberculata - Paulchoffatiidae
Galveodon nannothus n. gen., n. sp. Hahn and Hahn 1992
Hahn and Hahn 1992 1 specimen
P-2.H-5 (holotype i2)
Chondrichthyes - Rajiformes - Rhinobatidae
Rhinobatos sp. Link 1790
Canudo et al. 1997
Chondrichthyes - Hybodontiformes - Lonchidiidae
Lonchidion microselachos Estes and Sanchíz 1982
Sánchez-Hernández et al. 2007 5 specimens
MPG P-CR(m)-1, MPG P-CR(m)-2, MPG P-CR(m)-3, MPG P-CR(m)-4, MPG P-CR(m)-5
Chondrichthyes - Hybodontiformes - Polyacrodontidae
? Hybodus parvidens Woodward 1916
Canudo et al. 1997
    = Hybodus parvidens Woodward 1916
Ruiz-Omeñaca et al. 2004
recombined as Polyacrodus parvidens
see common names

Geography
Country:Spain State/province:Aragón County:Teruel
Coordinates: 40.6° North, 0.9° West (view map)
Paleocoordinates:30.7° North, 9.2° East
Basis of coordinate:estimated from map
Geographic resolution:outcrop
Time
Period:Cretaceous Epoch:Early/Lower Cretaceous
Stage:Barremian 10 m.y. bin:Cretaceous 2
Key time interval:Early/Lower Barremian
Age range of interval:125.77000 - 121.40000 m.y. ago
Stratigraphy
Formation:Camarillas
Local section:Galve Local bed:435 m
Local order:bottom to top
Stratigraphic resolution:group of beds
Stratigraphy comments: 5-I is the lower of two sub-units (the other is 5-II).
Lithology and environment
Primary lithology: mudstone
Secondary lithology: sandstone
Lithology description: "The thickest unit is lithofacies 5, composed of mudstones, sandstones and conglomerates. The sandstones and gravels are found in channels which may be 8 m thick (Diaz et al. 1984).
Environment:"channel"
Geology comments: "fluvially dominated channels.. low sinuosity channels with vertical acretion (Diaz et al. 1984)"
unidirectional currents
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Preservation of anatomical detail:good
Disassociated minor elements:all
Spatial resolution:parautochthonous
Collection methods and comments
Collection excludes:all microfossils,some macrofossils
Collection methods:field collection
Reason for describing collection:taxonomic analysis
Collectors:Herrero
Collection method comments: Material is in the J. M. Herrero collection (Galve).

Mudstones contain Unionids, and vertebrate remains, including archosaurs and fish teeth. Channels contain fragments of vertebrates and plant remains, occasionally silicified. It is unclear whether these taxa are present at the Cerrada Roya locality.
Metadata
Also known as:Teruel, Galve syncline, Cerrada Roya-Mina, Cerrada Roya Mina, La Mina
Database number:36368
Authorizer:J. Alroy, R. Benson, P. Barrett, M. Carrano, P. Mannion, R. Butler Enterer:R. Whatley, T. Cleary, R. Benson, P. Mannion, M. Carrano, R. Butler
Modifier:M. Carrano Research group:vertebrate
Created:2004-01-26 17:53:52 Last modified:2021-04-15 13:08:00
Access level:the public Released:2004-01-26 17:53:52
Creative Commons license:CC BY
Reference information

Primary reference:
7242.5% 24760M. Diaz, A. Yébenes, A. Goy and J. L. Sanz. 1984. Landscapes inhabited by Upper Jurassic/Lower Cretaceous archosaurs (Galve, Teruel, Spain). In W.-E. Reif & F. Westphal (ed.), Third Symposium on Mesozoic Terrestrial Ecosystems, Short Papers 67-72 [J. Alroy/R. Whatley/M. Uhen]

Secondary references:

