Laño L1A: Late/Upper Campanian, Spain
List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Reptilia
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Titanosaurinae indet.
Lydekker 1893
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Sanz 1986 | 461 specimens | ||||||||
orig. 1 tooth | ||||||||||
= Titanosauridae indet.
Lydekker 1885
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Astibia et al. 1990 | |||||||||
= Lirainosaurus astibiae n. gen., n. sp.
Sanz et al. 1999
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Pereda-Suberbiola et al. 2000 | |||||||||
Holotype: MCNA 7458, anteriormost caudal vertebra (Sanz et al., 1999: pl. 2; Díez Díaz et al., 2013: fig. 4). Paratypes: an incomplete braincase (MCNA 7439), two isolated teeth (MCNA 7440, 7441), ten dorsal vertebrae (MCNA 7442-7450, 7456), seven caudal vertebrae (MCNA 1812, 7451-7457), one scapula (MCNA 7459), one coracoid (MCNA 7460), one sternal plate (MCNA 7461), four humeri (MCNA 7462-7465), a fragment of ilium (MCNA 7466), a fragment of a pubis (MCNA 7467), three femora (MCNA 3160, 7468, 7470), a tibia (MCNA 7471), a fibula (MCNA 7472) and two osteoderms (MCNA 14473, 14474) (figured in Sanz et al., 1999). | ||||||||||
Reptilia
- Nodosauridae
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Nodosauridae indet.
Marsh 1890
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Astibia et al. 1990 | 1 specimen | ||||||||
Struthiosaurus sp.
Bunzel 1870
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Pereda-Suberbiola 1999 | |||||||||
a partial skeleton, including the synsacrum, portions of the pelvic girdle and hindlimbs, a dorsal vertebra, dorsal rib fragments and a dermal scute | ||||||||||
= cf. Struthiosaurus austriacus
Bunzel 1871
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Le Loeuff and Buffetaut 1995 | |||||||||
Reptilia
- Rhabdodontidae
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Ornithopoda indet.
Marsh 1881
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Astibia et al. 1987 | |||||||||
= Rhabdodon priscus
Matheron 1869
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Astibia et al. 1990 | |||||||||
= Rhabdodon sp.
Matheron 1869
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Pereda-Suberbiola and Sanz 1999 | |||||||||
four isolated teeth, three from the maxilla and one from the dentary, two dorsal vertebrae, four caudal vertebrae, one left humerus, left and right femora | ||||||||||
Reptilia
- Hadrosauridae
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Hadrosauridae indet.
Cope 1869
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Pereda-Suberbiola et al. 2000 | |||||||||
MCNA 10510, tooth | ||||||||||
Reptilia
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Coelurosauria indet.
Huene 1914
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Canudo and Ruiz-Omeñaca 2003 | |||||||||
cf. Paraves informal indet. A
Sereno 1997
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Isasmendi et al. 2022 | 7 specimens | ||||||||
teeth, MCNA 14523–14529 | ||||||||||
cf. Paraves informal indet. B
Sereno 1997
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Isasmendi et al. 2022 | 31 specimens | ||||||||
teeth, MCNA 14530–14546, 14548–14561 | ||||||||||
cf. Paronychodon sp.
Cope 1876
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Isasmendi et al. 2022 | 6 specimens | ||||||||
teeth, MCNA 14547, 14562–14565, 16755 | ||||||||||
cf. Euronychodon sp.
Antunes and Sigogneau-Russell 1991
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Pereda-Suberbiola et al. 2000 | |||||||||
synonym of Paronychodon | ||||||||||
Reptilia
- Dromaeosauridae
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Theropoda indet.
Marsh 1881
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Astibia et al. 1990 | |||||||||
= Dromaeosauridae indet.
Colbert and Russell 1969
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Pereda-Suberbiola et al. 2000 | |||||||||
3 forms (A, B,and C) | ||||||||||
Dromaeosauridae indet.
