Holbrook quarry, UCMP V3922: Aegean - Bithynian, Arizona
collected by S. Welles 1939, 1940

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Plantae indet. Haeckel 1866
carbonized remains
Polypodiopsida - Equisetales - Calamitaceae
Calamites sp. wood Suckow 1784
casts
Ptycholepiformes - Ptycholepididae
? Boreosomus sp. Stensiö 1921
1 specimen
ganoid scales
Reptilia
Reptilia indet. Laurenti 1768
50 elements
At least 50 elements, some of which may belong to Arizonasaurus and/or Anisodontosaurus.
? Arizonasaurus babbitti Welles 1947
1 specimen
UCMP 36231, partial maxilla
    = Archosauria indet. Cope 1869
Nesbitt 2005
Poposauroidea informal unnamed species
Nesbitt 2005
"Chatterjeea-like taxon"
Arizonasaurus babbitti n. gen., n. sp. Welles 1947
6 specimens
UCMP 36232, anterior of left maxilla. UCMP 36233, 37815, ilia, UCMP 37812, 37818, vertebrae, UCMP 37803, femur
Reptilia - Chirotheriidae
Chirotherium sp. Kaup 1835
Reptilia
Anisodontosaurus greeri n. gen., n. sp. Welles 1947
1 specimen
UCMP 37804, left mandible with teeth
Amphibia - Temnospondyli
Taphrognathus bradyi n. gen., n. sp. Welles 1947
recombined as Hadrokkosaurus bradyi
UCMP 36199 (holotype), mandibular ramus
Amphibia - Temnospondyli - Brachyopidae
Vigilius wellesi Warren and Marsicano 2000
Warren and Marsicano 2000 15 specimens
UCMP 36205, 36247, 36836, 37771, 75431, 75432, 75433, 75435, 76477, 76478, 76479, isolated skull fragments; UCMP 36205, mandible; UCMP 80855, atlas; UCMP 36841, clavicle; UCMP 37770, humerus
Amphibia - Temnospondyli - Heylerosauridae
Rhadalognathus boweni n. gen., n. sp. Welles 1947
UCMP 36221, anterior half of left mandibular ramus. Numerous other fragments
Amphibia - Temnospondyli
Eocyclotosaurus wellesi n. sp. Schoch 2000
Schoch 2000 2 specimens
UCMP 42841 (holotype), skull. UCMP 36220, postglenoid area
Cyclotosaurus randalli n. sp. Welles 1947
recombined as Eocyclotosaurus randalli
UCMP 36188 (holotype), skull roof. UCMP 36189, 36190, partial skulls. Numerous additional skull, jaw and skeletal elements
Coelacanthimorpha - Coelacanthiformes - Coelacanthidae
Moenkopia wellesi n. gen., n. sp. Schaeffer and Gregory 1961
Schaeffer and Gregory 1961 13 elements
UCMP 36193, complete basisphenoid (holotype). UCMP 37801, 36234, 36197, basisphenoids. UCMP 56228, parasphenoid. UCMP 37788, 37789, 37790, 37793, pterygoids. UCMP 37791, quadrate. UCMP 56078, ceratohyal. UCMP 73805, 73806, cleithra.
Chondrichthyes - Hybodontiformes - Hybodontidae
Leiacanthus sp. Agassiz 1837
32 elements
synonym of Hybodus
spines
see common names

