Napak (Napak Member): Burdigalian, Uganda
collected by W. W. Bishop 1958 - 1964

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
? Krapfiella sp.
Pickford et al. 1986
Burtoa nilotica
Pickford et al. 1986
Thapsia sp.
Pickford et al. 1986
Trochonanina sp.
Pickford et al. 1986
Tayloria sp.
Pickford et al. 1986
Gonaxis sp.
Pickford et al. 1986
Edentulina sp.
Pickford et al. 1986
Gulella sp.
Pickford et al. 1986
Ptychotrema sp.
Pickford et al. 1986
Mammalia - Primates - Dendropithecidae
Micropithecus clarki Fleagle and Simons 1978
Fleagle and Simons 1978
type from Napak IV; referred specimens from Napak I, IV, V, and IX
Mammalia - Primates
Lomorupithecus harrisoni n. gen., n. sp. Rossie and MacLatchy 2006
Rossie and MacLatchy 2006
type collected in 2002 from Napak IV
Cercopithecoidea indet. Gray 1821
Pickford et al. 1986
Mammalia - Rodentia - Thryonomyidae
Paraphiomys stromeri
Pickford et al. 1986
Paraphiomys pigotti Andrews 1914
Pickford et al. 1986
Mammalia - Rodentia
Diamantomys luederitzi
Pickford et al. 1986
Mammalia - Creodonta - Hyaenodontidae
Prionogale breviceps Schmidt-Kittler and Heizmann 1991
Morales et al. 2007 1 specimen
Mammalia - Creodonta
Metasinopa napaki Savage 1965
Savage 1965 1 specimen
Teratodon spekei Savage 1965
Morales et al. 2007 1 specimen
Metapterodon aff. stromeri Morales et al. 1998
Morales and Pickford 2017 1 specimen
NAP V 121'08 from Napak V
Pterodon africanus Andrews 1903
Savage 1965 1 specimen
    = Hyainailouros napakensis Ginsburg 1980
Morales and Pickford 2017
Isohyaenodon zadoki (Savage 1965)
Morales et al. 2007 1 specimen
    = Sectisodon occultus n. gen., n. sp. Morales and Pickford 2017
Morales and Pickford 2017
UM NAP V 178’08 from Napak V (holotype); referred specimens from Napak I and IV
Isohyaenodon pilgrimi (Savage 1965)
Morales et al. 2007 1 specimen
original and current combination Hyaenodon (Isohyaenodon) pilgrimi
Leakitherium sp. Savage 1965
Morales et al. 2007 1 specimen
Mammalia - Carnivora
Carnivora indet. Bowdich 1821
Pickford et al. 1986
said to be new
Mammalia - Carnivora - Amphicyonidae
Afrocyon sp. Arambourg 1961
Morales et al. 2016
NAP I 7'99 from Napak 1
Hecubides euryodon Savage 1965
Savage 1965 6 specimens
Mammalia - Carnivora - Nimravidae
Ginsburgsmilus napakensis Morales et al. 2001
Morales et al. 2007 1 specimen
Mammalia - Carnivora - Herpestidae
Ugandictis napakensis n. gen., n. sp. Morales et al. 2007
Morales et al. 2007 4 specimens
Mammalia - Carnivora - Viverridae
Kichechia zamanae Savage 1965
Savage 1965 2 specimens
Mammalia - Perissodactyla - Rhinocerotidae
Dicerorhinus leakeyi
Pickford et al. 1986
Mammalia - Perissodactyla - Chalicotheriidae
Chalicotherium rusingense Butler 1965
Pickford et al. 1986
Mammalia
Walangania africanus Whitworth 1958
Pickford et al. 1986
Mammalia - Tragulidae
Dorcatherium songhorensis Whitworth 1958
Pickford et al. 1986
Mammalia - Suidae
Nguruwe kijivium Pickford 1986
Pickford et al. 1986
Mammalia - Hyracoidea - Titanohyracidae
Megalohyrax championi (Arambourg 1933)
Pickford et al. 1986
recombined as Afrohyrax championi
Mammalia - Ptolemaiida - Kelbidae
Kelba quadeemae Savage 1965
Savage 1965 2 specimens
Reptilia - Testudines
Chelonia indet. (Latreille 1800)
Pickford et al. 1986
synonym of Testudines
Reptilia - Ophidia
Ophidia indet. Latreille 1804
Pickford et al. 1986
Gastropoda - Heterostropha - Achatinidae
Limicolaria sp. Schumacher 1817
Pickford et al. 1986
Achatina leakeyi
Pickford et al. 1986
Gastropoda - Heterostropha - Subulinidae
Subulinidae indet. Fischer and Crosse 1877
Pickford et al. 1986
Gastropoda - Heterostropha
Trochozonites sp. Pfeffer 1883
Pickford et al. 1986
Gastropoda - Maizaniidae
Maizania sp. Bourguignat 1889
Pickford et al. 1986
see common names

