Fox-Amphoux-La Bastide Neuve: Late/Upper Campanian, France
collected by P. Mechin, Germain, Palaios/ University of Poitiers/RBINS 1987, 1990, 2009, 2012

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Reptilia
Dinosauria indet. (Owen 1842)
Database 2015 2 specimens
CM acc. 33679 (field nos. 48-36, 48-45)
Garrigatitan meridionalis n. gen., n. sp. Díez Díaz et al. 2021
Díez Díaz et al. 2021
MMS/VBN.09.170 - holotype; referred specimens: MMS/VBN.02.99, 09.A.016, 09.47, 12B.12a, 12B.12b; tentatively referred specimens: MMS/VBN.12B.011, 12.82, 09.A.017, 00.13
Reptilia - Saltasauridae
Titanosauridae indet. Lydekker 1885
Le Loeuff and Buffetaut 1998
(1 measurement)
    = Saltasauridae indet. Sereno 1998
Chanthasit and Buffetaut 2009
Reptilia - Nodosauridae
Ankylosauria indet. Osborn 1923
Le Loeuff and Buffetaut 1998
    = Nodosauridae indet. Marsh 1890
Garcia et al. 2010
Reptilia - Rhabdodontidae
Ornithopoda indet. Marsh 1881
Le Loeuff and Buffetaut 1998
    = Rhabdodon sp. Matheron 1869
Chanthasit and Buffetaut 2009
Reptilia - Dromaeosauridae
Dromaeosauridae indet. Colbert and Russell 1969
Chanthasit and Buffetaut 2009 6 specimens
CM-203, 307, 186, 218, 537, 684
cf. Dromaeosauridae indet. Colbert and Russell 1969
Brilhante et al. 2022
MDE-D25 (proximal half of a hypertrophied ungual phalanx of the left pedal digit II)
Variraptor mechinorum n. gen., n. sp. Le Loeuff and Buffetaut 1998
Le Loeuff and Buffetaut 1998 3 specimens
MDE-D169, CM-645 (2 measurements)
Reptilia - Gargantuaviidae
Gargantuavis philoinos Buffetaut and Le Loeuff 1998
Buffetaut and Le Loeuff 2011
partial synsacrum; Mechin collection, n° 711 (femur)
Reptilia
Aves indet. Linnaeus 1758
Le Loeuff and Buffetaut 1998
    = Enantiornithes indet. Walker 1981
Buffetaut et al. 2000
Mechin collection no. 606 (complete right tibiotarsus)
Reptilia - Abelisauridae
Abelisauridae indet. Bonaparte and Novas 1985
Cavin et al. 2020
Reptilia - Azhdarchidae
Azhdarchidae indet. Nessov 1984
Buffetaut et al. 2006 1 specimen
Right humerus representing an individual with a 3 m wingspan
Reptilia
Crocodylidae indet. Cuvier 1807
Le Loeuff and Buffetaut 1998
    = Acynodon iberoccitanus Buscalioni et al. 1997
Martin 2007
ACAP-FX1 (a complete skull) and ACAP-FX2 (an almost complete skull with premaxillary and maxillary dentition)
Alligatoridae indet. Gray 1844
Le Loeuff and Buffetaut 1998
    = Allodaposuchus precedens Nopcsa 1928
Martin et al. 2016
MMS/VBN-12-42, a nearly complete skull of a juvenile; MMS/VBN-12-10A, a complete adult skull; MMS/VBN-12-10B, a right dentary associated with the large adult skull MMS/VBN-12- 10A; MMS/VBN-93-28, a dorsoventrally compressed subadult skull; MMS/VBN-12-10D, a juvenile skull table; MMS/VBN-93-29, isolated fragmentary left maxilla; MMS/VBN-93-30, isolated fragmentary right maxilla possibly corresponding to MMS/VBN-93-29; MMS/ VBN-12-10F, one dorsal osteoderm; MMS/VBN-02- 37, a left femur
    = Allodaposuchus sp. Nopcsa 1928
Blanco and Brochu 2016
Solemys gaudryi (Matheron 1869)
Tong et al. 2023
PAM 485, MDE T54, MDE T55
Reptilia - Testudines
Podocnemidae indet.
Le Loeuff and Buffetaut 1998 6 specimens
MDEt 10 (holotype), a dorsoventrally crushed skull; an almost complete skull (figs. 103, 104, 107, 281; PAM 511A, Tong et al., 1998: figs. 3–6); an incomplete lower jaw (figs. 235, 236; PAM 511B, Tong et al., 1998: figs. 7, 8); an almost complete and wellpreserved shell in which the plastron is complete and only the posterior part of the carapace is missing (lacking right sixth to eighth costal and part of seventh and eighth left costal bones, eighth to eleventh right peripheral and eleventh left peripheral bones, suprapygal and pygal bones; PAM 548, Tong et al., 1998: figs. 9, 10); a partial shell consisting of the anterior third of the carapace in poor condition and a complete plastron (MDEt 09, Tong et al., 1998: fig. 11); a right scapula-coracoid (MDEt 11); and several isolated plates, an almost complete shell, MHNM uncatalogued
    = Foxemys mechinorum n. gen., n. sp. Tong et al. 1998
Tong et al. 1998
Coelacanthimorpha - Coelacanthiformes - Mawsoniidae
Axelrodichthys megadromos Cavin et al. 2016
Cavin et al. 2020 1 specimen
R prearticular, collection Méchin 745
Actinopteri - Lepisosteiformes - Lepisosteidae
Lepisosteidae indet. Cuvier 1825
Cavin et al. 2020
Chondrichthyes - Hybodontiformes - Hybodontidae
Hybodontidae indet. (Agassiz 1834)
Cavin et al. 2020
see common names

