Sandelzhausen gravel pit: Langhian, Germany
collected by BSPG 1969-1975, 1994-2001

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Amphibia - Temnospondyli - Pelobatidae
Pelobates fahlbuschi n. sp. Böhme 2010
Böhme 2010 1080 elements
>1026 skull bones/bone fragments, 1 prearticular, 22 ilia, 29 vertebra + 1 urostyle
Eopelobates sp. Parker 1929
Böhme 2010 10 elements
7 skull bones, 3 ilia
Amphibia - Temnospondyli - Ranidae
Ranidae informal indet. nov. gen. et sp. Rafinesque 1814
Böhme 2010 11 elements
1 maxilla fragment, 1 ilium, 9 terminal phalanges
Pelophylax sp. Fitzinger 1843
Böhme 2010 40 elements
28 ilia and many further postcrania
Amphibia - Temnospondyli - Bufonidae
Bufo sp. Laurenti 1768
Böhme 2010 1 fragment
single left ilium fragment
Bufo cf. viridis Laurenti 1768
Böhme 2010 11 elements
1 braincase, 6 ilia, 3 sacral vertebrae + 1 urostyle
Amphibia - Temnospondyli
Latonia gigantea Lartet 1851
Böhme 2010 70 elements
42 maxillae and several other skull bones, 12 ilia and several further postcrania
Amphibia - Temnospondyli - Urodela
Salamandra sansaniensis Lartet 1851
Böhme 2010 95 elements
3 occipital bones, 92 vertebrae
Chelotriton sp. Pomel 1853
Böhme 2010 45 elements
11 skull bones/bone fragments, 6 dentaries, 26 vertebrae, several ribs
Triturus cf. marmoratus (Latreille 1800)
Böhme 2010 210 elements
3 occipital bones, 198 vertebrae, several cranial and postcranial remains
Triturus cf. vulgaris (Linnaeus 1758)
Böhme 2010 290 elements
285 vertebrae, several cranial and postcranial remains
Amphibia - Temnospondyli - Albanerpetontidae
Albanerpeton inexpectatum Estes and Hoffstetter 1976
Böhme 2010 8 elements
4 skull bones, 2 dentaries, 2 vertebrae
Reptilia - Phasianidae
cf. Palaeortyx intermedia Ballman 1966
Gohlich 2003
Reptilia
Diplocynodon cf. gracilis Vaillant 1872
Scherer 1973 408 elements
1 fragment of dentary ("Dentale-Fragment 1" BSPG 1959 II 417)
    = Diplocynodon cf. buetikonensis von Meyer 1855
Scherer 1981
additional specimens (BSPG 1959 II 34, 420 to 426, 436, 437, 680, 681, 2036-2042): 11 skull elements/fragments, 12 jaw fragments, 43 isolated teeth
    = Diplocynodon styriacus Hofmann 1887
Böhme 2010
synonym of Diplocynodon ungeri
"Diplocynodon cf. bueticonensis is a synonym of Diplocynodon styriacus according to Ginsburg & Bulot (1997)"; Böhme (2010) reports further 331 small isolated teeth (BSPG 1959 II 6526, 7539, 16363) which are said to have accumulated in the sedimentary environment due to tooth replacement in juveniles
Diplocynodon sp. Pomel 1847
Scherer 1973 98 elements
2 fragmentary dentaries ("Dentale-Fragment 2" BSPG 1959 II 418, "Dentale-Fragment 3" BSPG 1959 II 419) and 96 isolated teeth (BSPG 1959 II 427 to 435)
    = Diplocynodon cf. buetikonensis von Meyer 1855
Scherer 1981
    = Diplocynodon styriacus Hofmann 1887
Böhme 2010
synonym of Diplocynodon ungeri
Reptilia - Colubridae
"Colubrinae" indet. Oppel 1811
Szyndlar 2009 12 elements
10 trunk vertebrae and 2 maxillary fragments, "tiny"
? Telescopus sp. Waner 1830
Szyndlar 2009 2 elements
2 trunk vertebrae
"Coluber" sp. Linnaeus 1758
Szyndlar 2009 66 elements
65 trunk vertebrae + 1 maxillary fragment; the quotation marks are due to the fact that the author consciously uses the somewhat older concept of the genus (extant "colubrine" snakes by now assigned to the genera Hemorrhois, Hierophis, and Platyceps)
"Natricinae" indet. Bonaparte 1838
Szyndlar 2009 50 elements
50 trunk vertebrae
cf. Neonatrix sp. Holman 1973