Colbert and Russell 1969
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Isasmendi et al. 2022 | 4 specimens | ||||||||
teeth, MCNA 14623–14626 | ||||||||||
cf. Richardoestesia informal sp. A
Currie et al. 1990
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Isasmendi et al. 2022 | 35 specimens | ||||||||
teeth, MCNA 14566–14610, 14612–14622, 16768, 16779, 16780, 16786–16788, 16790, 16803, 16804, 16816–16823, 16825–16830, 16832, 16833, 16837, 16841, 16846–16849, 16851, 16852, 16857–16861 | ||||||||||
cf. Richardoestesia informal sp. B
Currie et al. 1990
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Isasmendi et al. 2022 | 53 specimens | ||||||||
MCNA 16765–16767, 16769–16778, 16781–16785, 16789, 16791–16802, 16805–16807, 16814, 16815, 16824, 16831, 16834–16836, 16838–16840, 16842–16845, 16850, 16853–16856 | ||||||||||
Reptilia
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Ornithomimosauria indet.
Barsbold 1976
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Pereda-Suberbiola et al. 2000 | |||||||||
= ? Ornithomimosauria indet.
Barsbold 1976
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Canudo and Ruiz-Omeñaca 2003 | |||||||||
Reptilia
- Abelisauridae
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? Abelisauridae indet.
Bonaparte and Novas 1985
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Astibia et al. 1990 | |||||||||
cf. Tarascosaurus sp.
Le Loeuff and Buffetaut 1991
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Le Loeuff and Buffetaut 1991 | 2 specimens | ||||||||
femora, MHNV | ||||||||||
= Neoceratosauria indet.
Novas 1991
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Pereda-Suberbiola et al. 2000 | |||||||||
= Abelisauridae indet.
Bonaparte and Novas 1985
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Isasmendi et al. 2021 | |||||||||
femora (missing) and MCNA 14077 (caudal vertebra) | ||||||||||
Arcovenator sp.
Tortosa et al. 2014
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Isasmendi et al. 2022 | 32 specimens | ||||||||
teeth, MCNA 1852, 1853, 1855, 2205, 2206, 8589, 8600, 10082, 14067, 14073, 14074, 14520–14522, 16756–16764, 16808–16813, 16862–16864 | ||||||||||
Reptilia
- Azhdarchidae
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Azhdarchidae indet.
Nessov 1984
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Astibia et al. 1990 | 295 specimens | ||||||||
= cf. Azhdarcho sp.
Nessov 1981
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Pereda-Suberbiola et al. 2000 | |||||||||
Reptilia
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Ischyrochampsa sp.
Vasse 1995
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Pereda-Suberbiola et al. 2000 | |||||||||
nomen dubium belonging to Neosuchia | ||||||||||
Alligatoridae indet.
Gray 1844
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Astibia et al. 1987 | |||||||||
= Acynodon iberoccitanus
Buscalioni et al. 1997
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Pereda-Suberbiola et al. 2000 | |||||||||
Crocodylidae indet.
Cuvier 1807
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Astibia et al. 1990 | 52 specimens | ||||||||
may be Allodaposuchus | ||||||||||
= aff. Allodaposuchus sp.
Nopcsa 1928
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Pereda-Suberbiola et al. 2000 | |||||||||
= Allodaposuchus precedens
Nopcsa 1928
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Buscalioni et al. 2001 | |||||||||
= Allodaposuchus sp.
Nopcsa 1928
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Puértolas-Pascual et al. 2016 | |||||||||
? Trematochampsidae indet.
Buffetaut 1974
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Astibia et al. 1990 | |||||||||
= Musturzabalsuchus buffetauti
Buscalioni et al. 1997
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Pereda-Suberbiola et al. 2000 | |||||||||
Helochelydra nopcsai
Lapparent de Broin and Murelaga 1999
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Lapparent de Broin and Murelaga 1999 | |||||||||
partial shell, lots of fragments | ||||||||||
Cryptodira indet.
Cope 1868
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Astibia et al. 1987 | |||||||||
= Dermatemydidae indet.
Baur 1888
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Astibia et al. 1990 | |||||||||
"Dermatemyididae" sic | ||||||||||
= Solemys vermiculata
Lapparent de Broin and Murelaga 1996
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Joyce 2017 | |||||||||
Solemys vermiculata n. sp.
Lapparent de Broin and Murelaga 1996
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Lapparent de Broin and Murelaga 1996 | 260 specimens | ||||||||
hypodigme around 260 shell fragments, most of them are peripherals and pleural fragments, around 25 fragments of plastron including the hologype | ||||||||||
Reptilia
- Testudines
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Dortoka vasconica n. gen., n. sp.