Geography
Country:France State/province:Provence-Alpes-Côte d'Azur
Coordinates: 43.6° North, 6.1° East (view map)
Paleocoordinates:34.3° North, 6.8° East
Basis of coordinate:estimated from map
Altitude:424 meters
Geographic resolution:outcrop
Time
Period:Cretaceous Epoch:Late/Upper Cretaceous
Stage:Campanian 10 m.y. bin:Cretaceous 7
*Period:Late/Upper Cretaceous
*International age/stage:Late/Upper Campanian - Early/Lower Maastrichtian
Key time interval:Late/Upper Campanian
Age range of interval:83.50000 - 70.60000 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Stratigraphic resolution:group of beds
Stratigraphy comments: part of the Fox-Amphoux syncline; lower Rognacian

Late Campanian age, based of magnetostratigraphic analyses and biostratigraphic information derived from charophyte and dinosaur eggshell biozones (Cincotta et al. 2015)
Lithology and environment
Primary lithology:yellow sandy claystone
Includes fossils?Y
Lithology description: "yellowish sandy clays"
Environment:fluvial indet.
Geology comments: "as a freshwater environment corresponding to fluvial setting of moder- ate energy with broad floodplain"
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils,mesofossils
Preservation of anatomical detail:medium
Articulated whole bodies:none
Associated major elements:some
Disassociated major elements:many
Disassociated minor elements:many
Fragmentation:occasional
Collection methods and comments
Collection methods:selective quarrying,mechanical,field collection
Reason for describing collection:taxonomic analysis
Collectors:P. Mechin, Germain, Palaios/ University of Poitiers/RBINS Collection dates:1987, 1990s, 2009, 2012
Metadata
Database number:28454
Authorizer:M. Carrano, P. Mannion Enterer:M. Carrano, P. Mannion
Modifier:P. Mannion Research group:vertebrate
Created:2003-02-14 10:41:51 Last modified:2022-08-17 04:28:53
Access level:the public Released:2003-02-14 10:41:51
Creative Commons license:CC BY
Reference information

Primary reference:

7841. J. Le Loeuff, E. Buffetaut, P. Méchin and A. Méchin-Salessy. 1992. The first record of dromaeosaurid dinosaurs (Saurischia, Theropoda) in the Maastrichtian of southern Europe: palaeobiogeographical implications. Bulletin de la Société géologique de la France 163(3):337-343 [M. Carrano/M. Carrano/M. Carrano]

