Las Hoyas (MCCM collection): Late/Upper Barremian, Spain

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Weitchselia reticulata
Pseudoastacus llopisi
Clavatoraxis robustus
Sanz et al. 2014
Turbomesodon praeclarus
Sanz et al. 2014
Iterophyllum lobatum
Sanz et al. 2014
Equisetopsida - Araucariaceae
Pagiophyllum sp. Heer 1881
Sanz et al. 2014
Brachyphyllum speciosa
spelled Brachyphilum
Equisetopsida - Cheirolepidiaceae
Cupressinocladus micromerum
Polypodiopsida
Cladophlebis browniana (Dunker 1846)
Polypodiopsida - Gleicheniales - Matoniaceae
Weichselia reticulata (Stokes and Webb 1824)
Sanz et al. 2014
Polypodiopsida - Schizaeales - Schizaeaceae
Ruffordia goepperti (Dunker 1844)
Sanz et al. 2014
original and current combination Cheilanthites goepperti
Angiospermae
Montsechia vidali (Zeiller 1902)
Bennettitales
Zamites sp. Brongniart 1828
Williamsonia sp. Carruthers 1870
spelled Williansonia
Pinopsida - Pinales - Taxodiaceae
Sphenolepis kurriana (Dunker 1846)
Bivalvia - Unionida - Unionidae
Unio sp. Philipsson 1788
Sanz et al. 2014
Insecta - Blattodea
Blattoidea indet. Handlirsch 1906
synonym of Blattaria
Insecta - Chresmodidae
Chresmoda aquatica Martínez-Delclòs 1989
Delclòs et al. 2008 6 specimens
LH-26.546, LH-13826, LH-13574, LH-92AM102, MCCM-LH 18023, MCCM-LH 18024,
Insecta - Hemiptera
Heteroptera indet. Latreille 1810
Insecta - Hemiptera - Naucoridae
Naucoridae indet. Fallen 1814
Insecta - Hemiptera - Corixidae
Velocorixinae indet. Popov 1986
Insecta - Hemiptera - Belostomatidae
Iberonepa roemerali Martínez-Delclòs et al. 1995
Sanz et al. 2014
Insecta - Hemiptera - Notonectidae
Notonectidae indet. (Latreille 1802)
original and current combination Notonectariae
Insecta - Hymenoptera - Scoliidae
Cretoscolia conquensis n. sp. Rasnitsyn and Martínez-Delclòs 2000
Rasnitsyn and Martínez-Delclòs 2000 1 specimen
MCCM LH 17331 (1 measurement)
Archaeoscolia hispanica n. sp. Rasnitsyn and Martínez-Delclòs 1999
Rasnitsyn and Martínez-Delclòs 1999 1 specimen
MCCM LH 13827 (2 measurements)
Insecta - Mecoptera - Orthophlebiidae
Orthophlebiinae indet. (Handlirsch 1906)
corrected as Orthophlebiidae
Insecta - Raphidioptera - Mesoraphidiidae
Mesoraphidiidae indet. Martynov 1925
Insecta - Neuroptera - Mesochrysopidae
Triangulochrysopa sanzi n. gen., n. sp. Nel et al. 2005
Nel et al. 2005 5 specimens
MCCM LH-8110, 18572, 18573, 18574, MCCM 92/2/3 (9 measurements)
Chimerochrysopa incerta n. gen., n. sp. Nel et al. 2005
Nel et al. 2005 1 specimen
MCCM LH 18588 (1 measurement)
Allopterus mayorgai n. sp. Nel et al. 2005
Nel et al. 2005 2 specimens
MCCM LH 18570 (5 measurements)
Armandochrysopa inexpecta n. sp. Nel et al. 2005
Nel et al. 2005 1 specimen
recombined as Kareninoides inexpectus
MCCM LH 18575 (3 measurements)
Tachinymphes paicheleri n. sp. Nel et al. 2005
Nel et al. 2005 10 specimens
MCCM LH 18576-8, LH 8040, LH 13175, MCCM LH 18579-18584 (7 measurements)
Tachinymphes penalveri n. sp. Nel et al. 2005
Nel et al. 2005 5 specimens
MCCM LH 18585, LH 18586-7, U.A. Madrid LH 8033, 8094 (8 measurements)
Nanochrysopa pumilio n. gen., n. sp. Nel et al. 2005
Nel et al. 2005 1 specimen
MCCM LH 13217 (2 measurements)
Insecta - Coleoptera - Coptoclavidae
Ovonectes pilosum n. gen., n. sp. Soriano et al. 2007
Soriano et al. 2007 2 specimens
MCCM-LH23853, 24514 (2 measurements)
Hispanoclavina diazromerali n. gen., n. sp. Soriano et al. 2007
Soriano et al. 2007 5 specimens
MCCM-LH16264, 15070, 16315, 22299, 23859 (2 measurements)
Hispanoclavina gratshevi n. sp. Soriano et al. 2007
Soriano et al. 2007 3 specimens
MCCM-LH15961, 15747, 23000 (2 measurements)
Megacoptoclava longiurogomphia n. gen., n. sp. Ponomarenko and Martínez-Delclòs 2000
Ponomarenko and Martínez-Delclòs 2000 1 specimen
MCCM LH-1263
Hoyaclava buscalionae n. gen., n. sp. Soriano et al. 2007
Soriano et al. 2007 50 specimens
MCCM-LH24508, 7269, 23060, 24517 (2 measurements)
Insecta - Coleoptera - Ademosynidae