Szyndlar 2009 4 elements
4 trunk vertebrae
Natrix sp. Laurenti 1768
Szyndlar 2009 32 elements
32 trunk vertebrae
cf. Natrix sp. Laurenti 1768
Szyndlar 2009 4 elements
4 trunk vertebrae, "large form"
Reptilia - Crotalidae
Vipera sp. Laurenti 1768
Szyndlar 2009 70 elements
"Oriental viper" (contrastingly, Szyndlar & Rage, 2002, ref no. 43793, list the Vipera sp. from Sandelzhausen as a member of the "aspis complex"), 19 vertebrae and >50 isolated venom fangs
Reptilia - Elapidae
Elapidae indet. Boié 1827
Szyndlar 2009 6 elements
6 trunk vertebrae
Naja sp. Laurenti 1768
Szyndlar 2009 6 elements
6 trunk vertebrae
Reptilia
Eoanilius sp. Rage 1974
Szyndlar 2009 10 elements
10 trunk vertebrae
Reptilia - Cordylidae
Cordylidae indet. Gray 1837
Böhme 2010 2 fragments
2 jaw fragments
Bavaricordylus cf. molassicus Böhme 2010
Böhme 2010 6 fragments
2 maxillary fragments, 2 dentary fragments, 2 isolated dentary teeth
Reptilia - Scincidae
Scincidae informal indet. 1 Gray 1825
Böhme 2010 2 fragments
2 dentary fragments
Scincidae informal indet. 2 Gray 1825
Böhme 2010 2 fragments
2 dentary fragments
Tropidophorus bavaricus n. sp. Böhme 2010
Böhme 2010 4 specimens
1 dentary with 12 tooth positions (= holotype, BSPG 1959 II 16395), 3 dentary fragments
Reptilia
Palaeoblanus informal sp. nov. Schleich 1988
Böhme 2010 2 specimens
1 intermaxilla, 1 posterior dentary fragment
Palaeoblanus tobieni Schleich 1988
Böhme 1999
Reptilia - Lacertidae
Lacerta informal sp. 1 Linnaeus 1758
Böhme 2010 1 element
"Lacerta sensu lato", left dentary
Lacerta informal sp. 2 Linnaeus 1758
Böhme 2010 10 specimens
"Lacerta sensu lato", 1 dentary + 7 fragments, 1 maxilla + 1 fragment
Lacerta informal sp. 3 Linnaeus 1758
Böhme 2010 21 specimens
"Lacerta sensu lato", 1 dentary + 14 fragments + 2 jaw fragments, 1 maxilla + 3 fragments
Edlartetia sansaniensis Augé and Rage 2000
Böhme 2010 1 fragment
1 dentary fragment
cf. Miolacerta tenuis Roček 1984
Böhme 2010 2 fragments
2 dentary fragments
Reptilia
Scincomorpha indet. (Camp 1923)
Böhme 2010 6 specimens
2 intermaxillae, 4 jaw fragments
Reptilia - Anguidae
Anguidae informal indet. sp. nov. Gray 1825
Böhme 2010 3 fragments
2 dentary fragments and 1 maxilla fragment
Ophisaurus sp. Daudin 1803
Böhme 2010 40 specimens
13 skull bones/bone fragments, 10 dentaries/fragments + 2 further jaw fragments, 4 vertebrae, several osteoderms
Ophisaurus informal sp. I sensu Roček, 1984 Daudin 1803
Böhme 2010 10 fragments
4(?) dentary fragments, 6 maxilla fragments
cf. Ophisaurus cf. spinari Klembara 1979
Böhme 2010 3 fragments
"sensu Roček, 1984"; 3 maxilla fragments
Pseudopus laurillardi (Lartet 1851)
Böhme 2010 25 elements
5 skull bones, 4 dentaries/fragments, 7 vertebrae, 1 isolated tooth, several osteoderms
Reptilia - Chamaeleonidae
Chamaeleo bavaricus n. sp. Schleich 1983
Schleich 1983 4 fragments
1 dentary fragment (=holotype, BSPG 1959 II 2066), 3 maxilla fragments (reported by Böhme, 2010)
Chamaeleo aff. caroliquarti Moody and Roček 1980
Böhme 2010 5 elements
nomen dubium belonging to Chamaeleo
5 skull bones/fragments
Reptilia - Testudines - Testudinidae
Geochelone cf. perpiniana
Schleich 1984
    = Titanochelon schleichi n. sp. Pappa et al. 2023
Pappa et al. 2023
SNSB-BSPG 1959 II 2033, a partial skeleton comprising most of the carapace, the plastron, and some appendicu- lar elements (scapula, humerus, radius, ulna, carpal ele- ments, tibia, fibula, tarsal elements), mounted in anatomical position.