Secondary references:

59690 A. Blanco and C. A. Brochu. 2016. Intra- and interspecific variability in allodaposuchid crocodylomorphs and the status of western European taxa. Historical Biology [P. Mannion/P. Mannion]
80976 N. S. Brilhante, T. Constância de França, F. Castro, L. Sanches Costa, P. J. Currie, S. A. Kugland Azevedo, and R. Delcourt. 2022. A dromaeosaurid-like claw from the Upper Cretaceous of southern France. Historical Biology [P. Mannion/G. Varnham]
30133 E. Buffetaut. 2008. Late Cretaceous pterosaurs from France: a review. Zitteliana B 28:249-255 [R. Butler/R. Butler]
76670 E. Buffetaut, D. Angst, P. Mechin and A. Mechin-Salessy. 2019. A femur of the giant bird Gargantuavis from the Late Cretaceous of Var (south-eastern France). Carnets natures 6:47-52 [P. Mannion/P. Mannion]
38140 E. Buffetaut and J. Le Loeuff. 2011. Gargantuavis philoinos: Giant bird or giant pterosaur?. Annales de Paléontologie 96:135-141 [P. Mannion/P. Mannion]
38141 E. Buffetaut, P. Mechin, and A. Mechin-Salessy. 2000. An archaic bird (Enantiornithes) from the Upper Cretaceous of Provence (southern France). Comptes Rendus de l'Académie des Sciences, Sciences de la Terre et des Planètes 331:557-561 [P. Mannion/P. Mannion]
30136 E. Buffetaut, P. Mechin, and A. Mechin-Salessy. 2006. An azhdarchid pterosaur from the Upper Cretaceous of Provence (southern France). In Z. Csiki (ed.), Mesozoic and Cenozoic vertebrates and paleoenvironments, tributes the career of Professor Dan Grigorescu; Editura Ars, Docendi, Bucharest 95-100 [R. Butler/R. Butler]
76814 L. Cavin, E. Buffetaut, Y. Dutour, G. Garcia, J. Le Loeuff, A. Méchin, P. Méchin, H. Tong, T. Tortosa, E. Turini, and X. Valentin. 2020. The last known freshwater coelacanths: new Late Cretaceous mawsoniid remains (Osteichthyes: Actinistia) from southern France. PLoS One 15(6):e0234183:1-19 [M. Carrano/M. Carrano]
29887 P. Chanthasit and E. Buffetaut. 2009. New data on the Dromaeosauridae (Dinosauria: Theropoda) from the Late Cretaceous of southern France. Bulletin de la Société Géologique de France 180(2):145-154 [M. Carrano/M. Carrano/M. Carrano]
54744 C. M. Database. 2015. [M. Carrano/M. Carrano]
66247 V. Díez Díaz, G. Garcia, X. Pereda Suberbiola, B. Jentgen-Ceschino, K. Stein, P. Godefroit, and X. Valentin. 2018. The titanosaurian dinosaur Atsinganosaurus velauciensis (Sauropoda) from the Upper Cretaceous of southern France: New material, phylogenetic affinities, and palaeobiogeographical implications. Cretaceous Research 91:429-456 [P. Mannion/P. Mannion/P. Mannion]
74890 V. Díez Díaz, G. Garcia, X. Pereda Suberbiola, B. Jentgen-Ceschino, K. Stein, P. Godefroit, and X. Valentin. 2021. A new titanosaur (Dinosauria: Sauropoda) from the Upper Cretaceous of Velaux-La-Bastide Neuve (southern France). Historical Biology 33(11):2998-3017 [P. Mannion/P. Mannion/M. Carrano]
76794 V. Díez Díaz and X. Pereda Suberbiola. 2010. Titanosaurian fossil record of Europe: a review. 8th Meeting of the European Association of Vertebrate Palaeontologists. Abstract Volume 30 [M. Carrano/M. Carrano/M. Carrano]
