Maramena: Turolian, Greece

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
cf. Belgrandiella indet.
Schmidt-Kittler et al. 1995
Mastus sp.
Schmidt-Kittler et al. 1995
Milax sp.
Schmidt-Kittler et al. 1995
or Tandonia sp.
Cadona sp.
Schmidt-Kittler et al. 1995
Ouzoceros aff. gracilis
Schmidt-Kittler et al. 1995
Norbertia hellenica n. gen., n. sp.
Schmidt-Kittler et al. 1995
Ommatotriton sp.
Georgalis et al. 2019
M1: a vertebra (UU MAA 7489) and two femurs (UU MAA 7248 and UU MAA 7509).
Gastropoda - Aciculidae
cf. Platyla sp. Moquin-Tandon 1855
Schmidt-Kittler et al. 1995
Gastropoda - Bithyniidae
Bithynia sp. Leach 1818
Schmidt-Kittler et al. 1995
Gastropoda - Heterostropha - Planorbidae
Gyraulus sp. Charpentier 1837
Schmidt-Kittler et al. 1995
Planorbarius sp. Dumeril 1806
Schmidt-Kittler et al. 1995
cf. conru-mantelli
Planorbis sp. Muller 1774
Schmidt-Kittler et al. 1995
Gastropoda - Heterostropha - Carychiidae
Carychium cf. nouleti Bourguignat 1857
Schmidt-Kittler et al. 1995
Gastropoda - Heterostropha - Cochlicopidae
Cochlicopa sp. Ferussac 1821
Schmidt-Kittler et al. 1995
Gastropoda - Heterostropha - Parmacellidae
Parmacella sp. Cuvier 1804
Schmidt-Kittler et al. 1995
nov
Gastropoda - Heterostropha - Limacidae
cf. Limax sp. Linnaeus 1758
Schmidt-Kittler et al. 1995
Gastropoda - Heterostropha - Pristilomatidae
Vitrea sp. Fitzinger 1833
Schmidt-Kittler et al. 1995
Gastropoda - Heterostropha - Clausiliidae
Clausiliidae indet. Gray 1855
Schmidt-Kittler et al. 1995
Gastropoda - Heterostropha
Helicoidea indet. Rafinesque 1815
Schmidt-Kittler et al. 1995
sp. nov.
Gastropoda - Heterostropha - Elonidae
Klikia goniostoma (Sandberger 1875)
Schmidt-Kittler et al. 1995
Bivalvia - Cardiida - Sphaeriidae
Pisidium casertanum (Poli 1791)
Schmidt-Kittler et al. 1995
original and current combination Cardium casertanum
Amphibia - Temnospondyli - Ranidae
Ranidae indet. Rafinesque 1814
Georgalis et al. 2019
M2: three ilia (UU MAA 7195); M3: three humeri (UU MAA 7231).
Pelophylax sp. Fitzinger 1843
Georgalis et al. 2019
M2: an angular (UU MAA 7194); M3: a coracoid (UU MAA 7233) and three ilia (UU MAA 7232).
Amphibia - Temnospondyli - Hylidae
Hyla sp. Laurenti 1768
Georgalis et al. 2019
M1: seven humeri (UU MAA 7119, UU MAA 7244, UU MAA 7251, and UU MAA 7474-UU MAA 7476) and an ilium (UU MAA 7118); M2: three humeri (UU MAA 7193).