Ademosynidae indet. Ponomarenko 1968
Insecta - Coleoptera
Archostemata indet. Kolbe 1908
Zygadenia diazromerali n. sp. Soriano and Delclos 2006
Soriano and Delclos 2006 1 specimen
recombined as Notocupes diazromerali
MCCM LH 12001 (4 measurements)
Zygadenia siniestri n. sp. Soriano and Delclos 2006
Soriano and Delclos 2006 3 specimens
recombined as Conexicoxa siniestri
MCCM LH 23515, MCCM LH 21291, MCCM LH 15819 (4 measurements)
Zygadenia martinclosas Soriano and Delclos 2006
Soriano and Delclos 2006 1 individual
recombined as Conexicoxa martinclosas
MCCM LH 20151
Zygadenia longicoxa n. sp. Soriano and Delclos 2006
Soriano and Delclos 2006 2 specimens
recombined as Brachilatus longicoxa
MCCM LH 928030, MCCM LH 23432
Zygadenia viridis n. sp. Soriano and Delclos 2006
Soriano and Delclos 2006 1 specimen
recombined as Brachilatus viridis
MCCMLH 1805 (2 measurements)
Insecta - Coleoptera - Cupedidae
Anaglyphites zherikhini n. sp. Soriano and Delclos 2006
Soriano and Delclos 2006 1 specimen
recombined as Mesocupes zherikhini
MCCM LH 15122 (2 measurements)
Priacma sanzii n. sp. Soriano and Delclos 2006
Soriano and Delclos 2006 1 specimen
nomen dubium belonging to Cupedinae
MCCM LH 24524 (2 measurements)
Insecta - Coleoptera - Ommatidae
Tetraphalerus ponomarenkoi n. sp. Soriano and Delclos 2006
Soriano and Delclos 2006 1 specimen
MCCM LH 78 (2 measurements)
Insecta - Coleoptera - Parandrexidae
Martynopsis laticollis n. gen., n. sp. Soriano et al. 2006
Soriano et al. 2006 2 specimens
MCCM LH-21091, 24513 (2 measurements)
Insecta - Coleoptera - Nemonychidae
Distenorrhinus (Parabrenthorrhinus) xavieri n. sp. Zherikhin and Gratshev 2003
Zherikhin and Gratshev 2003 1 specimen
MCCM LH 17330 (1 measurement)
Insecta - Odonata - Aeschnidiidae
Gigantoaeschnidium ibericus n. gen., n. sp. Nel and Martínez-Delclòs 1993
Fleck and Nel 2003 6 specimens
(6 measurements)
Sinaeschnidia martinezdelclosi n. sp. Fleck and Nel 2003
Fleck and Nel 2003 2 specimens
LH 17332, ADR-317-I (2 measurements)
Iberoaeschnidium conquensis n. gen., n. sp. Nel and Martínez-Delclòs 1993
Fleck and Nel 2003 1 specimen
LH-083-R (1 measurement)
Insecta - Ephemeroptera
Ephemeroptera indet. Hyatt and Arms 1890
Insecta - Diptera - Ptychopteridae
Eoptychoptera sp. Handlirsch 1906
Lukashevich and Arillo 2016 1 specimen
MCCM-LH 35084
Insecta - Diptera - Tipulidae
Leptotarsus ibericus n. sp. Ribeiro and Lukashevich 2014
Ribeiro and Lukashevich 2014 3 specimens
MCCM LH 28206, 18122, 29995 (2 measurements)
Leptotarsus buscalioniae n. sp. Ribeiro and Lukashevich 2014
Ribeiro and Lukashevich 2014 1 specimen
MCCM LH 35088 (2 measurements)
Leptotarsus contractus n. sp. Ribeiro and Lukashevich 2014
Ribeiro and Lukashevich 2014 1 specimen
MCCM LH 15419 (2 measurements)
Arachnida - Opiliones
Opiliones indet. Sundevall 1833
Palencia et al. 2019 1 specimen
MUPA LH-29969
Malacostraca - Decapoda - Atyidae
Oplophorus roselli Via 1971
recombined as Delclosia roselli
Delclosia martinelli Rabada 1993
Sanz et al. 2014
Malacostraca - Decapoda - Astacidae
Austropotamobius llopisi (Via 1971)
Sanz et al. 2014
unclassified
Undichna unisulca de Gibert et al. 1999
Moratalla et al. 2017
Coelacanthimorpha - Coelacanthiformes - Latimeriidae
? Holophagus leridae
? "Holophagus"
Amphibia - Temnospondyli
Anura indet. (Fischer von Waldheim 1813)
Sanz et al. 1988 1 individual
Iberobatrachus angelae n. gen., n. sp. Baez 2013
Baez 2013
MCCM-LH 11393a-b - holotype (articulated specimen lacking the hindlimbs and left forelimb, two-dimensionally preserved as part and counterpart on two slabs); referred specimen - MCCM-LH 15999a-b (incomplete, articulated specimen preserved as part and counterpart on two slabs)
Gracilibatrachus avallei n. gen., n. sp. Baez 2013
Baez 2013
MCCM-LH 21171a, b - holotype (articulated skeleton of a postmetamorphic individual); referred specimens: MCCMLH-17422a, b, 11386, 11387, 26251, 26803a, b (in part), and, tentatively, 11394a, b