Lapparent de Broin and Murelaga 1996
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Lapparent de Broin and Murelaga 1996 | |||||||||
Polysternon atlanticum n. sp.
Lapparent de Broin and Murelaga 1996
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Lapparent de Broin and Murelaga 1996 | 1 specimen | ||||||||
recombined as Iberoccitanemys atlanticum | ||||||||||
? Elochelys sp.
Nopsca 1931
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Lapparent de Broin and Murelaga 1999 | |||||||||
right partial epiplastron | ||||||||||
Reptilia
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Scincomorpha informal indet. A
Camp 1923
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Astibia et al. 1990 | |||||||||
= Scincomorpha indet.
Camp 1923
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Rage 1999 | |||||||||
two fragments of unidentified bones with two teeth, and one isolated tooth | ||||||||||
Reptilia
- Lacertilia
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Lacertilia informal indet. 1
Owen 1842
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Rage 1999 | |||||||||
numerous remains (teeth) | ||||||||||
Lacertilia informal indet. 2
Owen 1842
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Rage 1999 | |||||||||
teeth | ||||||||||
Lacertilia informal indet. 3
Owen 1842
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Rage 1999 | |||||||||
teeth | ||||||||||
Lacertilia indet.
Owen 1842
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Rage 1999 | |||||||||
vertebrae | ||||||||||
Reptilia
- Madtsoiidae
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Aniliidae indet.
Fitzinger 1826
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Astibia et al. 1990 | |||||||||
= Herensugea caristiorum
Rage 1996
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Rage 1999 | |||||||||
17 vertebrae, perhaps one poorly preserved vertebra and perhaps one caudal vertebra | ||||||||||
Madtsoia sp.
Simpson 1933
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Astibia et al. 1990 | |||||||||
= Madtsoia laurasiae
Rage 1996
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Rage 1999 | |||||||||
recombined as Menarana laurasiae | ||||||||||
8 more or less complete vertebrae, and perhaps 2 fragments | ||||||||||
Reptilia
- Iguanidae
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Scincomorpha informal indet. B
Camp 1923
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Astibia et al. 1990 | |||||||||
= Iguanidae indet.
Oppel 1811
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Rage 1999 | |||||||||
one fragment of maxilla with four teeth, two fragments of unidentified bones with two teeth, and two fragments of unidentified bone with one tooth | ||||||||||
Reptilia
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Amphisbaenia indet.
Gray 1844
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Astibia et al. 1990 | |||||||||
? Amphisbaenia indet.
Gray 1844
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Rage 1999 | |||||||||
10 vertebrae | ||||||||||
Mammalia
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Theria informal indet. B
Parker and Haswell 1897
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Astibia et al. 1990 | |||||||||
Eutheria indet.
(Huxley 1880)
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Gheerbrant and Astibia 1999 | |||||||||
Mammalia
- Zhelestidae
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Lainodon informal new species
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Gheerbrant and Astibia 2012 | |||||||||
L1AT 18 (talonid of left M/3) and L1AT 13 and (talonid of right M/1 or M/2) | ||||||||||
Lainodon orueetxebarriai n. gen., n. sp.
Gheerbrant and Astibia 1994
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Gheerbrant and Astibia 1994 | |||||||||
? Labes sp.
Pol et al. 1992
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Gheerbrant and Astibia 1999 | |||||||||
new species | ||||||||||
= Lainodon orueetxebarriai
Gheerbrant and Astibia 1994
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Gheerbrant and Astibia 2012 | |||||||||
L1AT 17, talonid of left M/1 or M/2 (fig. 2); L1AT 4, disto-labial fragment of left upper molar; and possibly L1AT11, damaged right M/1 or M/2 | ||||||||||
Lainodon cf. orueetxebarriai
Gheerbrant and Astibia 1994
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Gheerbrant and Astibia 2012 | |||||||||
L1AT 20 (distal part of right lower premolar) | ||||||||||
Theria informal indet. A
Parker and Haswell 1897
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Astibia et al. 1990 | |||||||||
may be Marsupialia | ||||||||||
= Lainodon sp.
Gheerbrant and Astibia 1994
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Gheerbrant and Astibia 1999 | |||||||||
= Lainodon ragei n. sp.