Testudo rectogularis
Schleich 1984
Reptilia - Testudines - Geoemydidae
Ocadia sophiae (Ammon 1911)
Hervet 2004
recombined as Mauremys sophiae
Clemmydopsis turnauensis (Meyer 1847)
Schleich 1984
Reptilia - Testudines - Trionychidae
Trionyx aff. bohemicus Liebus 1930
Schleich 1984
    = Trionychidae indet. Gray 1825
Georgalis and Joyce 2017
Schleich 1984
    = Trionychidae indet. Gray 1825
Georgalis and Joyce 2017
Mammalia - Proboscidea - Mammutidae
Zygolophodon turicensis (Schintz 1824)
Mammalia - Proboscidea - Gomphotheriidae
Gomphotherium angustidens Burmeister 1837
subspecies G. a. subtapiroides
Mammalia - Talpidae
Myxomygale hutchisoni (Ziegler 1985)
Hugueney and Maridet 2017
Mammalia - Rodentia - Cricetidae
Democricetodon aff. schaubi
Fahlbusch 1964 15 elements
n. sp., 15 teeth
Democricetodon minor (Lartet 1851)
Fahlbusch 1964 15 elements
n.ssp. gracilis, 15 teeth
Democricetodon affinis Schaub 1925
Fahlbusch 1964 40 elements
n.ssp. mutilus, 40 teeth
Cotimus bifidus n. sp. Fahlbusch 1964
Fahlbusch 1964 21 elements
recombined as Eumyarion bifidus
21 teeth
Actinopteri - Gobiiformes - Gobiidae
Gobius sp. Linnaeus 1758
Böhme 2010 3 elements
1 premaxilla, 2 jaw elements
Actinopteri - Anabantiformes - Channidae
Channa elliptica von Salis 1967
Böhme 2010 150 elements
68 vertebrae, 5 pharyngobranchial fragments, 1 premaxilla, 1 dentary fragment, >70 isolated teeth
Actinopteri - Cypriniformes - Cyprinidae
Palaeoleuciscus informal sp. A Obrhelová 1969
Böhme 2010 33 elements
= P. sp. A sensu Böhme (2002, Beitr. Paläont. 27); 9 isolated pharyngeal teeth, 17 fragments of pharyngeal bone with attached teeth, 1 operculum fragment, 6 first vertebrae
Palaeocarassius informal sp. A Obrhelová 1970
Böhme 2010 1755 elements
1659 isolated pharyngeal teeth, 1 pharyngeal bone, 13 serrated fin spines, 85 first vertebrae
Gastropoda - Heterostropha - Carychiidae
Carychium (Carychium) galli n. sp. Salvador 2015
Salvador 2015
Carychium nouleti nouleti and gibbum in Gall (1972); C. sp. 1 and 2 in Moser et al. (2009)
Carychium (Carychiella) eumicron (Bourguignat 1857)
Salvador 2015
Carychium eumicron peneckei in Gall (1972); Carychium sp. 3 in Moser et al. (2009)
Gastropoda - Heterostropha - Testacellidae
Testacella zellii Klein 1853
Salvador 2013
Testacella larteti larteti in Gall (1972); Testacella sp. in Moser et al. (2009)
Gastropoda - Heterostropha - Limacidae
Limax sp. Linnaeus 1758
Salvador and Rasser 2014
Gastropoda - Heterostropha - Vitrinidae
Vitrina sp. Draparnaud 1801
Salvador and Rasser 2014
Gastropoda - Heterostropha - Gastrodontidae
Janulus supracostatus (Sandberger 1873)
Salvador and Rasser 2014
Gastropoda - Heterostropha - Helicidae
Helicidae indet. Rafinesque 1815
Salvador 2013
Cepaea cf. eversa (Deshayes 1851)
Salvador 2013
Cepaea cf. sylvestrina (Schlotheim 1820)
Salvador 2013
Gastropoda - Heterostropha - Hygromiidae
Leucochroopsis kleinii (Klein 1847)
Salvador 2013
Gastropoda - Heterostropha - Elonidae
Tropidomphalus cf. incrassatus (Klein 1853)
Salvador 2013
recombined as Pseudochloritis incrassata
Gastropoda - Heterostropha - Helicodiscidae
Lucilla subteres (Clessin 1877)
Salvador and Rasser 2014
Gastropoda - Heterostropha - Vertiginidae
Vertigo callosa (Reuss 1849)
Salvador 2015