Hylaeobatrachus sp. Dollo 1884
Sanz et al. 2014
Amphibia - Temnospondyli - Urodela
Caudata indet. Scopoli 1777
    = Valdotriton gracilis n. gen., n. sp. Evans and Milner 1996
Evans and Milner 1996
MCCM-LH 2850a-b, 6021, 928001a-b, ADR unnumbered
Amphibia - Temnospondyli - Albanerpetontidae
Celtedens ibericus n. sp. McGowan and Evans 1995
McGowan and Evans 1995
LH 6020 - type (part and counterpart of a nearly complete skeleton, missing only part of the palate and tail); referred material (McGowan 2002): two complete specimens in part and counterpart (LH 030 R A & B; LH 15710 A & B)
Mammalia - Groeberida - Nototheriidae
Spinolestes xenarthrosus n. gen., n. sp. Martin et al. 2015
Martin et al. 2015
MCCM-LH 30000 - holotype (complete skeleton); MCCMLH30000B (counter slab)
Reptilia - Testudines
Toxochelyidae indet. Baur 1895
2 specimens
MCCM-LH 84, LH 254 (carapace fragment)
    = Hoyasemys jimenezi n. gen., n. sp. Pérez-García et al. 2012
Pérez-García et al. 2012
Holotype: MCCM-LH 84, an almost complete skeleton, missing the dorsal portion of the skull. The specimen is preserved on two slabs. One shows the ventral view of the skull, the five proximal cervical vertebrae, some dorsal vertebrae, some fragments of the peripherals and of the lat− eral part of the costal plates, the plastron, sacral and caudal vertebrae and the appendicular skeleton. The other slab preserves the partial carapace and some bones of the hindlimb.
Reptilia
Jucaraseps grandipes n. gen., n. sp. Bolet and Evans 2012
Bolet and Evans 2012 1 individual
LH 18505 (holotype), almost complete skeleton in dorsal view
Reptilia - Teiidae
Meyasaurus diazromerali n. sp. Evans and Barbadillo 1997
Evans and Barbadillo 1997 6 individuals
LH 370, LH 372, LH 33, LH 13510, LH 6026, LH 14317
Reptilia
Lacertilia indet. Owen 1842
2 individuals
    = Scandensia ciervensis n. gen., n. sp. Evans and Barbadillo 1998
Evans and Barbadillo 1998
Hoyalacerta sanzi n. gen., n. sp. Evans and Barbadillo 1999
Evans and Barbadillo 1999 1 individual
LH 11000 (holotype skeleton)
Crocodylomorpha indet. (Hay 1930)
Sanz et al. 2014
Reptilia - Gobiosuchidae
Gobiosuchidae indet. Osmolska 1972
Sanz et al. 2014
    = Cassissuchus sanziuami n. gen., n. sp. Buscalioni 2017
Buscalioni 2017
MCCM-LH 7991 - holotype; referred specimen: MCCMLH 6508
Reptilia
Montsecosuchus sp. Buscalioni and Sanz 1988
Sanz et al. 2014
Reptilia - Atoposauridae
Atoposauridae indet. (Gervais 1871)
Reptilia - Bernissartiidae
Bernissartidae indet. Dollo 1883
Sanz et al. 2014
corrected as Bernissartiidae
Reptilia
Mantellisaurus atherfieldensis (Hooley 1925)
Llandres Serrano et al. 2013
LH-30677, incomplete but fully-articulated right hindlimb
    = cf. Mantellisaurus sp. Paul 2007
Moratalla et al. 2017
Reptilia - Theropoda
Megalosauridae indet. (Fitzinger 1843)
Canudo and Ruiz-Omeñaca 2003
tooth
    = Theropoda indet. Marsh 1881
Canudo and Ruiz-Omeñaca 2003
Reptilia
? Pterosauria indet. (Kaup 1834)
Lockley et al. 1995 1 specimen
MCCM-LH-6500, isolated track
    = ? Theropoda indet. Marsh 1881
Vullo et al. 2009
Theropoda indet. (Marsh 1881)
Moratalla et al. 2017 1 individual
MCCM-LH-Y-1-001, trackway
Aves indet. Linnaeus 1758
Sanz et al. 2001 4 individuals
MCCM-LH 11386; 3 taxa in a pellet
Eoalulavis hoyasi n. gen., n. sp. Sanz et al. 1996
Sanz et al. 1996 1 specimen
Concornis lacustris n. gen., n. sp. Sanz and Buscalioni 1992
Sanz et al. 1995
MCCM-LH-2814
cf. Concornis lacustris Sanz and Buscalioni 1992
Cambra-Moo et al. 2006 2 specimens
MCCM-LH 21006 a-b, tibiotarsus and tarsometatarsus
cf. Iberomesornis sp. Sanz et al. 1990
Sanz and Buscalioni 1994
Aves indet. Linnaeus 1758
new genus and species
    = Iberomesornis romerali n. gen., n. sp. Sanz et al. 1990
Sanz et al. 1990
MCCM-LH 22R - holotype (partial skeleton)
Pelecanimimus polyodon n. gen., n. sp. Pérez-Moreno et al. 1994
Pérez-Moreno et al. 1994 1 individual
MCCM-LH 7777 (1 measurement)
Reptilia - Carcharodontosauridae