Gheerbrant and Astibia 2012
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Gheerbrant and Astibia 2012 | |||||||||
L1AT 15 - type (right dP/5); referred specimens: L1AT 5: left dP/5; L1AT 15: right dP/5; L1AT 16: left dP5/; L1AT 8, left M/1 (trigonid); L1AT 9, damaged right upper molar (protocone) | ||||||||||
Amphibia
- Temnospondyli
- Albanerpetontidae
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cf. Albanerpeton sp.
Estes and Hoffstetter 1976
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Astibia et al. 1990 | |||||||||
= Albanerpetontidae indet.
Fox and Naylor 1982
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Duffaud and Rage 1999 | |||||||||
five incomplete dentaries, distal part of 3 humeri | ||||||||||
Amphibia
- Caudata
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Salamandridae indet.
Gray 1825
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Astibia et al. 1990 | |||||||||
may be Salamandra | ||||||||||
= Caudata indet.
Scopoli 1777
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Duffaud and Rage 1999 | |||||||||
1 atlantal centrum, 13 fragmentary vertebrae, 1 humerus | ||||||||||
Amphibia
- Temnospondyli
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Anura indet.
(Fischer von Waldheim 1813)
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Duffaud and Rage 1999 | |||||||||
distal part of 2 humeri, 8 ilia, fragmentary maxillaries, 2 broken sphenethmoids | ||||||||||
Amphibia
- Temnospondyli
- Palaeobatrachidae
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Palaeobatrachidae indet.
Cope 1865
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Duffaud and Rage 1999 | |||||||||
43 prearticulars, 1 presacral vert, 2 synsacrums, 6 distal parts of humeri | ||||||||||
Amphibia
- Temnospondyli
- Discoglossidae
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Discoglossidae indet.
Günther 1859
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Astibia et al. 1990 | |||||||||
6 prearticulars, 5 presacral verts, 7 sacral verts, 1 urostyle, 7 ilia, 2 humeri | ||||||||||
Actinopteri
- Lepisosteiformes
- Lepisosteidae
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Lepisosteus sp.
Agassiz 1843
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Astibia et al. 1990 | |||||||||
= Atractosteus sp.
Rafinesque 1820
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Pereda-Suberbiola et al. 2000 | |||||||||
Actinopteri
- Amiiformes
- Palaeolabridae
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? Palaeolabrus sp.
Casier 1967
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Pereda-Suberbiola et al. 2000 | |||||||||
Actinopteri
- Elopiformes
- Phyllodontidae
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Phyllodontidae indet.
Sauvage 1875
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Pereda-Suberbiola et al. 2000 | |||||||||
see common names |
Geography
Country: | Spain | State/province: | Castilla y Leon | County: | Burgos |
Coordinates: | 42.7° North, 2.6° West (view map) | ||||
Paleocoordinates: | 33.0° North, 0.1° West | ||||
Basis of coordinate: | estimated from map | ||||
Altitude: | 747 meters | ||||
Geographic resolution: | outcrop |
Time
Period: | Cretaceous | Epoch: | Late/Upper Cretaceous |
Stage: | Campanian | 10 m.y. bin: | Cretaceous 7 |
*Period: | Late/Upper Cretaceous | *Epoch: | Early/Lower Maastrichtian |
Key time interval: | Late/Upper Campanian | ||
Age range of interval: | 83.50000 - 70.60000 m.y. ago | ||
* legacy (obsolete) database fields |
Stratigraphy
Formation: | Vitoria | ||||
Local section: | Lano section | Local bed: | S1U3 | ||
Local order: | bottom to top | ||||
Stratigraphic resolution: | bed | ||||
