Las Zabacheras (ZH): Kimmeridgian - Tithonian, Spain
collected by J. M. Herrero, A. Lapparent, D. D. Fernández-Galiano 1934, 1958-1959, 1982

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Reptilia
Sauropoda informal n. sp.
Lapparent 1960 21 specimens
"an undescribed genus," MPG LZ-20 (2 measurements)
    = Brachiosaurinae indet. Huene 1927
Diaz et al. 1984
    = Aragosaurus ischiaticus n. gen., n. sp. Sanz et al. 1987
Sanz et al. 1987
ZH-1 to ZH-19 (and IG specimens) - holotype (postcranial skeleton)
Reptilia - Camarasauridae
Camarasauridae indet. Cope 1877
Royo-Torres and Ruiz-Omeñaca 1999
Additional specimens reported on by Sanz (1982)
Reptilia
Sauropoda indet. Marsh 1878
forearm, tooth (MPZ 97/557)
    = Diplodocidae indet. Marsh 1884
Barco et al. 1999
    = ? Diplodocidae indet. Marsh 1884
Ruiz-Omeñaca et al. 2004
    = Diplodocimorpha indet. Calvo and Salgado 1995
Canudo et al. 2005
cf. Hypsilophodon sp. Huxley 1870
Buscalioni and Sanz 1984 1 specimen
tooth
Theropoda indet. (Marsh 1881)
Barco and Ruiz-Omeñaca 2001 1 specimen
MPZ 01/97
    = Tetanurae indet. Gauthier 1986
Canudo et al. 2005
    = Theropoda indet. Marsh 1881
Carrano et al. 2012
Actinopteri - Lepisosteiformes
Lepidotes sp. Agassiz 1832
Canudo et al. 1997
    = "Lepidotes" sp. Agassiz 1832
Ruiz-Omeñaca et al. 2004
Actinopteri - Amiiformes
Amiidae indet. Huxley 1861
Canudo et al. 1997
    = Amiiformes indet. Hay 1929
Ruiz-Omeñaca et al. 2004
Actinopteri - Pycnodontiformes
Pycnodontidae indet. (Agassiz 1833)
Canudo et al. 1997
    = Pycnodontiformes indet. Berg 1937
Ruiz-Omeñaca et al. 2004
see common names

Geography
Country:Spain State/province:Aragón County:Teruel
Coordinates: 40.7° North, 0.9° West (view map)
Paleocoordinates:31.6° North, 8.6° East
Basis of coordinate:stated in text
Geographic resolution:outcrop
Time
Period:Jurassic Epoch:Late/Upper Jurassic
Key time interval:Kimmeridgian - Tithonian
Age range of interval:157.30000 - 145.00000 m.y. ago
Stratigraphy
Formation:Villar del Arzobispo
Local section:Galve Local bed:Unit 3
Local order:bottom to top
Stratigraphic resolution:bed
Stratigraphy comments: Unit 3 has previously been suggested to be within the El Castellar Formation, and dated as Hauterivian–lower Barremian (see Canudo et al. 2012), but Royo-Torres et al. (2014) argue that Unit 3 is stratigraphically lower than the El Castellar Formation and belongs to the Villar del Arzobispo Formation, which is dated as upper Tithonian–lower Berriasian. Also called the "Zabachera beds."