Concavenator corcovatus n. gen., n. sp. Ortega et al. 2010
Ortega et al. 2010 1 individual
MCCM-LH 6666 (5 measurements)
Reptilia
Ornithocheiridae indet. (Seeley 1870)
Vullo et al. 2009 2 elements
MCCM-LH-17264, LH-21451, teeth
Reptilia - Istiodactylidae
Istiodactylidae indet. Howse et al. 2001
Vullo et al. 2009 2 elements
MCCM-LH-28572, LH-15448, teeth
Reptilia
Europejara olcadesorum n. gen., n. sp. Vullo et al. 2012
Vullo et al. 2012 1 specimen
MCCM-LH 9413, jaws and post-orbital elements of the skull preserved on part and counterpart slabs
Actinopteri - Pycnodontiformes
Pycnodontiformes indet. Berg 1937
genus incertae sedis
Actinopteri - Pycnodontiformes - Pycnodontidae
Macromesodon sp. Blake 1905
Wenz and Poyato-Ariza 1994
Coelodus subdiscus
Wenz and Poyato-Ariza 1994
Actinopteri - Pycnodontiformes
Eomesodon sp. Woodward 1918
Wenz and Poyato-Ariza 1994
Actinopteri - Semionotiformes - Macrosemiidae
Notagogus sp. Agassiz 1833
Notagogus ferreri
Actinopteri - Lepisosteiformes
Lepidotes sp. Agassiz 1832
Lepidotes microrbis
Sanz et al. 2014
Lepidotes tanyrhis n. sp. Wenz 2003
Wenz 2003
MCCM-LH-7410
Lepidotes microrhis n. sp. Wenz 2003
Wenz 2003
MCCM-LH-8010a,b
Actinopteri - Amiiformes - Caturidae
Caturus sp. Agassiz 1834
Actinopteri - Amiiformes - Amiidae
? Amiopsis sp. Kner 1863
Urocles woodwardi
synonym of Amiopsis
Vidalamia catalunica (Sauvage 1903)
Actinopteri - Gonorhynchiformes - Chanidae
Rubiesichthys gregalis Wenz 1984
Actinopteri
Gordichthys conquensis n. gen., n. sp. Poyato-Ariza 1994
Poyato-Ariza 1994
holotype: LH 1288 R
Actinopteri - Leptolepidiformes - Leptolepididae
? Ascalabos voithi
synonym of Leptolepis
Actinopteri - Pholidophoriformes - Pleuropholidae
Pleuropholis sp. Egerton 1858
unclassified
Planolites sp. Nicholson 1873
Moratalla et al. 2017
Treptichnus sp. Miller 1889
Moratalla et al. 2017
Lockeia sp. James 1879
Moratalla et al. 2017
Cruziana sp. d'Orbigny 1842
Moratalla et al. 2017
Palaeophycus sp. Hall 1847
Moratalla et al. 2017
Helminthoidichnites sp. Fitch 1850
Moratalla et al. 2017
see common names

Geography
Country:Spain State/province:Castilla-La Mancha County:Cuenca
Coordinates: 40.1° North, 1.9° West (view map)
Paleocoordinates:30.5° North, 8.1° East
Basis of coordinate:estimated from map
Altitude:428 meters
Geographic resolution:outcrop
Time
Period:Cretaceous Epoch:Early/Lower Cretaceous
Stage:Barremian 10 m.y. bin:Cretaceous 2
*Period:Early/Lower Cretaceous *Epoch:Neocomian - Gallic
*International age/stage:Late/Upper Hauterivian - Early/Lower Barremian
Key time interval:Late/Upper Barremian
Age range of interval:130.00000 - 125.45000 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Formation:La Huérguina
Local section:Las Hoyas Local bed:3
Local order:bottom to top
Stratigraphic resolution:group of beds
Stratigraphy comments: also called "Calizas de La Huérgina Formation" or "La Huérgina Limestones Formations"
Lithology and environment
Primary lithology:planar lamination "limestone"
Secondary lithology: packstone
Includes fossils?Y
Lithology description: Limestone with laminates; three facies described: Massive limestone, packstones with peloids and sand-sized fragments of Charophyte stems. Occasionally, in the base of these beds, levels of gastropodal wackestones appear; Flaggy limestones, sometimes nodular. Ostracodal mudstones with occasional thick laminations, laterally grading to confolute laminations; Laminated limestones, mudstones with laminations containing abundant fossils. Laminae are very continuous laterally
Environment:lacustrine - large Tectonic setting:rift
Geology comments: shallow to deep lacustrine facies; "produced in the context of a continental (freshwater) subtropical, seasonal summer wet, carbonate wetland that overlay a low-relief karstic terrain. The wetland was drained by carbonate-rich freshwater and comprised a typical environmental mosaic of swampy plains, ponds, lakes, ephemeral channels and sloughs."