recombined as Vertigo (Vertigo) callosa
Gastropoda - Heterostropha - Gastrocoptidae
Gastrocopta nouletiana (Dupuy 1850)
Salvador 2015
Gastrocopta acuminata (Klein 1846)
Salvador 2015
Gastropoda - Heterostropha - Chondrinidae
Granaria sp. Held 1837
Salvador 2015
Napaeus? sp. (in part) in Gall (1972); Granaria sp. 2 and ?Chondrinidae/?Enidae in Moser et al. (2009)
Granaria cf. grossecostata (Gottschick and Wenz 1919)
Salvador 2015
Napaeus? sp. (in part) in Gall (1972); Granaria sp. 1 in Moser et al. (2009)
Gastropoda - Heterostropha - Pyramidulidae
Pyramidula sp. Fitzinger 1833
Salvador 2015
Trunactellina sp. in Moser et al. (2009)
Gastropoda - Heterostropha - Strobilopsidae
Strobilops sp. Pilsbry 1893
Salvador 2015
Discus (Discus) pleuradra in Gall (1972); Strobilops (Strobilops) and S. (Discostrobilops) sp. in Moser et al. (2009)
Gastropoda - Heterostropha - Filholiidae
Triptychia sp. Sandberger 1876
Salvador 2015
Gastropoda - Heterostropha - Clausiliidae
Pseudidyla moersingensis (Boettger 1877)
Salvador 2015
Clausiliidae indet 1 and 2 in Moser et al. (2009)
Gastropoda - Heterostropha - Spiraxidae
Palaeoglandina sp. Wenz 1914
Salvador 2013
Palaeoglandina gracilis porrecta in Gall (1972)
Gastropoda - Heterostropha - Lymnaeidae
Lymnaea dilatata (Noulet 1854)
Salvador and Rasser 2014
Galba dupuyiana (Noulet 1845)
Salvador and Rasser 2014
Gastropoda - Heterostropha - Planorbidae
Gyraulus dealbatus (Braun 1851)
Salvador and Rasser 2014
synonym of Gyraulus applanatus
Gyraulus albertanus (Clessin 1877)
Salvador and Rasser 2014
Planorbarius cornu (Brongniart 1810)
Salvador and Rasser 2014
Ferrissia deperdita (Desmarest 1814)
Salvador and Rasser 2014
Gastropoda - Heterostropha - Valvatidae
Valvata sp. Müller 1774
Salvador 2013
Gastropoda - Discidae
Discus pleuradrus (Bourguignat 1881)
Salvador and Rasser 2014
Gastropoda - Neritidae
Theodoxus sp. Monfort 1810
Salvador 2013
Gastropoda - Pomatiidae
Pomatias sp. Stimpson 1860
Salvador 2013
Bivalvia - Cardiida - Sphaeriidae
Sphaerium sp. Scopoli 1777
Salvador 2013
see common names

Geography
Country:Germany State/province:Bavaria County:Kelheim
Coordinates: 48.6° North, 11.8° East (view map)
Paleocoordinates:48.9° North, 10.4° East
Basis of coordinate:stated in text
Altitude:450 meters
Geographic resolution:outcrop
Time
Period:Neogene Epoch:Miocene
Stage:Langhian 10 m.y. bin:Cenozoic 5
Key time interval:Langhian Mammal zone: OSM C
Age range of interval:15.98000 - 13.82000 m.y. ago
Age estimate:16 Ma (graphic correlation)
Stratigraphy
Geological group:Molasse Formation:Upper Freshwater Molasse
Stratigraphic resolution:group of beds
Stratigraphy comments: The fossiliferous bed, which has a mean thickness of about 2.5 m, belongs to the Upper Freshwater Molasse (OSM) informal formation. Its age assigment has slightly changed several times over the past decades. Currently it is regarded as belonging to the OSM unit C (i.e. sedimentary cycle OSM 3), corresponding to the lower part of the Middle Miocene. Studies on several mammal groups have revealed that the locality biochronologically now is unequivocally referrable to the deeper part of MN 5 and not, as earlier thought, to MN6. The absolute age of the locality is about 16 Ma (see Fahlbusch, 2003 and references therein).
