Hornsleasow Quarry: Early/Lower Bathonian, United Kingdom
collected by K. Gardner 1987

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Nodosariata - Robertinida - Epistominidae
Epistomina regularis
Metcalf et al. 1992
Trochamminidae
Trochammina canningensis
Metcalf et al. 1992
Lycopodiopsida - Selaginellales
Densoisporites velatus Dettmann 1963
Metcalf et al. 1992
Algae - Botryococcaceae
Botryococcus braunii Kutzing 1849
Metcalf et al. 1992
Coniferales - Podocarpaceae
Callialasporites sp. Sukh-Dev 1961
Metcalf et al. 1992
Anthozoa
Anthozoa indet. Ehrenberg 1834
Metcalf et al. 1992
"corals"
Gastropoda - Viviparidae
Viviparus cf. scoticus
Metcalf et al. 1992
Gastropoda - Naticidae
Naticidae indet. Guilding 1834
Metcalf et al. 1992
Gastropoda - Heterostropha - Valvatidae
Valvata cf. comes
Metcalf et al. 1992
Bivalvia
Bivalvia indet. Linnaeus 1758
Metcalf et al. 1992
Bivalvia - Pectinida - Limidae
? Plagiostoma sp. Sowerby 1814
Metcalf et al. 1992
Bivalvia - Ostreida - Gryphaeidae
Praeexogyra sp. Charles 1952
Metcalf et al. 1992
Ostracoda
Ostracoda indet. Latreille 1802
Metcalf et al. 1992
Echinoidea
Echinoidea indet. (Leske 1778)
Metcalf et al. 1992
Chondrichthyes - Hybodontiformes - Acrodontidae
Asteracanthus sp. Agassiz 1836
Metcalf et al. 1992
GLRCM G.50031
Amphibia - Anura
? Discoglossidae indet. Günther 1859
Metcalf et al. 1992
    = ? Anura indet. Fischer von Waldheim 1813
Metcalf and Walker 1994
"possible frog remains"
Amphibia - Temnospondyli
Caudata indet. Scopoli 1777
Metcalf et al. 1992
"salamanders"
Caudata informal indet. B Scopoli 1777
Evans and Milner 1994
? Marmorerpeton sp. Evans et al. 1988
Metcalf et al. 1992
GLRCM G.60406
Amphibia - Temnospondyli - Albanerpetontidae
? Albanerpetontidae indet. Fox and Naylor 1982
Evans and Milner 1994
    = Albanerpetontidae indet. Fox and Naylor 1982
Gardner and Böhme 2008
unclassified
Therapsida indet. Broom 1905
Evans and Milner 1994
Tritylodontidae
Tritylodontidae indet. Cope 1884
Evans and Milner 1994
Stereognathus sp. Charlesworth 1854
Metcalf et al. 1992 70 specimens
GLRCM G.53809
    = Stereognathus ooliticus Charlesworth 1854
Panciroli et al. 2017
Mammalia
Mammalia indet. Linnaeus 1758
Evans and Milner 1994
Mammalia - Multituberculata
Multituberculata indet. Cope 1884
Metcalf et al. 1992
Mammalia - Pantotheria
Eupantotheria indet. Kermack and Musset 1958
Metcalf et al. 1992
GLRCM G.66570; "eupantotheres"
? Eupantotheria indet. Kermack and Musset 1958
Metcalf et al. 1992
GLRCM G.66856
Docodonta
Docodonta indet. Kretzoi 1946
Metcalf et al. 1992
Reptilia - Testudines
Testudines indet. Batsch 1788
Metcalf et al. 1992
GLRCM G.66759
Reptilia - Squamata
aff. Squamata indet. Oppel 1811
Metcalf et al. 1992
"squamate affinites"
Reptilia - Crocodylia
Crocodylia indet. (Owen 1842)
Metcalf et al. 1992
GLRCM G.55545, 50002
Reptilia - Goniopholididae
Goniopholidae indet. (Cope 1875)
Metcalf et al. 1992 40 %-specimens
original and current combination Goniopholididae
"goniopholids"
Reptilia - Teleosauridae
? Teleosauridae indet. Geoffroy 1831
Metcalf et al. 1992
Reptilia - Fabrosauridae
? Phyllodon sp. Thulborn 1975
Metcalf et al. 1992
GLRCM G.53264; "fabrosaurid similar to Phyllodon sp."
    = ? Fabrosauridae indet. Galton 1972
Evans and Milner 1994
Reptilia
? Stegosauria indet. Marsh 1877
Metcalf et al. 1992
GLRCM G.64020
Reptilia - Hypsilophodontidae
cf. Alocodon sp.
Metcalf et al. 1992
"hypsilophodontid similar to Alocodon sp."
    = Hypsilophodontidae indet. Dollo 1882
Evans and Milner 1994
Reptilia
? Maniraptora indet. Gauthier 1986
Metcalf et al. 1992
GLRCM G.51422; small theropods, possibly maniraptoran or dromaeosaurid
    = Coelurosauria indet. Huene 1914
Evans and Milner 1994
Reptilia - Megalosauridae
Megalosaurus sp. Parkinson 1822
Metcalf et al. 1992
GLRCM G.66742
Reptilia
Ceratosauria indet. (Marsh 1884)
Metcalf et al. 1992
Cetiosaurus sp. Owen 1841
Metcalf et al. 1992
GLRCM G.50051
? Pterodactyloidea indet. Plieninger 1901
Metcalf et al. 1992
    = Pterosauria indet. Kaup 1834
Unwin 1996
Reptilia - Pterosauria
Pterosauria indet. Kaup 1834
Reptilia - Rhamphorhynchidae
Rhamphorhynchoidea indet. Plieninger 1901
Metcalf et al. 1992
GLRCM G.66839; "rhamphorhynchoid"
    = Rhamphocephalus sp. Seeley 1880
Unwin 1996
Teeth and postcranial remains
Reptilia
Lepidosauromorpha indet. Benton 1983
Evans and Milner 1994
Lepidosauromorpha informal indet. A Benton 1983
Evans and Milner 1994
Reptilia - Sphenodontia
Sphenodontidae indet. Cope 1871
Metcalf et al. 1992
    = Sphenodontia indet. Jaekel 1910
Metcalf and Walker 1994
Reptilia - Choristodera
Choristodera indet. Cope 1876
Evans and Milner 1994
Reptilia - Cteniogenyidae
Cteniogenys sp. Gilmore 1928
Metcalf et al. 1992
GLRCM G.66855, 51749
    = cf. Cteniogenys sp. Gilmore 1928
Metcalf and Walker 1994
"similar to those of the small diapsid reptile Cteniogenys sp."
Actinopteri - Pycnodontiformes - Pycnodontidae
Pycnodontidae indet. (Agassiz 1833)
Metcalf et al. 1992
Actinopteri - Lepisosteiformes
Lepidotes sp. Agassiz 1832
Metcalf et al. 1992
GLRCM G.50001
see common names