Taphonomy
Modes of preservation:body,adpression,trace,soft parts
Lagerst�tten type:conservation
Degree of concentration:concentrated
Size of fossils:macrofossils,mesofossils
Preservation of anatomical detail:excellent
Articulated whole bodies:many
Associated major elements:some
Disassociated major elements:some
Disassociated minor elements:some
Spatial resolution:parautochthonous
Collection methods and comments
Collection excludes:some macrofossils,some microfossils
Collection methods:selective quarrying,surface (in situ),field collection
Reason for describing collection:general faunal/floral analysis
Collection method comments: Repository: Museo de las Ciencias de Castilla-La Mancha, Cuenca (MCCM)
Metadata
Database number:28006
Authorizer:J. Alroy, M. Bell, M. Clapham, R. Benson, M. Carrano, P. Mannion, R. Butler Enterer:E. Leckey, M. Carrano, J. Alroy, J. Tennant, M. Oreska, M. Bell, R. Benson, M. Clapham, J. Karr, R. Butler, P. Mannion
Modifier:M. Carrano Research group:paleoentomology,vertebrate
Created:2003-01-20 12:47:22 Last modified:2021-01-22 10:43:48
Access level:the public Released:2003-01-20 12:47:22
Creative Commons license:CC BY
Reference information

Primary reference:

7671.5% 14640J. L. Sanz, S. Wenz, A. Yebenes, R. Estes, X. Martinez-Delclos, E. Jiminez-Fuentes, C. Dieguez, A. D. Buscalioni, L. J. Barbadillo and L. Via. 1988. An Early Cretaceous faunal and floral continental assemblage: Las Hoyas fossil site (Cuenca, Spain). Géobios 21(5):611-635 [J. Alroy/E. Leckey/J. Alroy]

Secondary references:

44133 A. M. Baez. 2013. Anurans from the Early Cretaceous Lagerstätte of Las Hoyas, Spain: New evidence on the Mesozoic diversification of crown-clade Anura. Cretaceous Research 41:90-106 [P. Mannion/P. Mannion/P. Mannion]
35206 G. Bechly, A. Nel, X. Martínez-Declòs, E. A. Jarzembowski, R. Coram, D. Martill, G. Fleck, F. Escuillié, M. M. Wisshak and M. Maisch. 2001. A revision and phylogenetic study of Mesozoic Aeshnoptera, with description of numerous new taxa (Insecta: Odonata: Anisoptera). Neue Paläontologische Abhandlungen 4:1-219 [M. Clapham/M. Clapham/M. Clapham]
41482 A. Bolet and S. E. Evans. 2012. A tiny lizard (Lepidosauria, Squamata) from the Lower Cretaceous of Spain. Palaeontology 55(3):491-500 [R. Benson/R. Benson]
62518 A. D. Buscalioni. 2017. The Gobiosuchidae in the early evolution of Crocodyliformes. Journal of Vertebrate Paleontology 37(3):e1324459:1-21 [P. Mannion/P. Mannion/P. Mannion]
17880ETE A. D. Buscalioni and M. Frenegal-Martinez. 2006. Archosaurian size bias in Jurassic and Cretaceous freshwater ecosystems. 9th International Symposium on Mesozoic Terrestrial Ecosystems and Biota, Abstracts and Proceedings Volume 9-12 [M. Carrano/M. Carrano/M. Carrano]
34089ETE O. Cambra-Moo, A. D. Buscalioni, J. Cubo, J. Castanet, M.-M. Loth, E. Margerie, and A. Ricqlès. 2006. Histological observations of enantiornithine bone (Saurischia, Aves) from the Lower Cretaceous of Las Hoyas (Spain). Comptes Rendus Palevol 5:685-691 [M. Carrano/M. Oreska/M. Carrano]
17647ETE J. I. Canudo and J. I. Ruiz-Omeñaca. 2003. Los restos directos de dinosaurios teropódos (excluyendo Aves) en España [Direct remains of theropod dinosaurs (excluding Aves) in Spain]. Ciencias de la Tierra 26:347-373 [M. Carrano/M. Carrano]
55113 E. Cuesta, I. Díaz-Martínez, F. Ortega and J. L. Sanz. 2015. Did all theropods have chicken-like feet? First evidence of a non-avian dinosaur podotheca. Cretaceous Research 56(1):53-59 [M. Carrano/M. Carrano]