The fossiliferous bed itself is subdivided into 6 (sub)layers (A to F from base to top). However, in almost all cases no particular horizon is specified for a particular fossil or taxon in the paleontological literature.
Lithology and environment
Primary lithology:shelly/skeletal,black,brown,gray,green sandy,conglomeratic marl
Secondary lithology: unlithified silty mudstone
Includes fossils?Y
Includes fossils?Y
Lithology description: The base of the fossiliferous bed rests on a paleorelief of about 30 cm. The (sub)layers A to D are composed of greenish-gray to brownish-black, marly conglomerates ("Geröllmergel") with decrease in quantity and size of pebbles from max. 10 cm in the lower part to few mm in the upper part of the fossiliferous bed. (Sub)layer A is partially well cemented by carbonate minerals. The pebbles mainly consist of quartz but also of various crystalline rock and hornfels, and some are flattened limestone pebbles. (Sub)layer C contains a thin lignite lens which, together with its lateral equivalents, served as guide bed dividing the "Untere Geröllmergel" below from the "Obere Geröllmergel" above. The (sub)layers A to D are roughly distinguishable from one another throughout the whole gravel pit, however, on a smaller scale rapid (often within few meters) facies changes occur. (Sub)layer E is a montmorillonite-rich bed of silty clay. The (sub)layers A to E all contain fossils: predominantly vertebrates and gastropod shell debris. (Sub)layer F is devoid of fossils. (Sub)layer B partially is characterized by structures which indicate synsedimentary movements such as slumps, but it cannot be ruled out that they may result from bioturbation. Almost all (sub)horizons of the fossiliferous bed are characterized by the presence of light grey to yellowish-white limestone concretions of various shape and size. These are the result of different diagenetic phenomena (e.g. subrecent precipitation from descendent meteoric water or early diagenetic processes related to fossil formation) (all informations on geology are from Fahlbusch 2003 and Böhme, 2010).
Environment:fluvial-lacustrine indet. Tectonic setting:foreland basin
Geology comments: (Sub)bed A is interpreted as a gravel bed channel. (Sub)beds B to D (lower) may represent an alluvial plain in which, after seasonal floods, small, short-lived ponds existed under semiarid to semihumid conditions. (Sub)beds D (upper) and E are interpreted as permanent water body, a small lake or pond, under semihumid to humid conditions (Böhme, 2010).
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils,mesofossils
Spatial orientation:random
Preservation of anatomical detail:variable
Abundance in sediment:abundant
Disassociated minor elements:all
Size sorting:well
Fragmentation:frequent
Temporal resolution:time-averaged
Collection methods and comments
Collection excludes:some macrofossils
Collection methods:bulk,surface (float),mechanical,acetic,sieve,survey of museum collection
Rock censused:15000000 cm3 (volume)
Reason for describing collection:taxonomic analysis
Museum repositories:BSP
Collectors:BSPG Collection dates:1969-1975, 1994-2001
Collection method comments: The fossil locality was discovered in 1959 by Richard Dehm (director of the BSPG at the time). After getting positive results from samples taken in 1962 systematic excavations including screenwashing took place from 1969 to 1975 (several weeks per year). In 1994 a second excavation period started which ended in 2001. During both periods a total of 1500 m3 has been quarried, accompanied by 150 t of screenwashed sediment (Fahlbusch, 2003). It should be noted, that amount of screenwashed samples varies between the (sub)layers and cannot be reconstructed (Böhme, 2010).