Geography
Country:United Kingdom State/province:England County:North Gloucestershire
Coordinates: 52.0° North, 1.8° West (view map)
Paleocoordinates:41.6° North, 8.6° East
Basis of coordinate:estimated from map
Altitude:153 meters
Geographic resolution:outcrop
Time
Period:Jurassic Epoch:Middle Jurassic
Stage:Bathonian 10 m.y. bin:Jurassic 4
*Period:Middle Jurassic
*International age/stage:Early/Lower Bathonian
Key time interval:Early/Lower Bathonian Zone: Zigzagiceras zigzag
Age range of interval:167.70000 - 164.70000 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Geological group:Great Oolite Formation:Chipping Norton Limestone
Stratigraphic resolution:bed
Stratigraphy comments: "Hornsleasow Clay Unit" placed between Chipping Norton Member (above) and Hook Norton Member (below). Upper part of unit is the "Green Clay" and lower part is the "Grey Clay". From the macrescens Subzone of the zigzag Zone
Lithology and environment
Primary lithology:planar lamination,bioturbation,pebbly carbonaceous claystone
Secondary lithology:desiccation cracks claystone
Includes fossils?Y
Includes fossils?Y
Lithology description: Primary (Grey Clay): "matrix-supported, but contains large, irregularu subrounded limestone cobbles (concentrated in the basal portion, which is interpreted as a regolith), with associated lime and lime-fraction silica sand. The clay is smectitic, with a high organic content, and it contains much partially lignitized wood...finely laminated. This lamination has been somewhat disrupted, possibly by bioturbation." Secondary (Green Clay): "more homogenous than the Grey Clay. Limestone clasts and sand make up only 3 percent of the composition by weight. The deposit has more silica and sand and is an illite/chlorite clay. Vertical rootlets pervade the layer, extending from the mud-cracked, upper surface...into the underlying Grey Clay...The plant material is somewhat oxidized."
Environment:karst indet.
Glacial or sequence phase:regressive
Geology comments: Clay units were deposited into the karstic surface of the Hook Norton Member. Clay deposition involved low paleocurrent activity and increasingly anoxic conditions. The unit "is interpreted as a paleosol. The depositional environment...is one of marshy, low-pH type. The partially decayed organic material within this deposit suggests relatively reducing conditions, and the presence of pyrolusite (MnO2)...is suggestive of bog or lake deposition...salinity was low to moderate...The Grey Clay is a waterlogged, boggy paleosol developing within a freshwater to brackish-water pond. The absence of dispersed kerogen indicates a low-energy depositional environment, with little fluvial influence...The Green Clay is also interpreted as a paleosol, deposited under more oxidizing, high-pH conditions.
Taphonomy
Modes of preservation:body
Lagerst�tten type:concentrate
Degree of concentration:-multiple events
Size of fossils:macrofossils,mesofossils,microfossils
Spatial orientation:random
Preservation of anatomical detail:poor
Articulated whole bodies:none
Associated major elements:some
Disassociated major elements:many
Disassociated minor elements:many
Fragmentation:frequent
Bioerosion:occasional
Feeding/predation traces:gastric dissolution
Temporal resolution:time-averaged
Spatial resolution:parautochthonous
Collection methods and comments
Collection methods:bulk,surface (in situ),sieve,field collection
Reason for describing collection:general faunal/floral analysis
Collectors:K. Gardner Collection dates:1987
Metadata
Also known as:Snowshill Quarry
Database number:31054
Authorizer:M. Carrano, R. Butler Enterer:M. Carrano, K. Maguire, R. Butler
Modifier:M. Carrano Research group:vertebrate
Created:2003-04-09 20:44:33 Last modified:2012-07-05 15:20:58
Access level:the public Released:2003-04-09 20:44:33
Creative Commons license:CC BY
Reference information