41244 X. Delclòs, A. Nel, D. Azar, G. Bechly, J. A. Dunlop, M. S. Engel, and S. W. Heads. 2008. The enigmatic Mesozoic insect taxon Chresmodidae (Polyneoptera): new palaeobiological and phylogenetic data, with the description of a new species from the Lower Cretaceous of Brazil. Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen 247:353-381 [M. Clapham/M. Clapham/M. Clapham]
37280 S. E. Evans and L. J. Barbadillo. 1997. Early Cretaceous lizards from Las Hoyas, Spain. Zoological Journal of the Linnean Society 119:23-49 [R. Benson/R. Benson]
26529ETE S. E. Evans and L. J. Barbadillo. 1998. An unusual lizard (Reptilia: Squamata) from the Early Cretaceous of Las Hoyas, Spain. Zoological Journal of the Linnean Society 124:235-265 [M. Carrano/M. Carrano]
37784 S. E. Evans and L. J. Barbadillo. 1999. A short-limbed lizard from the Lower Cretaceous of Spain. In D. M. Unwin (ed.), Cretaceous Fossil Vertebrates. Special Papers in Palaeontology 60:73-85 [R. Benson/R. Benson/M. Carrano]
25847ETE S. E. Evans and A. R. Milner. 1996. A metamorphosed salamander from the Early Cretaceous of Las Hoyas, Spain. Philosophical Transactions of the Royal Society of London B 351:627-646 [M. Carrano/M. Carrano]
35239 G. Fleck, G. Bechly, X. Martinez-Delclos, E. Jarzembowski, R. Coram and A. Nel. 2003. Phylogeny and classification of the Stenophlebioptera (Odonata: Epiproctophora). Annales de la Société Entomologique de France 39(1):55-93 [M. Clapham/J. Karr]
35493 G. Fleck and A. Nel. 2003. Revision of the Mesozoic family Aeschnidiidae (Odonata: Anisoptera). Zoologica 153:1-170 [M. Clapham/J. Karr]
28227ETE N. S. Gupta, O. Cambra-Moo, D. E. G. Briggs, G. D. Love, M. A. Fregenal-Martinez and R. E. Summons. 2008. Molecular taphonomy of macrofossils from the Cretaceous Las Hoyas Formation, Spain. Cretaceous Research 29:1-8 [M. Carrano/M. Carrano]
42228 D. W. E. Hone, H. Tischlinger, E. Frey and M. Röper. 2012. A new non-pterodactyloid pterosaur from the Late Jurassic of southern Germany. PLoS ONE 7(7):e38900 [R. Benson/R. Benson]
45545 M. Llandres Serrano, R. Vullo, J. Marugan-Lobon, F. Ortega, and A. D. Buscalioni. 2013. An articulated hindlimb of a basal iguanodont (Dinosauria, Ornithopoda) from the Early Cretaceous Las Hoyas Lagerstätte (Spain). Geological Magazine 150(3):527-576 [P. Mannion/J. Tennant/M. Carrano]
31125 M. Lockley, T. J. Logue, J. J. Moratalla, A. P. Hunt, R. J. Schultz and J. W. Robinson. 1995. The fossil trackway Pteraichnus is pterosaurian, not crocodilian: implications for the global distribution of pterosaur tracks. Ichnos 4:7-20 [R. Butler/R. Butler]
56924 E. D. Lukashevich and A. Arillo. 2016. New Eoptychoptera (Insecta: Diptera, Ptychopteridae) from the Lower Cretaceous of Spain. Cretaceous Research 58:254-264 [M. Clapham/M. Clapham]
56864 T. Martin, J. Marugán-Lobón, R. Vullo, H. Martín-Abad, Z.-X. Luo and A. D. Buscalioni. 2015. A Cretaceous eutriconodont and integument evolution in early mammals. Nature 526:380-384 [P. Mannion/P. Mannion]
49227 X. Martínez-Delclòs. 1993. Blátidos (Insecta, Blattodea) del Cretácico Inferior de España. Familias Mesoblattinidae, Blattulidae y Poliphagidae. Boletín Geológico y Minero 104:516-538 [M. Clapham/M. Clapham]
45506 X. Martínez-Delclòs, A. Nel, and Y. A. Popov. 1995. Systematics and functional morphology of Iberonepa romerali n. gen. and sp., Belostomatidae from the Spanish Lower Cretaceous (Insecta, Heteroptera). Journal of Paleontology 69:496-508 [M. Clapham/M. Clapham]
44194 G. McGowan and S. E. Evans. 1995. Albanerpetontid amphibians from the Cretaceous of Spain. Nature 373:143-145 [P. Mannion/P. Mannion]
42672 G. J. McGowan. 2002. Albanerpetontid amphibians from the Lower Cretaceous of Spain and Italy: a description and reconsideration of their systematics. Zoological Journal of the Linnean Society 135(1):1-32 [M. Carrano/M. Carrano]