Amphibia - Temnospondyli
Latonia sp. von Meyer 1843
Georgalis et al. 2019
M1: two fragmentary frontoparietals belonging to a single individual (UU MAA 7445 and UU MAA 7462), three premaxillae (UU MAA 7130), 66 maxillae (UU MAA 7120, UU MAA 7128, UU MAA 7131, UU MAA 7156, UU MAA 7249, UU MAA 7250, UU MAA 7458, and UU MAA 7464-UU MAA 7472), seven angulars (UU MAA 7154 and UU MAA 7529), an atlas (UU MAA 7459), 17 presacral vertebrae (UU MAA 7125, UU MAA 7158a, UU MAA 7442-UU MAA 7444, UU MAA 7452, UU MAA 7454, UU MAA 7457, UU MAA 7460, and UU MAA 7461), three sacral vertebrae (UU MAA 7455 and UU MAA 7522), four urostyles (UU MAA 7446, UU MAA 7450, and UU MAA 7521), a scapula (UU MAA 7520), four costae (UU MAA 7523), 14 humeri (UU MAA 7155, UU MAA 7158b, UU MAA 7453, and UU MAA 7463), and nine ilia (UU MAA 7447-UU MAA 7449, UU MAA 7456, and UU MAA 7528); M2: a fragmentary frontoparietal (UU MAA 7192); M3: seven maxillae (UU MAA 7223, UU MAA 7224, UU MAA 7227, UU MAA 7238, UU MAA 7239, UU MAA 7240, and UU MAA 7241), a presacral vertebra (UU MAA 7228), a humerus (UU MAA 7225), and an ilium (UU MAA 7229).
Amphibia - Temnospondyli - Urodela
Cryptobranchidae indet. Cope 1889
Georgalis et al. 2019
M1: an atlas (UU MAA 7441), probably also a second atlas (UU MAA 7530)
Salamandra sp. Laurenti 1768
Georgalis et al. 2019
M1: 12 trunk vertebrae (UU MAA 7494- UU MAA 7505) and six caudal vertebrae (UU MAA 7117, UU MAA 7491-UU MAA 7493, UU MAA 7506, and UU MAA 7507).
Salamandridae indet. Gray 1825
Georgalis et al. 2019
M1: an atlas (UU MAA 7490), four vertebrae (UU MAA 7153, UU MAA 7485, UU MAA 7488, and UU MAA 7243), and two ribs (UU MAA 7508 and UU MAA 7519).
Salamandrina sp. Fitzinger 1826
Georgalis et al. 2019
M1: two vertebrae (UU MAA 7483 and UU MAA 7484)
Lissotriton sp. Bell 1839
Georgalis et al. 2019
(Lissotriton vulgaris group) M1: two vertebrae (UU MAA 7517 and UU MAA 7518).
Mammalia - Rodentia - Gliridae
Myomimus maritsensis de Bruijn et al. 1970
70 specimens
approx.
Myomimus dehmi (de Bruijn 1966)
Schmidt-Kittler et al. 1995
Glis cf. minor Kowalski 1963
Schmidt-Kittler et al. 1995
Muscardinus pliocaenicus Kowalski 1963
Schmidt-Kittler et al. 1995
Mammalia - Rodentia - Sciuridae
Tamias informal sp. A Illinger 1811
Tamias + Protospermophilus=approx. 13 specimens
Hylopetes macedoniensis
Pliopetaurista + Hylopetes=approx. 50 specimens
? Protospermophilus sp. Gazin 1930
Tamias + Protospermophilus=approx. 13 specimens
Pliopetaurista dehneli
Pliopetaurista + Hylopetes=approx. 50 specimens
Miopetaurista thaleri Mein 1970
Schmidt-Kittler et al. 1995
Mammalia - Rodentia - Eomyidae
Keramidomys aff. carpathicus (Schaub and Zapfe 1953)
probably less than 3 specimens
Leptodontomys catalaunicus
Schmidt-Kittler et al. 1995
Mammalia - Rodentia - Castoridae
Castoridae indet. Gray 1821
probably less than 3 specimens
Mammalia - Rodentia - Muridae
Apodemus dominans
A. gudrunae+A.dominans=approx. 390 specimens
Apodemus gudrunae van de Weerd 1976
A. gudrunae+A.dominans=approx. 390 specimens
Micromys informal sp. A Dehne 1841
18 specimens
approx.
Occitanomys cf. neutrum de Bruijn 1976
40 specimens
approx.