Primary reference:

8380.5% 12660S. J. Metcalf and R. J. Walker. 1994. A new Bathonian microvertebrate locality in the English Midlands. In the Shadow of the Dinosaurs: Early Mesozoic Tetrapods, edited by N. C. Fraser and H.-D. Sues. Cambridge University Press, Cambridge 322-331 [M. Carrano/M. Carrano/J. Alroy]

Secondary references:

10557ETE S. E. Evans and A. R. Milner. 1994. Middle Jurassic microvertebrate assemblages from the British Isles. In the Shadow of the Dinosaurs: Early Mesozoic Tetrapods, N. C. Fraser and H.-D. Sues (eds.), Cambridge University Press 303-321 [M. Carrano/M. Carrano/M. Carrano]
52039 J. D. Gardner and M. Böhme. 2008. Review of the Albanerpetontidae (Lissamphibia), with comments on the paleoecological preferences of European Tertiary albanerpetontids. In J. T. Sankey & S. Baszio (ed.), Vertebrate Microfossil Assemblages: Their Role in Paleoecology and Paleobiogeography 178-218 [M. Carrano/M. Carrano]
27913ETE S. J. Metcalf, R. F. Vaughan, M. J. Benton, J. Cole, M. J. Simms and D. L. Dartnall. 1992. A new Bathonian (Middle Jurassic) microvertebrate site, within the Chipping Norton Limestone Formation at Honsleasow Quarry, Gloucestershire. Proceedings of the Geologists' Association 103:321-342 [M. Carrano/M. Carrano]
65143 E. Panciroli, S. Walsh, N. C. Fraser, S. L. Brusatte, and I. Corfe. 2017. A reassessment of the postcanine dentition and systematics of the tritylodontid Stereognathus (Cynodontia, Tritylodontidae, Mammaliamorpha), from the Middle Jurassic of the United Kingdom. Journal of Vertebrate Paleontology 37(5):e1351448:1-20 [M. Carrano/M. Carrano]
30893 D. M. Unwin. 1996. The fossil record of Middle Jurassic pterosaurs. In M. Morales (ed.), The Continental Jurassic, Museum of Northern Arizona Bulletin 60:291-304 [R. Butler/R. Butler]