28224ETE A. Badiola, J. I. Canudo, and G. Cuenca-Bescós. 2007. Los fósiles de multituberculados del Hauteriviense y Barremiense de la Península Ibérica [The multituberculate fossils from the Hauterivian and Barremian of the Iberian Peninsula]. IV Jornadas Internacionales sobre Paleontología de Dinosaurios y su Entorno, Salas de los Infantes, Spain 47-48 [M. Carrano/M. Carrano]
17666ETE J. L. Barco, J. I. Canudo, J. I. Ruiz-Omeñaca and R. Royo-Torres. 1999. Bones, teeth and tracks: about sauropod dinosaur remains from Aragón (northeastern Spain). In J. I. Canudo & G. Cuenca-Bescós (ed.), IV European Workshop on Vertebrate Paleontology. Programme and Abstracts. Field Guide 22-23 [M. Carrano/M. Carrano/M. Carrano]
37479 A. D. Buscalioni, E. Buffetaut, and J. L. Sanz. 1984. An immature specimen of the crocodilian Bernissartia from the Lower Cretaceous of Galve (Province of Teruel, Spain). Palaeontology 27(4):809-813 [P. Mannion/P. Mannion]
72435% 17900A. D. Buscalioni and J. L. Sanz. 1987. Cocodrilos del Cretacico Inferior de Galve (Teruel, España). Estudios Geologicos (Madrid) extr.:23-43 [J. Alroy/R. Whatley/J. Alroy]
46249 A. D. Buscalioni and J. L. Sanz. 1990. The small crocodile Bernissartia fagesii from the Lower Cretaceous of Galve (Teruel, Spain). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Sciences de la Terre 60:129-150 [P. Mannion/P. Mannion]
16800ETE J. I. Canudo, O. Amo, G. Cuenca-Bescós, A. Meléndez, J. I. Ruiz-Omeñaca and A. R. Soria. 1997. Los vertebrados del Tithónico-Barremiense de Galve (Teruel, España) [The Tithonian-Barremian vertebrates of Galve (Teruel, Spain)]. Cuadernos de Geologia Ibérica 23:209-241 [M. Carrano/M. Carrano]
16674ETE J. I. Canudo, M. Aurell, J. L. Barco, G. Cuenca-Bescós, and J. I. Ruiz-Omeñaca. 2005. Los dinosaurios de la Formación Villar del Arzobispo (Titónico medio–Berriasiense inferior) en Galve (Teruel) [The dinosaurs of the Villar del Arzobispo Formation (middle Tithonian–lower Berriasian) in Galve (Teruel)]. Geogaceta 38:39-42 [M. Carrano/M. Carrano]
53564 J. I. Canudo, J. L. Barco, R. Royo-Torres and J. I. Ruiz-Omeñaca. 2001. Los saurópodos (Dinosauria) del Tithónico (Jurásico Superior) y del Cretácico de Aragón [The sauropoda (Dinosauria) from the Tithonian (Upper Jurassic) and Cretaceous of Aragón]. I Jornadas Internacionales sobre Paleontología de Dinosaurios y Su Entorno Actas:309-318 [M. Carrano/M. Carrano/M. Carrano]
54616 G. Cuenca-Bescós, O. Amo, J. L. Barco, J. I. Canudo, R. Royo-Torres and J. I. Ruiz-Omeñaca. 1999. Dinosaurios de Aragón [Dinosaurs of Aragon]. Zubía 17:235-257 [M. Carrano/M. Carrano]
31500 P. M. Galton. 2009. Notes on Neocomian (Lower Cretaceous) ornithopod dinosaurs from England - Hypsilophodon, Valdosaurus, "Camptosaurus", "Iguanodon" - and referred specimens from Romania and elsewhere. Revue de Paléobiologie, Genève 28(1):211-273 [R. Butler/R. Butler]
38866 G. Hahn and R. Hahn. 1992. Neue Multituberculaten-Zähne aus der Unter-Kreide (Barremium) von Spanien (Galve un Uña). Geologica et Palaeontologica 26:143-162 [R. Benson/R. Benson/R. Benson]