Geography
Country:United States State/province:Arizona County:Navajo
Coordinates: 34.9° North, 110.3° West (view map)
Paleocoordinates:3.2° North, 38.8° West
Basis of coordinate:stated in text
Geographic resolution:outcrop
Time
Period:Triassic Epoch:Middle Triassic
Stage:Anisian 10 m.y. bin:Triassic 2
Key time interval:Aegean - Bithynian
Age range of interval:247.20000 - 242.00000 m.y. ago
Stratigraphy
Formation:Moenkopi Member:Holbrook
Stratigraphic resolution:bed
Stratigraphy comments: "In the Holbrook Member beneath the 'upper massive sandstone' that caps the Holbrook Member"
Lucas & Schoch (2002: Lethaia 35: 97-106) inferred the Holbrook member to be earliest Anisian (Aegean–Bithynian) in age based upon tetrapod biostratigraphy (Eocyclotosaurus used to correlate Holbrook Member to German Upper Bundsandstein) and magnetostratigraphy
Lithology and environment
Primary lithology:concretionary,nodular,green,red sandy siltstone
Includes fossils?Y
Lithology description: "Channel deposit of greenish, limy silt and sand with included lumps of chocolate shale and small greenish lime nodules with gray centres. Bone as waterworn fragments in matrix"
Environment:fine channel fill
Taphonomy
Modes of preservation:body,charcoalification
Size of fossils:macrofossils,mesofossils
Disassociated major elements:some
Disassociated minor elements:many
Spatial resolution:allochthonous
Collection methods and comments
Collection methods:selective quarrying,surface (in situ),mechanical,field collection
Reason for describing collection:taxonomic analysis
Museum repositories:UCMP
Collectors:S. Welles Collection dates:1939, 1940
Metadata
Also known as:Geronimo Quarry
Database number:34888
Authorizer:J. Alroy, R. Butler Enterer:E. Leckey, R. Butler
Modifier:R. Butler Research group:vertebrate
Created:2003-10-09 14:27:51 Last modified:2012-10-25 03:29:16
Access level:the public Released:2003-10-09 14:27:51
Creative Commons license:CC BY
Reference information

Primary reference:

9311.5% 45940S. P. Welles. 1947. Vertebrates from the Upper Moenkopi Formation of northern Arizona. University of California Publications in Geological Sciences 27(7):241-294 [J. Alroy/E. Leckey/E. Leckey]

Secondary references:

32399 S. G. Lucas and R. R. Schoch. 2002. Triassic temnospondyl biostratigraphy, biochronology and correlation of the German Bundsandstein and North American Moenkopi Formation. Lethaia 35:97-106 [R. Butler/R. Butler]
32383 M. Morales. 1987. Terrestrial fauna and fauna from the Triassic Moenkopi Formation of the southwestern United States. Journal of the Arizona-Nevada Academy of Science 22:1-19 [R. Butler/R. Butler]
32384 S. J. Nesbitt. 2005. The tetrapod fauna of the Moenkopi Formation in northern Arizona. In S. J. Nesbitt, W. G. Parker, R. B. Irmis (eds.), Guidebook to the Triassic Formations of the Colorado Plateau in northern Arizona, Mesa Southwest Museum Bulletin 9:25-32 [R. Butler/R. Butler]
32386 S. J. Nesbitt. 2005. Osteology of the Middle Triassic pseudosuchian archosaur Arizonasaurus babbitti. Historical Biology 17:19-47 [R. Butler/R. Butler]
32378 S. J. Nesbitt. 2005. The Moenkopi Formation along the Little Colorado River in eastern Arizona. In S. J. Nesbitt, W. G. Parker, R. B. Irmis (eds.), Guidebook to the Triassic Formations of the Colorado Plateau in northern Arizona, Mesa Southwest Museum Bulletin 9:13-23 [R. Butler/R. Butler]
32392 S. J. Nesbitt. 2005. A new archosaur from the upper Moenkopi Formation (Middle Triassic) of Arizona and its implications for rauisuchian phylogeny and diversification. Neues Jahrbuch für Geologie und Paläontologie Monatshefte 2005(6):332-346 [R. Butler/R. Butler]
43345 B. Schaeffer and J. T. Gregory. 1961. Coelacanth fishes from the continental Triassic of the western United States. American Museum Novitates 2036:1-18 [R. Butler/R. Butler]
32381 R. R. Schoch. 2000. The status and osteology of two new cyclotosaurid amphibians from the Upper Moenkopi Formation of Arizona (Amphibia: Temnospondyli; Middle Triassic). Neues Jahrbuch für Geologie und Paläontologie Abhandlungen 216(3):387-411 [R. Butler/R. Butler]
32401 A. Warren and C. Marsicano. 2000. A phylogeny of the Brachyopoidea (Temnospondyli, Stereospondyli). Journal of Vertebrate Paleontology 20:462-483 [R. Butler/R. Butler]
32390 S. P. Welles. 1969. Collecting Triassic vertebrates in the Plateau Province. Journal of the West 8(2):231-246 [R. Butler/R. Butler]