Almost all of the material is housed in the BSPG, bearing the catalogue signature BSPG 1959 II.
Taxonomic list comments:Szyndlar (2009) lists 130 caudal vertebrae, 2 dentary fragments, and 1 unidentified dentigerous bone as Serpentes indet.
Note: Occurence of Eumyarion bifidus reported by Heißig (1989) has been removed from the list due to duplicate (alternative combination of Cotimus bifidus). (TL)
Metadata
Also known as:Sandelzhausen 1, Kiesgrube Bachmaier-Karber; Sandelshausen
Database number:34615
Authorizer:J. Hunter, J. Mueller, M. Clapham, E. Vlachos, T. Cleary, P. Mannion, P. Holroyd, D. Nicholson Enterer:M. Hall, E. Vlachos, M. Clapham, T. Cleary, P. Mannion, P. Holroyd, D. Nicholson, T. Liebrecht
Modifier:G. Varnham Research group:vertebrate
Created:2003-09-12 07:54:45 Last modified:2022-02-11 10:09:19
Access level:the public Released:2003-09-12 07:54:45
Creative Commons license:CC BY
Reference information

Primary reference:
9144.5% 38460N. Schmidt-Kittler. 1972. Die obermiozäne Fossil-Lagerstätte Sandelzhausen. 6. Proboscidea (Mammalia). Mitteilungen der Bayerischen Staatssammlung für Paläontologie und Histor. Geologie 12:83-95 [J. Hunter/M. Hall/T. Liebrecht]

Secondary references:

67837 M. Böhme. 1999. Doppelschleichen (Sauria, Amphisbaenia) aus dem Untermiozän von Stubersheim 3 (Süddeutschland). Mitteilungen der Bayerischen Staatssammlung für Paläontologie und historischen Geologie 39:85-90 [T. Cleary/T. Cleary]
43888 M. Böhme. 2010. Ectothermic vertebrates (Actinopterygii, Allocaudata, Urodela, Anura, Crocodylia, Squamata) from the Miocene of Sandelzhausen (Germany, Bavaria) and their implications for environment reconstruction and palaeoclimate. Paläontologische Zeitschrift 84(1):3-41 [J. Mueller/T. Liebrecht]
12913 V. Fahlbusch. 1964. Die Cricetiden (Mamm.) der Oberen Süßwasser-Molasse Bayerns (Cricetids of the Upper Freshwater Molasse of Bavaria). Bayerische Akademie der Wissenschaften; mathematisch-naturwissenschaftliche Klasse; Abhandlungen, neue Folge 118:1-136 [J. Alroy/S. Kuemmell/S. Kuemmell]
52039 J. D. Gardner and M. Böhme. 2008. Review of the Albanerpetontidae (Lissamphibia), with comments on the paleoecological preferences of European Tertiary albanerpetontids. In J. T. Sankey & S. Baszio (ed.), Vertebrate Microfossil Assemblages: Their Role in Paleoecology and Paleobiogeography 178-218 [M. Carrano/M. Carrano]
77386 J. D. Gardner, A. Villa, S. Colombero, M. Venczel, and M. Delfino. 2021. A Messinian (latest Miocene) occurrence for Albanerpeton Estes & Hoffstetter, 1976 (Lissamphibia: Albanerpetontidae) at Moncucco Torinese, Piedmont Basin, northwestern Italy, and a review of the European Cenozoic record for albanerpetontids. Geodiversitas 43(14):391-404 [M. Carrano/M. Carrano]
62147 G. L. Georgalis and W. G. Joyce. 2017. A Review of the Fossil Record of Old World Turtles of the Clade Pan-Trionychidae. Bulletin of the Peabody Museum of Natural History 58(1):115-208 [P. Holroyd/P. Holroyd/P. Mannion]
40668 U. B. Gohlich. 2003. The avifauna of the Grund Beds (Middle Miocene, Early Badenian, northern Austria). Annalen des Naturhistorischen Museums in Wien, Serie A 104:237-249 [M. Uhen/M. Uhen/T. Liebrecht]