Stratigraphy comments: in a sector of the south limb of the Miranda-Trevino syncline, level is also called L1A
lower pat of subchron C32n Also assigned to Sedano Fm. or its equivalent |
Lithology and environment
Primary lithology: | "cross stratification",hardground conglomeratic sandstone |
Secondary lithology: | black siltstone |
Includes fossils? | Y |
Includes fossils? | Y |
Lithology description: Multidirectional cross-laminated sands with microconglomerates in the bottom-sets, rarely more than 1m thick. Strata become massive toward the top, and are frequently intersected by abrasion surfaces showing an iron-mineralized crust or hard-ground. The most important bone accumulation is between two of these hardgrounds. The unit is topped by two black silt beds. | |
Environment: | coastal indet. |
Glacial or sequence phase: | regressive |
Geology comments: upper foreshore and backshore environments to a subcontinental, plant-rich perhaps swampy environment. The vertebrate bearing levels indicate a continental depositional environment affected by high-energy episodes, a fluviatile environment. |
Taphonomy
Modes of preservation: | body,permineralized,replaced with pyrite |
Size of fossils: | macrofossils,mesofossils,microfossils |
Disassociated minor elements: | all |
Collection methods and comments
Collection excludes: | difficult macrofossils |
Collection methods: | sieve,field collection |
Collection size: | 33 specimens |
Reason for describing collection: | taxonomic analysis |
Collection method comments: mostly chelonian shell fragments (30) |
Metadata
Also known as: | Laño Quarry | ||
Database number: | 28406 | ||
Authorizer: | J. Alroy, P. Mannion, M. Carrano | Enterer: | E. Leckey, M. Carrano, P. Mannion |
Modifier: | M. Carrano | Research group: | vertebrate |
Collections that are a subset of this one: | |||
Created: | 2003-02-12 10:11:45 | Last modified: | 2023-11-20 13:24:51 |
Access level: | the public | Released: | 2003-02-12 10:11:45 |
Creative Commons license: | CC BY |
Reference information
Primary reference:
55315. | B. Vila, M. Suñer, A. Santos-Cubedo, J. I. Canudo, B. Poza and A. Galobart. 2011. Saurischians through time. In A. Galobart, M. Suñer, & B. Poza (eds.), Dinosaurs of Eastern Iberia 130-168 [M. Carrano/M. Carrano] |
Secondary references:
7903 | H. Astibia, E. Buffetaut, A. D. Buscalioni, H. Cappetta, C. Corral, R. Estes, F. Garcia-Garmilla, J. J. Jaeger, E. Jiminez-Fuentes, J. Le Loeuff, J. M. Mazin, X. Orue-Etexebarria, J. Pereda-Suberbiola, J. E. Powell, J. C. Rage, J. Rodriguez-Lazaro, J. L. Sanz and H. Tong. 1990. The fossil vertebrates from the Lano (Basque Country, Spain); new evidence on the composition and affinities of the Late Cretaceous continental faunas of Europe. Terra Nova 2:460-466 [J. Alroy/E. Leckey/M. Uhen] | |
7885 | H. Astibia, F. Garcia-Garmilla, X. Orue-Etxebarria and J. Rodriguez-Lazaro. 1987. The Cretaceous-Tertiary boundary in a sector of the south limb of the Miranda-Trevino synclinal: the first appearance of Chelonia and Archosauria in the Basque Country. Cretaceous Research 8:15-27 [J. Alroy/E. Leckey/E. Leckey] | |
83613 | M. Aurell, D. Torromé, J. M. Gasca, P. Calvín, M. Pérez-Pueyo, J. Parrilla-Bel, E. Medrano-Aguado, C. Martín-Closas, A. Vicente, P. Sierra-Campos, and J. I. Canudo. 2022. Latest Cretaceous palaeogeographic evolution of northeast Iberia: Insights from the Campanian continental Montalbán subbasin (Spain). Earth-Science Reviews 235:10425 [M. Carrano/M. Carrano/P. Mannion] | |
86530 | G. Botfalvai, E. Prondvai, and A. 2021. Living alone or moving in herds? A holistic approach highlights complexity in the social lifestyle of Cretaceous ankylosaurs. Cretaceous Research 118:104633 [M. Carrano/M. Carrano] | |