38868 G. Hahn and R. Hahn. 2001. Multituberculaten-Zähne aus der Unter-Kreide (Barremium) von Pié Pajarón (Prov. Cuenca, Spanien). Palaeontologische Zeitschrift 74(4):587-589 [R. Benson/R. Benson]
14365 Z. Kielan-Jaworowska, R. L. Cifelli, and Z.-X. Luo. 2004. Mammals from the age of dinosaurs: Origins, evolution, and structure. Columbia University Press, New York 1-630 [R. Whatley/R. Whatley/R. Whatley]
43066 A. Pérez-García and X. Murelaga. 2012. Galvechelone lopezmartinezae gen. et sp. nov. a new cryptodiran turtle in the Lower Cretaceous of Europe. Palaeontology 55(5):937-944 [R. Benson/R. Benson/M. Carrano]
46755 A. Pérez-García, T. Scheyer, and X. Murelaga. 2013. The turtles from the uppermost Jurassic and Early Cretaceous of Galve (Iberian Range, Spain): anatomical, systematic, biostratigraphic and palaeobiogeographical implications. Cretaceous Research 44:64-82 [P. Mannion/J. Tennant/P. Mannion]
58341 A. Richter. 1994. Lacertilia aus der Unteren Kreide von Una und Galve (Spanien) und Anoual (Marokko). Berliner Geowissenschaftliche Abhandlungen Reihe E 14:1-147 [P. Barrett/T. Cleary]
55492 J. I. Ruiz-Omeñaca, J. I. Canudo, and G. Cuenca-Bescós. 2002. Los dinosaurios del Jurásico y del Cretácico de la Cordillera Ibérica aragonesa [The Jurassic and Cretaceous dinosaurs of the Aragonese Iberian Cordillera]. Actas del I Encuentro Nacional de Estudios sobre la Cordillera Ibérica (ciencias de la naturaleza) 119-130 [M. Carrano/M. Carrano]
55500 J. I. Ruiz-Omeñaca and J. I. Canudo. 2003. Dinosaurios (Saurischia, Ornithischia) en el Barremiense (Cretácico Inferior) de la península Ibérica [Dinosaurs (Saurischia, Ornithischia) in the Barremian (Lower Cretaceous) of the Iberian peninsula]. In F. Pérez Lorente (ed.), Dinosaurios y Otros Reptiles Mesozóicos de España 269-312 [M. Carrano/M. Carrano]
19754ETE J. I. Ruiz-Omeñaca, J. I. Canudo, M. Aurell, B. Bádenas, J. L. Barco, G. Cuenca-Bescós, and J. Ipas. 2004. Estado de las investigaciones sobre los vertebrados del Jurásico Superior y Cretácico Interior de Galve (Teruel) [State of the investigations into the vertebrates of the Upper Jurassic and Lower Cretaceous of Galve (Teruel)]. Estudios Geológicos 60:179-202 [M. Carrano/M. Carrano]
32167ETE B. Sánchez-Hernández, M. J. Benton, and D. Naish. 2007. Dinosaurs and other fossil vertebrates from the Late Jurassic and Early Cretaceous of the Galve area, NE Spain. Palaeogeography, Palaeoclimatology, Palaeoecology 249:180-215 [R. Butler/R. Butler/M. Carrano]
72415% 17900J. L. Sanz, A. D. Buscalioni, M.-L. Casanovas and J.-V. Santafé. 1987. Dinosaurios del Cretácico inferior de Galve (Teruel, España) [Dinosaurs from the Lower Cretaceous of Galve (Teruel, Spain)]. Estudios Geologicos, Volumen Extraordinario Galve-Tremp 45-64 [J. Alroy/R. Whatley/M. Carrano]
59838 J. P. Tennant, P. D. Mannion, and P. Upchurch. 2016. Evolutionary relationships and systematics of Atoposauridae (Crocodylomorpha: Neosuchia): implications for the rise of Eusuchia. Zoological Journal of the Linnean Society 177:854-936 [P. Mannion/P. Mannion]