Laño L1B (S2U1): Late/Upper Campanian, Spain

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Actinopteri - Lepisosteiformes - Lepisosteidae
Atractosteus sp. Rafinesque 1820
Pereda-Suberbiola et al. 2000
Reptilia - Madtsoiidae
Madtsoia laurasiae Rage 1996
Pereda-Suberbiola et al. 2000
recombined as Menarana laurasiae
Reptilia - Testudines
Pereda-Suberbiola et al. 2000
Reptilia
Pereda-Suberbiola et al. 2000
Reptilia - Azhdarchidae
cf. Azhdarcho sp. Nessov 1981
Pereda-Suberbiola et al. 2000
Reptilia
Lirainosaurus astibiae Sanz et al. 1999
Pereda-Suberbiola et al. 2000
Neoceratosauria indet. Novas 1991
Pereda-Suberbiola et al. 2000 4 specimens
    = Theropoda informal indet. 1 Marsh 1881
Torices et al. 2015
four teeth (MCNA 22051, 4520, 14521, and 1852)
Theropoda informal indet. 2 (Marsh 1881)
Torices et al. 2015 2 specimens
MCNA 1853, 14522, teeth
Coelurosauria indet. (Huene 1914)
Torices et al. 2015 42 specimens
MCNA 14523–14565, teeth
Reptilia - Dromaeosauridae
? Dromaeosauridae indet. Colbert and Russell 1969
Torices et al. 2015 1 specimen
MCNA 14622
? Pyroraptor olympius Allain and Taquet 2000
Torices et al. 2015 4 specimens
Four teeth (MCNA 14623–14626)
? Richardoestesia sp. Currie et al. 1990
Torices et al. 2015 64 specimens
sixty-four isolated teeth (MCNA 14566–14621)
Reptilia - Gargantuaviidae
Gargantuavis philoinos Buffetaut and Le Loeuff 1998
Angst et al. 2017
MCNA 2583
Reptilia - Nodosauridae
Struthiosaurus sp. Bunzel 1870
Pereda-Suberbiola et al. 2000
Reptilia - Rhabdodontidae
Rhabdodon sp. (Matheron 1869)
Pereda-Suberbiola et al. 2000
Reptilia
Musturzabalsuchus buffetauti Buscalioni et al. 1997
Pereda-Suberbiola et al. 2000
Acynodon iberoccitanus Buscalioni et al. 1997
Pereda-Suberbiola et al. 2000
Chondrichthyes - Lamniformes - Otodontidae
Cretolamna sp. Glikman 1958
Cretolamna appendiculata (Agassiz 1835)
recombined as Cretalamna appendiculata
Chondrichthyes - Rajiformes - Sclerorhynchidae
Ganopristis leptodon Arambourg 1935
Chondrichthyes - Myliobatiformes - Rhombodontidae
Rhombodus binkhorsti Dames 1881
see common names

Geography
Country:Spain State/province:Castilla y Leon County:Burgos
Coordinates: 42.7° North, 2.6° West (view map)
Paleocoordinates:33.0° North, 0.1° West
Basis of coordinate:estimated from map
Altitude:747 meters
Geographic resolution:outcrop
Time
Period:Cretaceous Epoch:Late/Upper Cretaceous
Stage:Campanian 10 m.y. bin:Cretaceous 7
*Period:Late/Upper Cretaceous *Epoch:Maastrichtian
Key time interval:Late/Upper Campanian
Age range of interval:83.60000 - 72.10000 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Formation:Vitoria
Local section:Lano section Local bed:S2U1
Local order:bottom to top
Stratigraphic resolution:bed
Stratigraphy comments: separated from underlying S1U3 by an unconformity, level also called L1B
Lithology and environment
Primary lithology:dolomitic sandy "limestone"
Includes fossils?Y
Lithology description: marine sediments passing upward to continental rocks. Mainly calcareous-dolomite composition, lacking, in general, current structures. Calcarenites and calcirudites
Environment:shallow subtidal indet.
Glacial or sequence phase:regressive
Geology comments: unit begins with marine sediments passing upwards to continental rocks
Taphonomy
Modes of preservation:body
Size of fossils:mesofossils,microfossils
Disassociated major elements:some
Disassociated minor elements:all
Collection methods and comments
Collection excludes:some genera
Collection methods:sieve,field collection
Reason for describing collection:taxonomic analysis
Metadata
Database number:28765
Authorizer:J. Alroy, P. Mannion, M. Carrano Enterer:E. Leckey, M. Carrano, P. Mannion, J. Tennant
Modifier:M. Carrano Research group:vertebrate
Created:2003-02-24 11:43:53 Last modified:2019-02-13 12:39:26
Access level:the public Released:2003-02-24 11:43:53
Creative Commons license:CC BY
Reference information