Mammalia - Rodentia - Cricetidae
Kowalskia fahlbuschi Bachmayer and Wilson 1970
recombined as Neocricetodon fahlbuschi
Kowalskia + Cricetus= approx. 83 specimens
"Cricetus" informal sp. A Leske 1779
Kowalskia + Cricetus= approx. 83 specimens
Mammalia - Lagomorpha - Leporidae
Alilepus turolensis
Alilepus + Prolagus = 41 specimens
Mammalia - Lagomorpha - Ochotonidae
Prolagus michauxi
Alilepus + Prolagus = 41 specimens
Mammalia - Primates - Cercopithecidae
Mesopithecus pentelicus Wagner 1839
Schmidt-Kittler et al. 1995
Mammalia - Erinaceidae
Schizogalerix sp. Engesser 1980
Doukas 1989
    = Schizogalerix macedonica n. sp. Doukas et al. 1995
Doukas et al. 1995
Erinaceus sp. Linnaeus 1758
Doukas 1989
    = Erinaceus samsonowiczi Sulimski 1959
Doukas et al. 1995
Erinaceus samsonowiczi Sulimski 1959
Doukas 2005
Mammalia - Soricidae
Soricidae informal sp. A Fischer 1814
Doukas 1989
    = Asoriculus gibberodon
Doukas et al. 1995
Petenyia sp. Kormos 1934
Doukas 1989
    = Petenyia hungarica Kormos 1934
Doukas et al. 1995
Deinsdorfia kerkhoffi n. sp. Doukas et al. 1995
Doukas et al. 1995
Soricidae informal sp. B Fischer 1814
Doukas 1989
    = Sorex sp. Linnaeus 1758
Doukas et al. 1995
Amblycoptus sp. Kormos 1926
Doukas 1989
    = Amblycoptus jessiae n. sp. Doukas et al. 1995
Doukas et al. 1995
Mammalia - Talpidae
Desmanella sp. Engesser 1972
Doukas 1989
    = Desmanella dubia Rümke 1976
Doukas et al. 1995
Talpidae informal sp. A (Fischer de Waldheim 1817)
Doukas 1989
    = Urotrichus sp. Temminck 1841
Doukas et al. 1995
Dibolia dekkersi Rümke 1985
Rümke 1985
    = Ruemkelia dekkersi
Doukas et al. 1995
synonym of Archaeodesmana
Talpidae informal sp. B (Fischer de Waldheim 1817)
Doukas 1989
    = Talpa fossilis Petenyi 1864
Doukas et al. 1995
Mammalia - Chiroptera
Chiroptera indet. Blumenbach 1779
Schmidt-Kittler et al. 1995
Mammalia - Perissodactyla - Rhinocerotidae
Rhinocerotidae indet. Gray 1821
Schmidt-Kittler et al. 1995
Mammalia - Perissodactyla - Equidae
Hipparion sp. de Christol 1832
Schmidt-Kittler et al. 1995
2 species
Mammalia - Suidae
Korynochoerus palaeochoerus
Schmidt-Kittler et al. 1995
Mammalia - Cervidae
Pliocervus graecus n. sp.
Schmidt-Kittler et al. 1995
Mammalia - Bovidae
Gazella sp. Blaineville 1816
Schmidt-Kittler et al. 1995
Boselaphini indet. Knottnerus-Meyer 1997
Schmidt-Kittler et al. 1995
Tragoportax gaudryi Kretzoi 1941
Schmidt-Kittler et al. 1995
Tragoportax cf. amalthea (Roth and Wagner 1854)
Schmidt-Kittler et al. 1995
Mammalia - Giraffidae
Samotherium cf. boissieri Forsyth Major 1888
Schmidt-Kittler et al. 1995
Mammalia - Carnivora - Hyaenidae
Chasmaporthetes sp. Hay 1921
Schmidt-Kittler et al. 1995
Mammalia - Carnivora - Viverridae
Viverridae indet. Gray 1821
Schmidt-Kittler et al. 1995
Mammalia - Carnivora - Mephitidae
Promephitis sp. Gaudry 1861
Schmidt-Kittler et al. 1995
Mammalia - Carnivora - Mustelidae
Martes lefkonensis n. sp.