78477 J. J. Moratalla, J. Marugán-Lobón, H. Martín-Abad, E. Cuesta, and A. D. Buscalioni. 2017. A new trackway possibly made by a trotting theropod at the Las Hoyas fossil site (Early Cretaceous, Cuenca Province, Spain): Identification, bio-dynamics and palaeoenvironmental implications. Palaeontologia electronica 20(3):59A:1-14 [M. Carrano/M. Carrano]
8943ETE M. B. Mostovski and X. Martínez-Delclòs. 2000. New Nemestrinoidea (Diptera: Brachycera) from the Upper Jurassic - Lower Cretaceous of Eurasia, taxonomy, and palaeobiology. Entomological Problems 31(2):137-148 [C. Labandeira/A. Spencer-Lee/J. Karr]
39071 A. Nel, X. Delclos, and A. Hutin. 2005. Mesozoic chrysopid-like Planipennia: a phylogenetic approach (Insecta: Neuroptera). Annales de la Société Entomologique de France 41:29-69 [M. Clapham/M. Clapham/M. Clapham]
35441 A. Nel and X. Martínez-Delclòs. 1993. Nuevos Zygoptera y Anisoptera (Insecta: Odonata) en el Cretacico inferior de Espana. Estudios Geológicos 49:351-359 [M. Clapham/J. Karr/M. Clapham]
33846ETE F. Ortega, F. Escaco, and J. L. Sanz. 2010. A bizarre, humped Carcharodontosauria (Theropoda) from the Lower Cretaceous of Spain. Nature 467:203-206 [M. Carrano/M. Carrano/M. Carrano]
67846 L. Palencia, E. Peñalver, C. E. Prieto and F. J. Poyato-Ariza. 2019. First fossil harvestmen (Arachnida: Opiliones) from Spain and notes on the fossil record of Opiliones. Palaeontologia Electronica 22(1.5A):1-18 [M. Clapham/M. Clapham/M. Clapham]
42000 A. Pérez-García, M. S. de la Fuente, and F. Ortega. 2012. A new freshwater basal eucryptodiran turtle from the Early Cretaceous of Spain. Acta Palaeontologica Polonica 57(2):285-298 [R. Benson/R. Benson/P. Mannion]
12449ETE B. P. Pérez-Moreno, J. L. Sanz, A. D. Buscalioni, J. J. Moratalla, F. Ortega and D. Rasskin-Gutman. 1994. A unique multitoothed ornithomimosaur from the Lower Cretaceous of Spain. Nature 370:363-367 [M. Carrano/M. Carrano/M. Carrano]
61983 B. P. Pérez-Moreno and J. L. Sanz. 1997. Las Hoyas. In P. J. Currie & K. Padian (ed.), Encyclopedia of Dinosaurs 398-402 [M. Carrano/M. Carrano]
48667 A. Ponomarenko and X. Martínez-Delclòs. 2000. New beetles (Insecta: Coleoptera) from the Lower Cretaceous of Spain. Acta Geologica Hispanica 35:47-52 [M. Clapham/M. Clapham]
47043 F. J. Poyato-Ariza. 1994. A new Early Cretaceous Gonorynchiform fish (Teleostei: Ostariophysi) from Las Hoyas (Cuenca, Spain). Occassional papers of the Museum of Natural History, The University of Kansas, Lawrence, Kansas 164:1-37 [M. Bell/M. Bell]
39638 A. P. Rasnitsyn and X. Martínez-Delclòs. 1999. New Cretaceous Scoliidae (Vespida=Hymenoptera) from the Lower Cretaceous of Spain and Brazil. Cretaceous Research 20:767-772 [M. Clapham/M. Clapham]
39604 A. P. Rasnitsyn and X. Martínez-Delclòs. 2000. Wasps (Insecta: Vespida = Hymenoptera) from the Early Cretaceous of Spain. Acta Geologica Hispanica 35:65-95 [M. Clapham/M. Clapham/M. Clapham]
50631 G. C. Ribeiro and E. D. Lukashevich. 2014. New Leptotarsus from the Early Cretaceous of Brazil and Spain: the oldest members of the family Tipulidae (Diptera). Zootaxa 3753:347-363 [M. Clapham/M. Clapham]
55500 J. I. Ruiz-Omeñaca and J. I. Canudo. 2003. Dinosaurios (Saurischia, Ornithischia) en el Barremiense (Cretácico Inferior) de la península Ibérica [Dinosaurs (Saurischia, Ornithischia) in the Barremian (Lower Cretaceous) of the Iberian peninsula]. In F. Pérez Lorente (ed.), Dinosaurios y Otros Reptiles Mesozóicos de España 269-312 [M. Carrano/M. Carrano]
70303 J. L. Sanz. 2000. Yacimientos ibéricos de dinosaurios [Iberian dinosaur localities]. Dinosaurios: Los Señores del Pasado [Dinosaurs: The Lords of the Land] 267-299 [M. Carrano/M. Carrano]