Primary reference:
7903. H. Astibia, E. Buffetaut, A. D. Buscalioni, H. Cappetta, C. Corral, R. Estes, F. Garcia-Garmilla, J. J. Jaeger, E. Jiminez-Fuentes, J. Le Loeuff, J. M. Mazin, X. Orue-Etexebarria, J. Pereda-Suberbiola, J. E. Powell, J. C. Rage, J. Rodriguez-Lazaro, J. L. Sanz and H. Tong. 1990. The fossil vertebrates from the Lano (Basque Country, Spain); new evidence on the composition and affinities of the Late Cretaceous continental faunas of Europe. Terra Nova 2:460-466 [J. Alroy/E. Leckey/M. Uhen]

Secondary references:

61633 D. Angst, E. Buffetaut, J. C. Corral and X. Pereda-Suberbiola. 2017. First record of the Late Cretaceous giant bird Gargantuavis philoinos from the Iberian Peninsula. Annales de Paléontologie 103:135-139 [P. Mannion/P. Mannion/P. Mannion]
17647ETE J. I. Canudo and J. I. Ruiz-Omeñaca. 2003. Los restos directos de dinosaurios teropódos (excluyendo Aves) en España [Direct remains of theropod dinosaurs (excluding Aves) in Spain]. Ciencias de la Tierra 26:347-373 [M. Carrano/M. Carrano]
43435 V. Díez Díaz, X. Pereda Suberbiola, and J. L. Sanz. 2012. Juvenile and adult teeth of the titanosaurian dinosaur Lirainosaurus (Sauropoda) from the Late Cretaceous of Iberia. Geobios 45:265-274 [M. Carrano/M. Carrano]
7842 J. Le Loeuff and E. Buffetaut. 1991. Tarascosaurus salluvicus nov. gen., nov. sp., dinosaure théropode du Crétacé supérieur du sud de la France [Tarascosaurus salluvicus nov. gen., nov. sp., a theropod dinosaur from the Upper Cretaceous of southern France]. Géobios 25(5):585-594 [M. Carrano/M. Carrano/M. Carrano]
9513 X. Pereda-Suberbiola. 1999. Ankylosaurian dinosaur remains from the Upper Cretaceous of Lano (Iberian Peninsula). Estudios del Museo de Ciencias Naturales de Alava 14(1):273-288 [J. Alroy/E. Leckey/E. Leckey]
9559 X. Pereda-Suberbiola, H. Asibia, X. Murelaga, J. J. Elzorza, and J. J. Gomez-Alday. 2000. Taphonomy of the Late Cretaceous dinosaur-bearing beds of the Lano Quarry (Iberian Peninsula). Palaeogeography, Palaeoclimatology, Palaeoecology 157:247-275 [J. Alroy/E. Leckey/E. Leckey]
14565ETE X. Pereda-Suberbiola, H. Astibia, and E. Buffetaut. 1995. New remains of the armoured dinosaur Struthiosaurus from the Late Cretaceous of the Iberian peninsula (Laño locality, Basque-Cantabric basin). Bulletin de la Société Géologique de France 166(2):207-211 [M. Carrano/M. Carrano/M. Carrano]
9512 X. Pereda-Suberbiola and J. Sanz. 1999. The ornithopod dinosaur Rhabdodon from the Upper Cretaceous of Lano (Iberian Peninsula). Estudios del Museo de Ciencias Naturales de Alava 14(1):257-272 [J. Alroy/E. Leckey/E. Leckey]
41499 A. Pérez-García, T. M. Scheyer, and X. Murelaga. 2012. New interpretations of Dortoka vasconica Lapparent de Broin and Murelaga, a freshwater turtle with an unusual carapace. Cretaceous Research 36:151-161 [R. Benson/R. Benson]
13503ETE J. L. Sanz, J. E. Powell, J. Le Loeuff, R. Martinez, and X. Pereda-Suberbiola. 1999. Sauropod remains from the Upper Cretaceous of Laño (northcentral Spain). Titanosaur phylogenetic relationships. Estudios del Museo de Ciencias Naturales de Alava 14(especial 1):235-255 [M. Carrano/M. Carrano/M. Carrano]
19710ETE F. Torcida Fernández-Baldor. 2005. Los dinosaurios de Castilla y León [The dinosaurs of Castilla y León]. Patrimonio Histórico de Castilla y León 6(23):23-34 [M. Carrano/M. Carrano/M. Carrano]
56091 A. Torices, P. J. Currie, J. I. Canudo and X. Pereda-Suberbiola. 2015. Theropod dinosaurs from the Upper Cretaceous of the South Pyrenees Basin of Spain. Acta Palaeontologica Polonica 60(3):611-626 [P. Mannion/J. Tennant/M. Carrano]