Schmidt-Kittler et al. 1995
Lutra affinis Gervais 1859
Schmidt-Kittler et al. 1995
Promeles macedonicus
Schmidt-Kittler et al. 1995
Mammalia - Proboscidea - Choerolophodontidae
Choerolophodon pentelicus
Schmidt-Kittler et al. 1995
Reptilia - Testudines
Testudines indet. Batsch 1788
Georgalis et al. 2019
M1: an incomplete unidentified neural (UU MAA 7136), three unidentified peripherals (UU MAA 7137, UU MAA 7140, and UU MAA 7142), and several shell fragments (UU MAA 7138, UU MAA 7139, UU MAA 7147 [five fragments]); M3: an unidentified peripheral (UU MAA 7254), a shell fragment (UU MAA 7252), and several fragments of different elements (one caudal vertebra, one pelvic fragment, three neurals, four costals, one peripheral, and 24 shell fragments) (UU MAA 7255).
Reptilia - Testudines - Testudinidae
? Testudinidae indet. Batsch 1788
Georgalis et al. 2019
M1: an isolated ?limb osteoderm (UU MAA 7135).
Testudinata indet. (Oppel 1811)
Schmidt-Kittler et al. 1995
    = Testudo sp. Linnaeus 1758
Vlachos 2015
isolated plates
Reptilia - Testudines - Geoemydidae
Mauremys aristotelica Vlachos et al. 2019
Georgalis et al. 2019
M1: three incomplete unidentified neurals (UU MAA 7143, UU MAA 7144, and UU MAA 7146); M3: a nuchal (UU MAA 7283), a fragmentary unidentified neural (UU MAA 7253), a left costal VI (UU MAA 7284), unidentified costals (UU MAA 7286 and UU MAA 7288), a left peripheral III (UU MAA 7289), a right peripheral III (UU MAA 7290), unidentified peripherals (UU MAA 7285, UU MAA 7287, UU MAA 7291, UU MAA 7292, UU MAA 7293, UU MAA 7295, and UU MAA 7297), and a right hyoplastron (UU MAA 7296).
Reptilia - Squamata
Squamata indet. Oppel 1811
Georgalis et al. 2019
M1: a premaxilla (UU MAA 7093), eight maxillae (UU MAA 7342, UU MAA 7355, UU MAA 7356, UU MAA 7360, and UU MAA 7428), six dentaries (UU MAA 7103, UU MAA 7105, and UU MAA 7427), 14 fragments of tooth bearing bones (UU MAA 7102, UU MAA 7311, UU MAA 7354, UU MAA 7393, UU MAA 7405, and UU MAA 7406), three axes (UU MAA 7346 and UU MAA 7378), three post-axial cervical vertebrae (UU MAA 7322, UU MAA 7348, and UU MAA 7789), 13 trunk vertebrae (UU MAA 7110, UU MAA 7345, UU MAA 7382-UU MAA 7384, UU MAA 7421), two cloacal vertebrae (UU MAA 7380 and UU MAA 7381), eight caudal vertebrae (UU MAA 7347, UU MAA 7385-UU MAA 7387, and UU MAA 7390), a fragment of caudal vertebra (UU MAA 7310), 11 ribs (UU MAA 7111 and UU MAA 7419), seven humeri (UU MAA 7309 and UU MAA 7352), two pelvic girdles (UU MAA 7349 and UU MAA 7376), 14 femurs (UU MAA 7109, UU MAA 7308, UU MAA 7351, UU MAA 7377, and UU MAA 7420), and a tibia (UU MAA 7350); M2: a fragment of a presacral vertebra (UU MAA 7299) and a caudal vertebra (UU MAA 7300); M3: a fragment of tooth bearing bone (UU MAA 7339) and a caudal vertebra (UU MAA 7221).
Reptilia
Serpentes indet. (Linnaeus 1758)
Szyndlar 1995 6 elements
6 precaudal and caudal vertebrae (AMPG MA 190 to 195), "may have belonged either to Naja or Vipera"
Scolecophidia indet. (Duméril and Bibron 1844)
Georgalis et al. 2019
M1: three trunk vertebrae (UU MAA 7373-UU MAA 7375).