Most recently been placed in the lower section of the Galve Formation and assigned a probable late Berriasian–early Valanginian age (Aurell et al., 2016).
Villar del Arzobispo is now considered Kimmeridgian-Tithonian age (Campo-Sotos et al. 2017)
Lithology and environment
Primary lithology:red lime mudstone
Secondary lithology:lenticular,current ripples,bioturbation sandy "siliciclastic"
Includes fossils?Y
Lithology description: Formed predominately of red limes, with rare thick lenses of terrigenous deposits (Molina and Yebenes 1987) - described as "red clays" by Lapparent (1960). Unit 3: "The coarser terrigenous material (fine to medium sand) is not typically channelized, but most frequently occurs in lenticular bedding and isolated dunes (Diaz et al. 1984)."
Environment:estuary/bay
Geology comments: "Unit 3 is characterized by the disappearance of the carbonate facies and predominance of mudstones. This is interpreted as a muddy intertidal flat deposit...Despite a large discontinuity existing within unit 3, or at its upper or lower boundaries, the intertidal facies can be integrated withing[sic].." terregenous delta-derived material prograding into a shallow lagoon (Diaz et al. 1987).
Taphonomy
Modes of preservation:body,trace
Size of fossils:macrofossils,mesofossils
Preservation of anatomical detail:good
Articulated whole bodies:some
Disassociated major elements:some
Disassociated minor elements:many
Collection methods and comments
Collection methods:selective quarrying,surface (in situ),mechanical,field collection
Reason for describing collection:taxonomic analysis
Collectors:J. M. Herrero, A. Lapparent, D. D. Fernández-Galiano Collection dates:1934, 1958-1959, 1982
Collection method comments: Material is in the J. M. Herrero collection (Galve). Discovered by Herrero in 1958 and initially collected by him and Fernández-Galiano that year; Lapparent visited the site with Herrero, P. Atrián, and G. Gómez to further collect.

The Aragosaurus holotype specimens are distributed between two collections: (1) the material recovered by Dimas Fernández-Galiano in the 1950s, which is deposited at the Museo Provincial de Teruel with the acronym ‘IG’; and (2) the material excavated by J.M. Herrero and the team of author J.L.S. in the 1980s, with acronym ‘ZH’, now in the Museo Paleontológico de Galve. In 2001, on the occasion of the inauguration of the Dinópolis museum in Galve, the material (fragments of vertebrae and ribs, coracoids, humerus, carpal, metacarpus, and ischia) deposited by Dimas Fernández- Galiano in the Museo Provincial de Teruel was moved to a satellite museum of Dinópolis in Galve (today belonging to the Museo Aragonés de Paleontologia), with the other specimens remaining in the Museo Paleontológico de Galve.
Metadata
Also known as:Teruel, Galve syncline, La Carretera
Database number:36366
Authorizer:J. Alroy, M. Carrano Enterer:R. Whatley, M. Carrano
Modifier:G. Varnham Research group:vertebrate
Created:2004-01-26 12:35:07 Last modified:2022-05-10 06:20:34
Access level:the public Released:2004-01-26 12:35:07
Creative Commons license:CC BY
Reference information

Primary reference:

19730.ETE D. Fernández Galiano. 1960. Yacimientos de dinosaurios en Galve (Teruel) [Dinosaur localities in Galve (Teruel)]. Boletin de la Real Sociedad Española de Historia Natural, Sección Geológica 58:95-96 [M. Carrano/M. Carrano]

Secondary references:

17666ETE J. L. Barco, J. I. Canudo, J. I. Ruiz-Omeñaca and R. Royo-Torres. 1999. Bones, teeth and tracks: about sauropod dinosaur remains from Aragón (northeastern Spain). In J. I. Canudo & G. Cuenca-Bescós (ed.), IV European Workshop on Vertebrate Paleontology. Programme and Abstracts. Field Guide 22-23 [M. Carrano/M. Carrano/M. Carrano]
40453 J. L. Barco and J. I. Ruiz-Omeñaca. 2001. Primeros restos postcraneales de terópodo (Dinosauria, Saurischia) en la Formación Villar del Arzobispo (Tithónico-Berriasiense): Un centro vertebral caudal del yacimiento Carretera (Galve, Teruel) [First postcranial remains of a theropod (Dinosauria, Saurischia) in the Villar del Arzobispo Formation (Tithonian–Berriasian): a caudal vertebral centrum from the Carretera locality (Galve, Teruel)]. In G. Meléndez, Z. Herrera, G. Delvene & B. Azanza (eds.), XVII Jornadas de la Sociedad Española de Paleontología. Los Fósiles y la Paleogeografía. Ayuntamiento de Albarracín y Fundación Conjunto Paleontólogico de Teruel, Albarracin, Spain 5.1:247-254 [M. Carrano/M. Carrano]
31068ETE A. D. Buscalioni and J. L. Sanz. 1984. Los Arcosaurios (Reptilia) del Jurásico superior – Cretácico inferior de Galve (Teruel, España) [The archosaurs (Reptilia) from the Upper Jurassic–Lower Cretaceous of Galve (Teruel, Spain)]. Teruel 71:9-30 [M. Carrano/M. Carrano]
31067 A. D. Buscalioni and J. L. Sanz. 1987. Lista faunistica de los vertebrados de Galve (Teruel) [Faunal list of the vertebrates of Galve (Teruel)]. Estudios Geologicos, Volumen Extraordinario Galve-Tremp 65-67 [M. Carrano/M. Carrano]
16800ETE J. I. Canudo, O. Amo, G. Cuenca-Bescós, A. Meléndez, J. I. Ruiz-Omeñaca and A. R. Soria. 1997. Los vertebrados del Tithónico-Barremiense de Galve (Teruel, España) [The Tithonian-Barremian vertebrates of Galve (Teruel, Spain)]. Cuadernos de Geologia Ibérica 23:209-241 [M. Carrano/M. Carrano]
16674ETE J. I. Canudo, M. Aurell, J. L. Barco, G. Cuenca-Bescós, and J. I. Ruiz-Omeñaca. 2005. Los dinosaurios de la Formación Villar del Arzobispo (Titónico medio–Berriasiense inferior) en Galve (Teruel) [The dinosaurs of the Villar del Arzobispo Formation (middle Tithonian–lower Berriasian) in Galve (Teruel)]. Geogaceta 38:39-42 [M. Carrano/M. Carrano]
53564 J. I. Canudo, J. L. Barco, R. Royo-Torres and J. I. Ruiz-Omeñaca. 2001. Los saurópodos (Dinosauria) del Tithónico (Jurásico Superior) y del Cretácico de Aragón [The sauropoda (Dinosauria) from the Tithonian (Upper Jurassic) and Cretaceous of Aragón]. I Jornadas Internacionales sobre Paleontología de Dinosaurios y Su Entorno Actas:309-318 [M. Carrano/M. Carrano/M. Carrano]
48100 J. I. Canudo, J. M. Gasca, M. Moreno-Azanza and M. Aurell. 2012. New information about the stratigraphic position and age of the sauropod Aragosaurus ischiaticus from the Early Cretaceous of the Iberian Peninsula. Geological Magazine 149(2):252-263 [M. Carrano/M. Carrano]
41617 M. T. Carrano, R. B. J. Benson, and S. D. Sampson. 2012. The phylogeny of Tetanurae (Dinosauria: Theropoda). Journal of Systematic Palaeontology 10(2):211-300 [M. Carrano/M. Carrano]
70544 A. Cobos, R. Royo-Torres, and L. Alcalá. 2012. Restos postcraneales de ornitópodo en el Cretácico Inferior de El Castellar (Subcuenca de Peñagolosa, Teruel) [Ornithopod postcranial remains in the Lower Cretaceous of El Castellar (Peñagolosa Subbasin, Teruel)]. Geogaceta 52:181-184 [M. Carrano/M. Carrano]
54616 G. Cuenca-Bescós, O. Amo, J. L. Barco, J. I. Canudo, R. Royo-Torres and J. I. Ruiz-Omeñaca. 1999. Dinosaurios de Aragón [Dinosaurs of Aragon]. Zubía 17:235-257 [M. Carrano/M. Carrano]
72425% 24760M. Diaz, A. Yébenes, A. Goy and J. L. Sanz. 1984. Landscapes inhabited by Upper Jurassic/Lower Cretaceous archosaurs (Galve, Teruel, Spain). In W.-E. Reif & F. Westphal (ed.), Third Symposium on Mesozoic Terrestrial Ecosystems, Short Papers 67-72 [J. Alroy/R. Whatley/M. Uhen]