13500ETE J. L. Sanz, J. F. Bonaparte, and A. Lacasa. 1988. Unusual Early Cretaceous birds from Spain. Nature 331:433-435 [M. Carrano/M. Carrano/M. Carrano]
19339ETE J. L. Sanz, A. D. Buscalioni, J. J. Moratalla, V. Francés, and M. Antón. 1990. Los reptiles Mesozoicos del registro español [The Mesozoic reptiles from the Spanish record]. Monografias del Museo Nacional de Ciencias Naturales 2:1-79 [M. Carrano/M. Carrano]
13505ETE J. L. Sanz and A. D. Buscalioni. 1994. Géobios, Mémoire Spéciale 16:213-217 [M. Carrano/M. Carrano/M. Carrano]
24393ETE J. L. Sanz, L. M. Chiappe, and A. D. Buscalioni. 1995. The osteology of Concornis lacustris (Aves: Enantiornithes) from the Lower Cretaceous of Spain and a reexamination of its phylogenetic relationships. American Museum Novitates 3133:1-23 [M. Carrano/M. Carrano]
13506ETE J. L. Sanz, L. M. Chiappe, B. P. Pérez-Moreno, A. D. Buscalioni, J. J. Moratalla, F. Ortega, and F. J. Poyato-Ariza. 1996. An Early Cretaceous bird from Spain and its implications for the evolution of avian flight. Nature 382:442-445 [M. Carrano/M. Carrano/M. Carrano]
25164ETE J. L. Sanz, L. M. Chiappe, Y. Fernández-Jalvo, F. Ortega, B. Sánchez-Chillón, F. J. Poyato-Ariza, and B. P. Pérez-Moreno. 2001. An Early Cretaceous pellet. Nature 409:998-999 [M. Carrano/M. Carrano]
70301 J. L. Sanz, C. Dieguéz, and F. J. Poyato-Ariza. 2000. Die Unter-Kreide von Las Hoyas, Cuenca, Spanien [The Lower Cretaceous of Las Hoyas, Cuenca, Spain]. In D. Meischner (ed.), Europäische Fossillagerstätten [European Fossil Lagerstätten] 155-160, 254-5 [M. Carrano/M. Carrano]
55576 J. L. Sanz, F. Ortega, and M. Shibata. 2014. Dinosaurios Maravillosos de España [Marvelous Dinosaurs from Spain] 1-155 [M. Carrano/M. Carrano]
47397 P. A. Selden and D. Penney. 2003. Lower Cretaceous spiders (Arthropoda: Arachnida: Araneae) from Spain. Neues Jahrbuch für Geologie und Paläontologie, Monatshefte 2003:175-192 [M. Clapham/M. Clapham]
35032 C. Soriano and X. Delclos. 2006. New cupedid beetles from the Lower Cretaceous of Spain and the palaeogeography of the family. Acta Palaeontologica Polonica 51:185-200 [M. Clapham/J. Karr]
40087 C. Soriano, A. G. Kirejtshuk, and X. Delclòs. 2006. The Mesozoic Laurasian family Parandrexidae (Insecta: Coleoptera:, new species from the Lower Cretaceous of Spain. Comptes Rendus Palevol 5:779-784 [M. Clapham/M. Clapham/M. Clapham]
5066ETE 778C. Soriano, A. G. Ponomarenko, and X. Delclòs. 2007. Coptoclavid beetles (Coleoptera: Adephaga) from the Lower Cretaceous of Spain: A new feeding strategy in beetles. Palaeontology 50(2):525-536 [C. Labandeira/C. Labandeira/J. Karr]
31124 R. Vullo, Á D. Buscalioni, J. Marugán-Lobón and J. J. Moratalla. 2009. First pterosaur remains from the Early Cretaceous Lagerstätte of Las Hoyas, Spain: palaeoecological significance. Geological Magazine 146:931-936 [R. Butler/R. Butler]
42229 R. Vullo, J. Marugán-Lobón, A. W. A. Kellner, A. D. Buscaloni, B. Gomez, M. Fuente, and J. J. Moratalla. 2012. A new crested pterosaur from the Early Cetaceous of Spain: the first European tapejarid (Pterodactyloidea: Azhdarchoidea). PLoS ONE 7(7):e38900 [R. Benson/R. Benson]
45786 S. Wenz. 2003. Les Lepidotes (Actinopterygii, Semionotiformes) de Cretace inferieur (Barremien) de Las Hoyas (Province de Cuenca, Espagne). Geodiversitas 25(3):481-499 [M. Bell/M. Bell/M. Bell]
47044 S. Wenz and F. J. Poyato-Ariza. 1994. Les Actiopterygiens juveniles du Cretace Inferieur du Montsec et de Las Hoyas (Espagne). Geobios 16:203-212 [M. Bell/M. Bell]
39570 V. V. Zherikhin and V. G. Gratshev. 2003. A new weevil-beetle (Insecta, Coleoptera, Nemonychidae) from the Lower Cretaceous of Spain. Paleontological Journal 37:407-408 [M. Clapham/M. Clapham]