Reptilia - Elapidae
Naja sp. Laurenti 1768
Szyndlar 1995 25 elements
25 fragmentary precaudal vertebrae (AMPG MA 163 to 187)
Reptilia - Crotalidae
Vipera sp. Laurenti 1768
Szyndlar 1995 2 elements
"oriental viper", 2 fragmentary precaudal vertebrae (AMPG MA 188, 189)
Reptilia - Colubridae
Colubrinae indet. Oppel 1811
Szyndlar 1995 36 elements
36 fragmentary precaudal vertebrae (AMPG MA 101 to 136)
Paraxenophis spanios n. gen., n. sp. Georgalis et al. 2019
Georgalis et al. 2019
Holotype. Specimen UU MAA 7645 (a mid-trunk vertebra) (Figure 27). The holotype originates from M1 site. Paratypes. Specimen UU MAA 7536 (a posterior trunk vertebra) (Figure 29.1-5); UU MAA 7533 (a posteriormost trunk vertebra) (Figure 29.6-8). Both paratypes originate from M1 site.
Natricinae indet. Bonaparte 1838
Szyndlar 1995 26 elements
26 fragmentary precaudal vertebrae (AMPG MA 137 to 162)
Periergophis micros n. gen., n. sp. Georgalis et al. 2019
Georgalis et al. 2019
Holotype. Specimen UU MAA 7615 (a posterior trunk vertebra) (Figures 22 and 24.1-2). The holotype originates from M1 site. Paratypes. Specimen UU MAA 7551 (a cervical vertebra) (Figure 23.1-2); UU MAA 7614 (a midtrunk vertebra) (Figure 23.3-8); UU MAA 7618 (a cloacal vertebra) (Figure 23.9-12); UU MAA 7636 (a caudal vertebra) (Figures 23.13-16, 24.3). All paratypes originate from M1 site.
Natrix aff. rudabanyaensis
Georgalis et al. 2019
M1: 34 precloacal vertebrae (UU MAA 7656-UU MAA 7689) and seven caudal vertebrae (UU MAA 7690-UU MAA 7694, UU MAA 7893, and UU MAA 7904); M3: four precloacal vertebrae (UU MAA 7741-UU MAA 7744).
Natrix sp. Laurenti 1768
Georgalis et al. 2019
M1: six precloacal vertebrae (UU MAA 7695-UU MAA 7700).
Reptilia - Lacertidae
Lacertidae indet. Bonaparte 1831
Georgalis et al. 2019
M1: three maxillae (UU MAA 7031, UU MAA 7032, and UU MAA 7438), a quadrate (UU MAA 7096), three pterygoids (UU MAA 7097-UU MAA 7099), 16 dentaries (UU MAA 7048-UU MAA 7061, UU MAA 7095, and UU MAA 7424), 10 fragments of tooth bearing bones (UU MAA 7087-UU MAA 7092, UU MAA 7094, UU MAA 7391, and UU MAA 7802), and an isolated tooth (UU MAA 7392); M3: a dentary (UU MAA 7220).
Reptilia
Laterata indet. Vidal and Hedges 2005
Georgalis et al. 2019
M1: 29 dentaries (UU MAA 7026, UU MAA 7106, UU MAA 7357, UU MAA 7361, UU MAA 7379, UU MAA 7402-UU MAA 7404, UU MAA 7435, and UU MAA 7436); M3: a dentary (UU MAA 7206).
Reptilia - Scincidae
? Scincidae indet. Gray 1825
Georgalis et al. 2019
M1: two dentaries (UU MAA 7027 and UU MAA 7107).
Reptilia - Cordylidae
aff. Palaeocordylus sp. Roček 1984
Georgalis et al. 2019
M1: 26 maxillae (UU MAA 7046, UU MAA 7047, UU MAA 7077-UU MAA 7086, UU MAA 7101, UU MAA 7273, UU MAA 7358, UU MAA 7362-UU MAA 7365, UU MAA 7370-UU MAA 7372, UU MAA 7395, UU MAA 7396, and UU MAA 7437), 57 dentaries (UU MAA 7001-UU MAA 7025, UU MAA 7034-UU MAA 7036, UU MAA 7039, UU MAA 7040, UU MAA 7104, UU MAA 7108, UU MAA 7266, UU MAA 7267, UU MAA 7270, UU MAA 7279, UU MAA 7359, UU MAA 7366-UU MAA 7369, UU MAA 7397-UU MAA 7401, UU MAA 7425, and UU MAA 7426), and six fragments of tooth bearing bones (UU MAA 7037 and UU MAA 7038); M3: a maxilla (UU MAA 7340), eight dentaries (UU MAA 7199-UU MAA 7205, and UU MAA 7209), and a fragment of tooth bearing bone (UU MAA 7210).