58591 S. Hervet. 2004. Systématique du groupe "Palaeochelys sensu lato – Mauremys" (Chelonii, Testudinoidea) du Tertiaire d'Europe occidentale: principaux résultats [Systematic of the "Palaeochelys sensu lato – Mauremys" group (Chelonii, Testudinoidea) from the Tertiary of Western Europe: principal results. Annales de Paléontologie 90(1):13-78 [D. Nicholson/D. Nicholson/D. Nicholson]
61807 M. Hugueney and O. Maridet. 2017. Evolution of Oligo-Miocene talpids (Mammalia, Talpidae) in Europe: focus on the genera Myxomygale and Percymygale n. gen. Historical Biology [P. Mannion/P. Mannion]
70379 M. Ivanov. 2001. Changes in the composition of the European snake fauna during the Early Miocene and at the Early/Middle Miocene transition. Palaeontologische Zeitschrift 74(4):563-573 [T. Cleary/T. Cleary]
85112 I. Pappa, E. Vlachos, and M. Moser. 2023. A new species of a giant tortoise from Sandelzhausen (MN5, Burdigalian/Langhian boundary, Early/Middle Miocene, South Germany). The Anatomical Record 1-14 [E. Vlachos/E. Vlachos]
85570 R. B. Salvador. 2013. The fossil pulmonate snails of Sandelzhausen (Early/Middle Miocene, Germany): Succineidae, Testacelloidea and Helicoidea. Zootaxa 3721:157-171 [M. Clapham/M. Clapham]
85582 R. B. Salvador. 2013. The fossil land and freshwater mollusks of Sandelzhausen (Early/Middle Miocene, Germany): Caenogastropoda, Neritimorpha, lower Heterobranchia and Bivalvia. Strombus 20:19-26 [M. Clapham/M. Clapham]
85569 R. B. Salvador. 2015. The fossil pulmonate snails of Sandelzhausen (Early/Middle Miocene, Germany): Ellobiidae, Pupilloidea, and Clausilioidea. Paläontologische Zeitschrift 89:37-50 [M. Clapham/M. Clapham]
85571 R. B. Salvador and M. W. Rasser. 2014. The fossil pulmonate snails of Sandelzhausen (Early/Middle Miocene, Germany) (Hygrophila, Punctoidea and limacoids). Archiv für Molluskenkunde 143:187-202 [M. Clapham/M. Clapham]
43901 E. Scherer. 1973. Die obermiozäne Fossil-Lagerstätte Sandelzhausen. 9. Crocodilia. Mitteilungen der Bayerischen Staatssammlung für Paläontologie und Historische Geologie 13:103-114 [J. Mueller/T. Liebrecht]
43902 E. Scherer. 1981. Die mittelmiozäne Fossil-Lagerstätte Sandelzhausen. 12. Crocodylia (abschließender Bericht). Mitteilungen der Bayerischen Staatssammlung für Paläontologie und Historische Geologie 21:81-87 [J. Mueller/T. Liebrecht]
43939 H.-H. Schleich. 1983. Die mittelmiozäne Fossil-Lagerstätte Sandelzhausen. 13. Chamaeleo bavaricus sp. nov., ein neuer Nachweis aus dem Jungtertiär Süddeutschlands. Mitteilungen der Bayerischen Staatssammlung für Paläontologie und Historische Geologie 23:77-81 [J. Mueller/T. Liebrecht/T. Liebrecht]
66150 H.H. Schleich. 1984. Neogene Testudines of Germany. Their stratigraphical and ecological evaluation. Studia Geologica Salmanticensia (Studia Palaeocheloniologica 1), volumen especial 1:249-267 [E. Vlachos/E. Vlachos]
43680 Z. Szyndlar. 2009. Snake fauna (Reptilia: Serpentes) from the Early/Middle Miocene of Sandelzhausen and Rothenstein 13 (Germany). Paläontologische Zeitschrift 83(1):55-66 [J. Mueller/T. Liebrecht]