14444 | ETE | A. D. Buscalioni, F. Ortega, D. B. Weishampel and C. M. Jianu. 2001. A revision of the crocodyliform Allodaposuchus precedens from the Upper Cretaceous of the Hateg Basin, Romania. Its relevance in the phylogeny of Eusuchia. Journal of Vertebrate Paleontology 21(1):74-86 [M. Carrano/M. Carrano/M. Carrano] |
17647 | ETE | J. I. Canudo and J. I. Ruiz-Omeñaca. 2003. Los restos directos de dinosaurios teropódos (excluyendo Aves) en España [Direct remains of theropod dinosaurs (excluding Aves) in Spain]. Ciencias de la Tierra 26:347-373 [M. Carrano/M. Carrano] |
32749 | ETE | J. Company, X. Pereda Suberbiola, and J. I. Ruiz-Omenaca. 2009. Nuevos restos fosiles del dinosaurio Lirainosaurus (Sauropoda, Titanosauria) en el Cretacico Superior (Campaniano-Maastrichtiano) de la Peninsula Iberica. Ameghiniana 46(2):391-405 [M. Carrano/M. Carrano/M. Carrano] |
19147 | ETE | F. M. Dalla Vecchia. 2006. Telmatosaurus and the other hadrosaurids of the Cretaceous European Archipelago. An overview. Natura Nascosta 32:1-55 [M. Carrano/M. Carrano] |
56054 | F. M. Dalla Vecchia. 2014. An overview of the latest Cretaceous hadrosauroid record in Europe. In D. A. Eberth & D. C. Evans (ed.), Hadrosaurs 268-297 [M. Carrano/M. Carrano] | |
38464 | V. Díez Díaz, X. Pereda Suberbiola, and J. L. Sanz. 2011. Braincase anatomy of the titanosaurian sauropod Lirainosaurus astibiae from the Late Cretaceous of the Iberian Peninsula. Acta Palaeontologica Polonica 56(3):521-533 [M. Carrano/M. Carrano] | |
43435 | V. Díez Díaz, X. Pereda Suberbiola, and J. L. Sanz. 2012. Juvenile and adult teeth of the titanosaurian dinosaur Lirainosaurus (Sauropoda) from the Late Cretaceous of Iberia. Geobios 45:265-274 [M. Carrano/M. Carrano] | |
46308 | V. Díez Díaz, X. Pereda Suberbiola, and J. L. Sanz. 2013. The axial skeleton of the titanosaur Lirainosaurus astibiae (Dinosauria: Sauropoda) from the latest Cretaceous of Spain. Cretaceous Research 43:145-160 [P. Mannion/J. Tennant/J. Tennant] | |
48052 | V. Díez Díaz, X. Pereda Suberbiola, and J. L. Sanz. 2013. Appendicular skeleton and dermal armour of the Late Cretaceous titanosaur Lirainosaurus astibiae (Dinosauria: Sauropoda) from Spain. Palaeontologia Electronica 16(2):19A [P. Mannion/J. Tennant/P. Mannion] | |
9509 | S. Duffaud and J.-C. Rage. 1999. Amphibians from the Upper Cretaceous of Lano (Basque Country, Spain). Estudios del Museo de Ciencias Naturales de Alava 14(1):111-120 [J. Alroy/E. Leckey/J. Head] | |
76795 | F. Escaso, A. Pérez-García, F. Ortega and J. L. Sanz. 2010. Ankylosaurian evidence from the Upper Cretaceous of South- Central Pyrenees (Lleida, Spain): a reappraisal. 8th Meeting of the European Association of Vertebrate Palaeontologists. Abstract Volume 33 [M. Carrano/M. Carrano] | |
7914 | E. Gheerbrant and H. Astibia. 1994. Un nouveau mammifere du Maastrichtien de Lano. Comptes Rendus de l'Academie des Sciences 318:1125-1131 [J. Alroy/E. Leckey/M. Uhen] | |
7791 | 5% 960 | E. Gheerbrant and H. Astibia. 1999. The Upper Cretaceous mammals from Lano (Spanish Basque Country). Estudios del Museo de Ciencias Naturales de Alava 14(1):295-323 [J. Alroy/E. Leckey/E. Leckey] |
44341 | E. Gheerbrant and H. Astibia. 2012. Addition to the Late Cretaceous Laño mammal faunule (Spain) and to the knowledge of European “Zhelestidae” (Lainodontinae nov.). Bulletin de la Société Géologique de France 183(6):537-546 [P. Mannion/P. Mannion] | |
84728 | E. Isasmendi, A. Torices, J. I. Canudo and X. Pereda-Suberbiola. 2021. Abelisaurid dinosaurs from the Upper Cretaceous Laño site (Iberian Peninsula). Ciências da Terra Procedia 1:38-41 [P. Mannion/P. Mannion/P. Mannion] | |
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