Reptilia - Agamidae
Agaminae indet.
Georgalis et al. 2019
M1: a premaxilla (UU MAA 7076), three maxillae (UU MAA 7043-UU MAA 7045); five dentaries (UU MAA 7041, UU MAA 7042, UU MAA 7075, UU MAA 7276, and UU MAA 7305); 15 fragments of tooth bearing bones (UU MAA 7063, UU MAA 7065-UU MAA 7074, UU MAA 7264, UU MAA 7303, UU MAA 7304, and UU MAA 7394); five caudal vertebrae (UU MAA 7062, UU MAA 7064, and UU MAA 7302); M2: a frontal (UU MAA 7191); M3: a premaxilla (UU MAA 7341), a maxilla (UU MAA 7208), and a left dentary (UU MAA 7207).
Reptilia
Anguimorpha indet. Fürbringer 1900
Georgalis et al. 2019
M1: a dentary (UU MAA 7353).
Reptilia - Varanidae
Varanus sp. Merrem 1820
Georgalis et al. 2019
M1: six teeth (UU MAA 7029, UU MAA 7030, and UU MAA 7148-UU MAA 7151) and 12 caudal vertebrae (UU MAA 7028, UU MAA 7278, UU MAA 7388, UU MAA 7389, UU MAA 7429-UU MAA 7434, UU MAA 7805, and UU MAA 7806); M3: a caudal vertebra (UU MAA 7198).
Reptilia - Anguidae
Anguinae indet. Gray 1825
Georgalis et al. 2019
M1: 188 osteoderms (UU MAA 7160-UU MAA 7170, UU MAA 7262, UU MAA 7265, UU MAA 7268, UU MAA 7271, UU MAA 7275, UU MAA 7306, and UU MAA 7314-UU MAA 7318); M2: eight osteoderms (UU MAA 7187, UU MAA 7188, and UU MAA 7298).
Ophisaurus sp. Daudin 1803
Georgalis et al. 2019
M1: a maxilla (UU MAA 7324) and two dentaries (UU MAA 7176 and UU MAA 7177).
Anguis sp. Linnaeus 1758
Georgalis et al. 2019
M1: 37 trunk vertebrae (UU MAA 7181, UU MAA 7263, UU MAA 7274, UU MAA 7280, UU MAA 7332-UU MAA 7334, UU MAA 7411, UU MAA 7418, and UU MAA 7804), a cloacal vertebra (UU MAA 7269); M3: two trunk vertebrae (UU MAA 7217 and UU MAA 7218).
Pseudopus sp. Merrem 1820
Georgalis et al. 2019
M1: a maxilla (UU MAA 7171), a dentary (UU MAA 7407), a fragment of tooth bearing bone (UU MAA 7307), two trunk vertebrae (UU MAA 7172 and UU MAA 7328), and a cloacal vertebra (UU MAA 7329); M3: three trunk vertebrae (UU MAA 7212-UU MAA 7214).
Pisces
Pisces indet. Linnaeus 1758
Schmidt-Kittler et al. 1995
synonym of Osteichthyes
see common names

Geography
Country:Greece State/province:Central Macedonia County:Serres
Coordinates: 41.2° North, 23.5° East (view map)
Paleocoordinates:40.9° North, 23.2° East
Basis of coordinate:estimated from map
Geographic resolution:small collection
Time
Period:Neogene Epoch:Miocene
10 m.y. bin:Cenozoic 6
*Period:Tertiary *Local age/stage:Middle Turolian - Late/Upper Turolian
Key time interval:Turolian
Age range of interval:8.70000 - 5.33300 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Formation:Lefkon
Stratigraphy comments: may be the same level as Ano Metochi 2-3
Lithology and environment
Primary lithology: conglomerate
Secondary lithology: sandy marl
Includes fossils?Y
Lithology description: A polymict poorly sorted conglomerate containing pebbles of schist, gneiss and granite. This coarse-grained series grades upwards into arkosic sands and finally into clays with occasional lignites. Within the top part of the formation, two "landslide" breccias occur. The lower of these consists of metamorphic rocks, the upper one of granite.