76809 Y. Dutour and T. Tortosa. 2010. Aix-en-Provence Basin—between eggs and dinosaurs. 8th Meeting of the European Association of Vertebrate Palaeontologists. Abstract Volume 86-93 [M. Carrano/M. Carrano]
33197ETE G. Garcia, S. Amico, F. Fournier, E. Thouand, and X. Valentin. 2010. A new titanosaur genus (Dinosauria, Sauropoda) from the Late Cretaceous of southern France and its paleobiogeographic implications. Bulletin de la Société géologique de France 181(3):269-277 [M. Carrano/M. Carrano]
80599 E. Isasmendi, A. Torices, J. I. Canudo, P. J. Currie, and X. Pereda-Suberbiola. 2022. Upper Cretaceous European theropod palaeobiodiversity, palaeobiogeography and the intra‐Maastrichtian faunal turnover: new contributions from the Iberian fossil site of Laño. Papers in Palaeontology 8(1):e1419:1-38 [M. Carrano/M. Carrano]
59044 Y. Laurent, J. Le Loeuff, M. Bilotte, E. Buffetaut, and G. S. Odin. 2001. Campanian-Maastrichtian continental-marine connection at the Aquitaine-Pyrenees-Provence area (S France). In G. S. Odin (ed.), The Campanian-Maastrichtian Boundary. Developments in Palaeontology and Stratigraphy 19:657-674 [M. Carrano/M. Carrano]
36728ETE J. Le Loeuff. 1998. Les dinosaures du Crétacé supérieur de l'Île Ibéro-Armoricaine [The dinosaurs of the Upper Cretaceous of the Ibero-Armorican Island]. In A. M. G. de Carvalho, A. M. Andrade, V. F. dos Santos, J. Cascalho, R. Taborda, (eds.), I Encontro Internacional sobre Paleobiologia dos Dinossáurios. Museu Nacional de História Natural, Lisboa 49-64 [M. Carrano/M. Carrano]
7840 J. Le Loeuff and E. Buffetaut. 1998. A new dromaeosaurid theropod from the Upper Cretaceous of southern France. Oryctos 1:105-112 [M. Carrano/M. Carrano/M. Carrano]
37002 J. E. Martin. 2007. New material of the Late Cretaceous globidontan Acynodon iberoccitanus (Crocodylia) from southern France. Journal of Vertebrate Paleontology 27(2):362-372 [P. Mannion/P. Mannion]
76450 M. Pincemaille-Quillevere. 2000. Rhabdodon priscus (Ornithopoda, Dinosauria) sites from southern France. In M.-C. Buchy (ed.), 5th European Workshop on Vertebrate Palaeontology, Program. Abstracts. Excursion GuidesGiganotosaurus carolinii 61-62 [M. Carrano/M. Carrano]
83913 H. Tong, E. Buffetaut, P. Méchin, A. Méchin-Salessy, and J. Claude. 2023. A Solemys Skull from the Late Cretaceous of Southern France. Diversity 15:58:1-17 [P. Mannion/P. Mannion]
24140ETE H. Tong, E. S. Gaffney, and E. Buffetaut. 1998. Foxemys, a new side-necked turtle (Bothremydidae: Pelomedusoides) from the Late Cretaceous of France. American Museum Novitates 3251:1-19 [M. Carrano/M. Carrano]
76807 T. Tortosa, Y. Dutour, and G. Cheylan. 2010. New discoveries and review of Late Cretaceous titanosaurs from Provence, France. 8th Meeting of the European Association of Vertebrate Palaeontologists. Abstract Volume 84 [M. Carrano/M. Carrano]
55429 B. Vila, A. Galobart, J. I. Canudo, J. Le Loeuff, J. Dinarès-Turell, V. Riera, O. Oms, T. Tortosa, and R. Gaete. 2012. The diversity of sauropod dinosaurs and their first taxonomic succession from the latest Cretaceous of southwestern Europe: Clues to demise and extinction. Palaeogeography, Palaeoclimatology, Palaeoecology 350–352:19-38 [M. Carrano/M. Carrano]