Csarnota locality 2: Ruscinian - Villanyian, Hungary

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Megalobatrachidae indet.
2 specimens
Procamptoceros cf. brivatense
Myriapoda
Diplopoda indet. Gervais 1844
Gastropoda
Gastropoda indet. Cuvier 1795
Mammalia - Bovidae
Hemitragus cf. bonali
Gazellospira cf. torticornis
Mammalia - Cervidae
? Cervus pyrenaicus (Mennecart et al. 2017)
recombined as Croizetoceros pyrenaicus
Small Cervidae (Cervus pyrenaicus Deperet?)
Cervus (? Rusa) sp. (Smith 1827)
original and current combination Rusa
Cervus (?Rusa) sp. indet.
Megaceros cf. dupuisi
synonym of Megaloceros
(unlikely according to Kretzoi)
"Alces" sp. Gray 1821
Alces sp. indet.
Mammalia - Carnivora - Felidae
Felis sp. Linnaeus 1758
1 specimen
Felis s. 1. indet. (size of catus)
? Lynx sp. Kerr 1792
Lynx (?) sp. indet.
Mammalia - Carnivora - Ursidae
Ursus sp. Linnaeus 1758
few. Ursus s. 1. sp. indet.
Mammalia - Carnivora - Mustelidae
Mustela aff. palerminea
few.
Baranogale beremendensis
few.
Xenictis pilgrimi
few.
Mammalia - Carnivora - Canidae
Canidae indet. Fischer de Waldheim 1817
few. Canidae I-III indet.
Mammalia - Soricidae
Petenyia hungarica Kormos 1934
100 specimens
c. 100
Sorex sp. Linnaeus 1758
Sorex s. 1. sp.
Blarinoides mariae Sulimski 1959
few
Episoriculus gibberodon Petenyi 1864
100 specimens
c. 100
Beremendia fissidens (Petényi 1864)
400 specimens
c. 400
Petenyiella gracilis
Mammalia - Talpidae
Talpa csarnotana
100 specimens
c. 100
Talpa cf. fossilis Petenyi 1864
rare
Desmana sp. Guldenstaedt 1777
Desmana sp. indet. I-II, rare
Mammalia - Erinaceidae
Hemiechinus cf. samsonowiczi
few
Mammalia - Primates - Cercopithecidae
Macaca sp. Lacépède 1799
Macaca sp. indet. sporadic
cf. Dolichopithecus sp. Depéret 1889
cf. Dolichopithecus sp. indet.
Mammalia - Lagomorpha - Ochotonidae
Ochotonoides csarnotanus
1-2 specimens
Mammalia - Lagomorpha - Leporidae
Hypolagus cf. beremendensis
few.
Mammalia - Rodentia - Sciuridae
Pliopetes hungaricus
few
Sciurus sp. Linnaeus 1758
1 specimen
Sciurus sp. indet.
Pliopetaurista pliocaenica
2 specimens
? Eutamias sp. Trouessart 1880
Eutamias sp. indet. few
Mammalia - Rodentia - Gliridae
Glis minor Kowalski 1963
few
Muscardinus sp. Kaup 1829
few
Glirurus gemmula
1 specimen
Dryomimus eliomyoides Kretzoi 1959
1000 specimens
c. 1000
Mammalia - Rodentia - Cricetidae
Prospalax priscus
600 specimens
c. 600
Propliomys hungaricus
abundant
Promimomys cor
1 specimen
Cseria gracilis
synonym of Mimomys
common
Cricetinus europaeus
few
Baranomys loczyi
few
Dolomys nehringi
common
Mammalia - Rodentia - Muridae
Rhagapodemus frequens
common
Micromys praeminutus
few
Apodemus dominans
8000 specimens
c. 8000
Mammalia - Chiroptera
Chiroptera indet. Blumenbach 1779
common
Reptilia - Lacertidae
Lacerta ruscinensis
several thousand
Lacerta aff. agilis Linneaus 1758
few
Reptilia - Ophidia
Ophidia indet. Latreille 1804
abundant
Reptilia - Colubridae
Elaphe quatuorlineata (Lacépède 1789)
Venczel and Vardai 2000 1 element
a single vertebra (GMH uncatalogued)
Reptilia - Anguidae
Anguis sp. Linnaeus 1758
several thousand
Ophisaurus intermedius
common
Reptilia - Varanidae
Varanus deserticolus Bolkay 1913
few
    = Varanus sp. Merrem 1820
Villa et al. 2018