Environment:alluvial fan
Taphonomy
Modes of preservation:body
Size of fossils:mesofossils
Collection methods and comments
Collection methods:field collection
Reason for describing collection:taxonomic analysis
Museum repositories:AMPG
Metadata
Database number:32189
Authorizer:J. Alroy, E. Vlachos, L. van den Hoek Ostende, J. Mueller Enterer:E. Leckey, E. Vlachos, T. Liebrecht, L. van den Hoek Ostende
Modifier:T. Liebrecht Research group:vertebrate
Created:2003-06-09 16:30:14 Last modified:2012-11-25 17:53:22
Access level:the public Released:2003-06-09 16:30:14
Creative Commons license:CC BY
Reference information

Primary reference:

8408. H. de Bruijn. 1989. Smaller mammals from the Upper Miocene and Lower Pliocene of the Strimon basin, Greece. Part 1. Rodentia and Lagomorpha. Bollettino della Societa Paleontologica Italiana 28(2-3):189-195 [J. Alroy/E. Leckey/E. Leckey]

Secondary references:

84095% 15420C. S. Doukas. 1989. Smaller mammals from the Upper Miocene and Lower Pliocene of the Strimon basin, Greece. Part 2. Insectivora. Bollettino della Societa Palaeontologica Italiana 28(2-3):213-216 [J. Alroy/E. Leckey/E. Leckey]
73905 C. S. Doukas. 2005. Greece. In L. W. van den Hoek Ostende, C. S. Doukas, J. W. F. Reumer (eds.), The Fossil Record of the Eurasian Neogene Insectivores (Erinaceomorpha, Soricomorpha, Mammalia), Part I. Scripta Geologica Special Issue 5:99-112 [E. Vlachos/E. Vlachos]
19740 C. S. Doukas, L. W. van den Hoek Ostende, C. D. Theocharopoulos and J. W. F. Reumer. 1995. The Vertebrate Locality Maramena (Macedonia, Greece) at the Turolian-Ruscinian Boundary (Neogene). 5. Insectivora (Erinaceidae, Talpidae, Soricidae, Mammalia). Münchner Geowissenschaftliche Abhandlungen (A) 28:43-64 [L. van den Hoek Ostende/L. van den Hoek Ostende/T. Liebrecht]
70786 G. L. Georgalis, A. Villa, M. Ivanov, D. Vasilyan, and M. Delfino. 2019. Fossil amphibians and reptiles from the Neogene locality of Maramena (Greece), the most diverse European herpetofauna at the Miocene/Pliocene transition boundary. Palaeontologia Electronica 22.3.68:1-99 [E. Vlachos/E. Vlachos]
23416 C. G. Rümke. 1985. A review of fossil and recent Desmaninae (Talpidae, Insectivora). Utrecht Micropaleontological Bulletins 4:1-241 [L. van den Hoek Ostende/L. van den Hoek Ostende]
61128 N. Schmidt-Kittler, H. de Bruijn, and C. Doukas. 1995. The vertebrate locality Maramena (Macedonia, Greece) at the Turolian-Ruscinian Boundary (Neogene). 1. General Introduction. Munchner Goewiss. Abh 28:9-18 [E. Vlachos/E. Vlachos]
43602 Z. Szyndlar. 1995. The Vertebrate Locality Maramena (Macedonia, Greece) at the Turolian-Ruscinian Boundary (Neogene). 4. Serpentes (Squamata, Reptilia). Münchner Geowissenschaftliche Abhandlungen (A) 28:39-42 [J. Mueller/T. Liebrecht]
65163 E. Vlachos. 2015. The Fossil Chelonians of Greece. Systematics - Evolution - Stratigraphy - Palaeoecology. Scientific Annals of the School of Geology, Aristotle University of Thessaloniki, Greece 173:1-479 [E. Vlachos/E. Vlachos/E. Vlachos]