Geography
Country:Uganda
Coordinates: 2.1° North, 34.2° East (view map)
Paleocoordinates:1.7° North, 32.1° East
Basis of coordinate:based on political unit
Geographic resolution:local area
Time
Period:Neogene Epoch:Miocene
Stage:Burdigalian 10 m.y. bin:Cenozoic 5
*Period:Tertiary *Epoch:Early/Lower Miocene
Key time interval:Burdigalian
Age range of interval:20.44000 - 15.98000 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Stratigraphic resolution:member
Stratigraphy comments: "localities I, IV, V, and IX are intercalated with volcanic tuffs... and biotites from these tuffs at Napak I have produced dates of 19.5 +/- 2 Ma and 18.3 +/- 0.4 Ma" after being "adjusted for new constants" but "the younger date is discrepantly low" so an age of "between 19 and 20 Ma" (i.e., Burdigalian) is favored (Rossie and MacLatchy 2006)
Lithology and environment
Primary lithology: not reported
Environment:"floodplain"
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils,mesofossils
Collection methods and comments
Collection excludes:species names
Collection methods:field collection
Reason for describing collection:general faunal/floral analysis
Collectors:W. W. Bishop Collection dates:1958 - 1964
Collection method comments: Includes subcollections:I, IV, V, IX, X
Metadata
Also known as:Napak I, IV, V, IX, X
Database number:28441
Authorizer:J. Alroy, M. Uhen, P. Holroyd Enterer:J. Wertheim, M. Uhen, P. Holroyd, J. Alroy
Modifier:M. Uhen Research group:vertebrate
Created:2003-02-13 15:20:10 Last modified:2013-06-27 14:29:02
Access level:the public Released:2003-02-13 15:20:10
Creative Commons license:CC BY
Reference information

Primary reference:
4810.ETE 519W. W. Bishop and J. D. Clark. 1965. Background to Evolution in Africa. University of Chicago Press, Chicago [A. Behrensmeyer/A. Behrensmeyer/M. Kosnik]

Secondary references:

29409 J. G. Fleagle and E. L. Simons. 1978. Micropithecus clarki, a small ape from the Miocene of Uganda. American Journal of Physical Anthropology 49(4):427-440 [J. Alroy/J. Alroy/J. Alroy]
47356 J. Morales, M. Pickford, and D. Soria. 2007. New carnivoran material (Creodonta, Carnivora and Incertae sedis) from the Early Miocene of Napak, Uganda. Paleontological Research 11(1):71-84 [M. Uhen/M. Uhen]
67427 J. Morales, M. Pickford, and A. Valenciano. 2016. Systematics of African Amphicyonidae, with descriptions of new material from Napak (Uganda) and Grillental (Namibia). Journal of Iberian Geology 42(2):131-150 [P. Holroyd/P. Holroyd]
64590 J. Morales and M. Pickford. 2017. New Hyaenodonts (Ferae, Mammalia) from the early Miocene of Napak (Uganda), Koru (Kenya) and Grillental (Namibia). Fossil Imprint 73(3-4):332-359 [P. Holroyd/P. Holroyd]
78325% 23740M. Pickford, B. Senut, D. Hadoto, J. Musisi, and C. Kariira. 1986. Nouvelles decouvertes dans le Miocene inferiur de Napak, Ouganda Oriental [New discoveries in lower Miocene strata at Napak, Eastern Uganda]. Comptes Rendus de l'Academie des Sciences, Serie 2 302:47-52 [J. Alroy/J. Wertheim/J. Alroy]
29411 J. B. Rossie and L. MacLatchy. 2006. A new pliopithecoid genus from the early Miocene of Uganda. Journal of Human Evolution 50(5):568-586 [J. Alroy/J. Alroy]
47344 R. J. G. Savage. 1965. Fossil Mammals of Africa: 19 The Miocene Carnivora of East Africa. Bulletin of the British Museum (Natural History) Geology 10(8):241-316 [M. Uhen/M. Uhen]