Reptilia
Aves indet. Linnaeus 1758
Reptilia - Phasianidae
Tetrao macropus Jánossy 1967
few
Francolinus capeki Lambrecht 1933
sporadic. Francolinus capeki wezensis
Reptilia - Testudines - Testudinidae
Testudo cf. lambrechti
rare
Amphibia - Temnospondyli - Albanerpetontidae
Albanerpeton sp. Estes and Hoffstetter 1976
Venczel and Gardner 2003
"new species"
    = Albanerpeton pannonicus n. sp. Venczel and Gardner 2005
Venczel and Gardner 2005
corrected as Albanerpeton pannonicum
MAFI V.22000 - holotype (fused pair of nearly complete premaxillae, articulated with anterior end of right maxilla); referred material: fragmentary skulls (n = 24), each consisting of either a maxillary arcade comprising a maxilla articulated with some combination of a jugal, lacrimal, prefrontal and palatal bones or an incomplete skull roof comprising fused frontals articulated with some combination of a prefrontal, lacrimal and nasal: MAFI V.22003 ⁄ 1, V.22011, V.22012 ⁄ 1–3, V.22015, V.22021 ⁄ 1–2, V.22033 ⁄ 2, V.22034, V.22039, V.22053, V.22055, V.22075, V.22076 ⁄ 1–2, V.22077, V.22093 ⁄ 1–3, V.22094 ⁄ 1, V.22100 ⁄ 1–2, V.22111; fused premaxillae (n = 6): MAFI V.22017 ⁄ 1–2, V.22023, V.22026 ⁄ 1, V.22041 ⁄ 1, V.22072 ⁄ 1; isolated premaxillae (n = 38): MAFI V.22006 ⁄ 1–5, V.22007, V.22018 ⁄ 1–3, V.22026 ⁄ 2, V.22030 ⁄ 1–2, V.22036 ⁄ 1–4, V.22041 ⁄ 2, V.22050, V.22057, V.22067, V.22072 ⁄ 2–3, V.22078 ⁄ 1–2, V.22083, V.22089 ⁄ 1–3, V.22095, V.22101 ⁄ 1–2, V.22105, V.22106 ⁄ 1–3, V.22113 ⁄ 1–3; isolated maxillae (n = 50): MAFI V.22002 ⁄ 1–6, V.22013 ⁄ 1–10, V.22014 ⁄ 1–2, V.22028 ⁄ 1, V.22038 ⁄ 1–3, V.22043 ⁄ 1–2, V.22048, V.22056 ⁄ 5, V.22070 ⁄ 1–4, V.22087 ⁄ 1–2, V.22092 ⁄ 1–4, V.22099 ⁄ 1–4, V.22110 ⁄ 1–10; fused frontals, not articulated with other bones (n = 39): MAFI V.22003 ⁄ 2–3; V.22004 ⁄ 1–3, V.22005 ⁄ 1–4, V.22016 ⁄ 1–7, V.22022, V.22025, V.22029, V.22035, V.22040, V.22045, V.22060, V.22063, V.22066 ⁄ 1–2, V.22071, V.22081, V.22082 ⁄ 1–2, V.22088, V.22094 ⁄ 2, V.22104 ⁄ 1–5, V.22112 ⁄ 1–2; mandibles (n = 20): MAFI V.22009, V.22010 ⁄ 1–4, V.22020 ⁄ 1–4, V.22032 ⁄ 1–2, V.22047, V.22052, V.22059, V.22062, V.22065, V.22080, V.22097, V.22108 ⁄ 1–2; dentaries (n = 302): MAFI V.22001 ⁄ 1– 18, V.22008 ⁄ 1–38, V.22019 ⁄ 1–37, V.22024 ⁄ 1–2, V.22027 ⁄ 1–8, V.22031 ⁄ 1–10, V.22037 ⁄ 1–24, V.22042 ⁄ 1–3, V.22044 ⁄ 1–6, V.22046 ⁄ 1–10, V.22049, V.22051 ⁄ 1–4, V.22056 ⁄ 1–4, V.22058 ⁄ 1– 3, V.22061 ⁄ 1–10, V.22064 ⁄ 1–3, V.22068 ⁄ 1–12, V.22073 ⁄ 1–4, V.22079, V.22084, V.22085 ⁄ 1–11, V.22090 ⁄ 1–17, V.22096 ⁄ 1–23, V.22102 ⁄ 1–11, V.22107 ⁄ 1–25, V.22114 ⁄ 1–16; presacral vertebrae (n = 10): MAFI V.22028 ⁄ 2, V.22054, V.22074 ⁄ 1, V.22086 ⁄ 1–2, V.22098 ⁄ 1, V.22103 ⁄ 1, V.22109 ⁄ 1–2, V.22114 ⁄ 17; humeri (n = 16): MAFI V.22033 ⁄ 1, V.22069, V.22074 ⁄ 2, V.22086 ⁄ 3, V.22091 ⁄ 1–5, V.22098 ⁄ 2–4, V.22103 ⁄ 2–3, V.22109 ⁄ 3–4.
Amphibia - Temnospondyli - Urodela
Salamandra sp. Laurenti 1768
rare
Triturus sp. Rafinesque 1815
sporadic
Amphibia - Temnospondyli
Anura indet. (Fischer von Waldheim 1813)
(minly Bufo (listed seperately) c. 20,000 specimens in total)
Amphibia - Temnospondyli - Bufonidae
Bufo sp. Laurenti 1768
Pisces
Pisces indet. Linnaeus 1758
synonym of Osteichthyes
see common names