43435 V. Díez Díaz, X. Pereda Suberbiola, and J. L. Sanz. 2012. Juvenile and adult teeth of the titanosaurian dinosaur Lirainosaurus (Sauropoda) from the Late Cretaceous of Iberia. Geobios 45:265-274 [M. Carrano/M. Carrano]
25010ETE A. F. d. Lapparent. 1960. Los dos dinosaurios de Galve (provincia de Teruel, España) [The two dinosaurs from Galve (Teruel province, Spain)]. Teruel 24:177-197 [M. Carrano/M. Carrano]
25015ETE A. F. d. Lapparent. 1966. Nouveaux gisements de reptiles Mesozoiques en Espagne [New localities of Mesozoic reptiles in Spain]. Notas y Comunicaciones del Instituto Geologico y Minero de España 84:103-110 [M. Carrano/M. Carrano/M. Carrano]
17670ETE R. Royo-Torres and J. I. Ruiz-Omeñaca. 1999. The pubis of Aragosaurus ischiaticus Sanz, Buscalioni, Casanovas & Santafé, 1987, a camarasaurid sauropod from the Upper Hauterivian (Lower Cretaceous) of Galve, Teruel, Spain. In J. I. Canudo & G. Cuenca-Bescós (ed.), IV European Workshop on Vertebrate Paleontology. Programme and Abstracts. Field Guide 96-97 [M. Carrano/M. Carrano/M. Carrano]
51804 R. Royo-Torres, P. Upchurch, P. D. Mannion, R. Mas, A. Cobos, F. Gascó, L. Alcalá and J. L. Sanz. 2014. The anatomy, phylogenetic relationships, and stratigraphic position of the Tithonian–Berriasian Spanish sauropod dinosaur Aragosaurus ischiaticus. Zoological Journal of the Linnean Society 171(3):623-655 [P. Mannion/P. Mannion/M. Carrano]
55492 J. I. Ruiz-Omeñaca, J. I. Canudo, and G. Cuenca-Bescós. 2002. Los dinosaurios del Jurásico y del Cretácico de la Cordillera Ibérica aragonesa [The Jurassic and Cretaceous dinosaurs of the Aragonese Iberian Cordillera]. Actas del I Encuentro Nacional de Estudios sobre la Cordillera Ibérica (ciencias de la naturaleza) 119-130 [M. Carrano/M. Carrano]
19754ETE J. I. Ruiz-Omeñaca, J. I. Canudo, M. Aurell, B. Bádenas, J. L. Barco, G. Cuenca-Bescós, and J. Ipas. 2004. Estado de las investigaciones sobre los vertebrados del Jurásico Superior y Cretácico Interior de Galve (Teruel) [State of the investigations into the vertebrates of the Upper Jurassic and Lower Cretaceous of Galve (Teruel)]. Estudios Geológicos 60:179-202 [M. Carrano/M. Carrano]
32167ETE B. Sánchez-Hernández, M. J. Benton, and D. Naish. 2007. Dinosaurs and other fossil vertebrates from the Late Jurassic and Early Cretaceous of the Galve area, NE Spain. Palaeogeography, Palaeoclimatology, Palaeoecology 249:180-215 [R. Butler/R. Butler/M. Carrano]
56555 J. L. Sanz. 1984. Las faunas españolas de dinosaurios [The Spanish dinosaur faunas]. I Congreso Español de Geología, Segovia 497-506 [M. Carrano/M. Carrano]
70303 J. L. Sanz. 2000. Yacimientos ibéricos de dinosaurios [Iberian dinosaur localities]. Dinosaurios: Los Señores del Pasado [Dinosaurs: The Lords of the Land] 267-299 [M. Carrano/M. Carrano]
72375% 24740J. L. Sanz, A. D. Buscalioni, M. L. Casanovas and J. V. Santafe. 1984. The archosaur fauna from the Upper Jurassic/Lower Cretaceous of Galve (Teruel, Spain). In W.-E. Reif & F. Westphal (ed.), Third Symposium on Mesozoic Terrestrial Ecosystems, Short Papers 207-210 [J. Alroy/R. Whatley/M. Carrano]
72415% 17900J. L. Sanz, A. D. Buscalioni, M.-L. Casanovas and J.-V. Santafé. 1987. Dinosaurios del Cretácico inferior de Galve (Teruel, España) [Dinosaurs from the Lower Cretaceous of Galve (Teruel, Spain)]. Estudios Geologicos, Volumen Extraordinario Galve-Tremp 45-64 [J. Alroy/R. Whatley/M. Carrano]
19339ETE J. L. Sanz, A. D. Buscalioni, J. J. Moratalla, V. Francés, and M. Antón. 1990. Los reptiles Mesozoicos del registro español [The Mesozoic reptiles from the Spanish record]. Monografias del Museo Nacional de Ciencias Naturales 2:1-79 [M. Carrano/M. Carrano]