Geography
Country:Hungary State/province:Barany
Coordinates: 45.9° North, 18.2° East (view map)
Paleocoordinates:45.8° North, 18.1° East
Basis of coordinate:based on nearby landmark
Time
Period:Neogene Epoch:Pliocene
10 m.y. bin:Cenozoic 6
Key time interval:Ruscinian - Villanyian
Age range of interval:5.33300 - 2.58000 m.y. ago
Stratigraphy
Stratigraphy comments: Csarnotian stage, Mid. Pliocene, MN15 (according to Fejfar et al., 1997). Janossy puts it slightly later (in the Villanyian). Villafranchian.
Lithology and environment
Primary lithology: not reported
Lithology description: 'red clay.'
Environment:karst indet.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils,mesofossils
Collection methods and comments
Collection methods:bulk,field collection
Reason for describing collection:biostratigraphic analysis
Collection method comments: This is a reconstructed faunal list, including several collections excavated from 1910 onwards. The original site lists (1910-1930s) do not give exact locality info., but the later excavations uncovered 25 strata in a trench 1m x 1.5m and 3m deep.
Taxonomic list comments:'Two Pliogallus species described by Galliard have been deleted from the list.'
Metadata
Also known as:Csarnóta 2
Database number:35354
Authorizer:A. Turner, J. Head, J. Mueller, M. Carrano, P. Mannion Enterer:H. O'Regan, T. Liebrecht, M. Carrano, P. Mannion, G. Varnham
Modifier:M. Carrano Research group:PACED
Created:2003-12-05 04:26:08 Last modified:2021-07-14 11:11:08
Access level:the public Released:2003-12-05 04:26:08
Creative Commons license:CC BY
Reference information

Primary reference:
9534.PACED D. Janossy. 1986. Pleistocene vertebrate faunas of Hungary. Developments in Palaeontology and Stratigraphy, 8. Elsevier, Amsterdam 1-208 [A. Turner/H. O'Regan/H. O'Regan]

Secondary references:

77386 J. D. Gardner, A. Villa, S. Colombero, M. Venczel, and M. Delfino. 2021. A Messinian (latest Miocene) occurrence for Albanerpeton Estes & Hoffstetter, 1976 (Lissamphibia: Albanerpetontidae) at Moncucco Torinese, Piedmont Basin, northwestern Italy, and a review of the European Cenozoic record for albanerpetontids. Geodiversitas 43(14):391-404 [M. Carrano/M. Carrano]
23463ETE M. Venczel and J. Gardner. 2003. The geologically youngest albanerpetontid amphibian. Journal of Vertebrate Paleontology 23(3, suppl.):106A [M. Carrano/M. Carrano]
44192 M. Venczel and J. D. Gardner. 2005. The geologically youngest albanerpetontid amphibian, from the Lower Pliocene of Hungary. Palaeontology 48(6):1273-1300 [P. Mannion/P. Mannion]
54746 M. Venczel and G. Vardai. 2000. The Genus Elaphe in the Carpathian Basin: Fossil Record. Nymphaea – Folia Naturae Bihariae 28:65-82 [J. Mueller/T. Liebrecht]
69946 A. Villa, J. Abella, D. M. Alba, S. Almecija, A. Bolet, G. D. Koufos, F. Knoll, A. H. Lujan, J. Morales, J. M. Robles, I. M. Sanchez and M. Delfino. 2018. Revision of Varanus marathonensis (Squamata, Varanidae) based on historical and new material: morphology, systematics, and paleobiogeography of the European monitor lizards. PLoS ONE 13(12):1-46 [E. Vlachos/E. Vlachos]