Las Hoyas (MCCM collection): Late/Upper Barremian, Spain
List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
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Weitchselia reticulata
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Pseudoastacus llopisi
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Clavatoraxis robustus
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Sanz et al. 2014 | |||||||||
Turbomesodon praeclarus
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Sanz et al. 2014 | |||||||||
Iterophyllum lobatum
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Sanz et al. 2014 | |||||||||
Equisetopsida
- Araucariaceae
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Pagiophyllum sp.
Heer 1881
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Sanz et al. 2014 | |||||||||
Brachyphyllum speciosa
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spelled Brachyphilum | ||||||||||
Equisetopsida
- Cheirolepidiaceae
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Cupressinocladus micromerum
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Polypodiopsida
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Cladophlebis browniana
(Dunker 1846)
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Polypodiopsida
- Gleicheniales
- Matoniaceae
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Weichselia reticulata
(Stokes and Webb 1824)
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Sanz et al. 2014 | |||||||||
Polypodiopsida
- Schizaeales
- Schizaeaceae
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Ruffordia goepperti
(Dunker 1844)
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Sanz et al. 2014 | |||||||||
original and current combination Cheilanthites goepperti | ||||||||||
Angiospermae
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Montsechia vidali
(Zeiller 1902)
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Bennettitales
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Zamites sp.
Brongniart 1828
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Williamsonia sp.
Carruthers 1870
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spelled Williansonia | ||||||||||
Pinopsida
- Pinales
- Taxodiaceae
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Sphenolepis kurriana
(Dunker 1846)
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Bivalvia
- Unionida
- Unionidae
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Unio sp.
Philipsson 1788
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Sanz et al. 2014 | |||||||||
Insecta
- Blattodea
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Blattoidea indet.
Handlirsch 1906
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synonym of Blattaria | ||||||||||
Insecta
- Chresmodidae
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Chresmoda aquatica
Martínez-Delclòs 1989
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Delclòs et al. 2008 | 6 specimens | ||||||||
LH-26.546, LH-13826, LH-13574, LH-92AM102, MCCM-LH 18023, MCCM-LH 18024, | ||||||||||
Insecta
- Hemiptera
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Heteroptera indet.
Latreille 1810
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Insecta
- Hemiptera
- Naucoridae
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Naucoridae indet.
Fallen 1814
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Insecta
- Hemiptera
- Corixidae
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Velocorixinae indet.
Popov 1986
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Insecta
- Hemiptera
- Belostomatidae
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Iberonepa roemerali
Martínez-Delclòs et al. 1995
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Sanz et al. 2014 | |||||||||
Insecta
- Hemiptera
- Notonectidae
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Notonectidae indet.
(Latreille 1802)
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original and current combination Notonectariae | ||||||||||
Insecta
- Hymenoptera
- Scoliidae
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Cretoscolia conquensis n. sp.
Rasnitsyn and Martínez-Delclòs 2000
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Rasnitsyn and Martínez-Delclòs 2000 | 1 specimen | ||||||||
MCCM LH 17331 (1 measurement) | ||||||||||
Archaeoscolia hispanica n. sp.
Rasnitsyn and Martínez-Delclòs 1999
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Rasnitsyn and Martínez-Delclòs 1999 | 1 specimen | ||||||||
MCCM LH 13827 (2 measurements) | ||||||||||
Insecta
- Mecoptera
- Orthophlebiidae
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Orthophlebiinae indet.
(Handlirsch 1906)
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corrected as Orthophlebiidae | ||||||||||
Insecta
- Raphidioptera
- Mesoraphidiidae
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Mesoraphidiidae indet.
Martynov 1925
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Insecta
- Neuroptera
- Mesochrysopidae
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Triangulochrysopa sanzi n. gen., n. sp.
Nel et al. 2005
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Nel et al. 2005 | 5 specimens | ||||||||
MCCM LH-8110, 18572, 18573, 18574, MCCM 92/2/3 (9 measurements) | ||||||||||
Chimerochrysopa incerta n. gen., n. sp.
Nel et al. 2005
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Nel et al. 2005 | 1 specimen | ||||||||
MCCM LH 18588 (1 measurement) | ||||||||||
Allopterus mayorgai n. sp.
Nel et al. 2005
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Nel et al. 2005 | 2 specimens | ||||||||
MCCM LH 18570 (5 measurements) | ||||||||||
Armandochrysopa inexpecta n. sp.
Nel et al. 2005
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Nel et al. 2005 | 1 specimen | ||||||||
recombined as Kareninoides inexpectus | ||||||||||
MCCM LH 18575 (3 measurements) | ||||||||||
Tachinymphes paicheleri n. sp.
Nel et al. 2005
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Nel et al. 2005 | 10 specimens | ||||||||
MCCM LH 18576-8, LH 8040, LH 13175, MCCM LH 18579-18584 (7 measurements) | ||||||||||
Tachinymphes penalveri n. sp.
Nel et al. 2005
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Nel et al. 2005 | 5 specimens | ||||||||
MCCM LH 18585, LH 18586-7, U.A. Madrid LH 8033, 8094 (8 measurements) | ||||||||||
Nanochrysopa pumilio n. gen., n. sp.
Nel et al. 2005
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Nel et al. 2005 | 1 specimen | ||||||||
MCCM LH 13217 (2 measurements) | ||||||||||
Insecta
- Coleoptera
- Coptoclavidae
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Ovonectes pilosum n. gen., n. sp.
Soriano et al. 2007
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Soriano et al. 2007 | 2 specimens | ||||||||
MCCM-LH23853, 24514 (2 measurements) | ||||||||||
Hispanoclavina diazromerali n. gen., n. sp.
Soriano et al. 2007
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Soriano et al. 2007 | 5 specimens | ||||||||
MCCM-LH16264, 15070, 16315, 22299, 23859 (2 measurements) | ||||||||||
Hispanoclavina gratshevi n. sp.
Soriano et al. 2007
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Soriano et al. 2007 | 3 specimens | ||||||||
MCCM-LH15961, 15747, 23000 (2 measurements) | ||||||||||
Megacoptoclava longiurogomphia n. gen., n. sp.
Ponomarenko and Martínez-Delclòs 2000
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Ponomarenko and Martínez-Delclòs 2000 | 1 specimen | ||||||||
MCCM LH-1263 | ||||||||||
Hoyaclava buscalionae n. gen., n. sp.
Soriano et al. 2007
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Soriano et al. 2007 | 50 specimens | ||||||||
MCCM-LH24508, 7269, 23060, 24517 (2 measurements) | ||||||||||
Insecta
- Coleoptera
- Ademosynidae
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Ademosynidae indet.
Ponomarenko 1968
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Insecta
- Coleoptera
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Archostemata indet.
Kolbe 1908
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Zygadenia diazromerali n. sp.
Soriano and Delclos 2006
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Soriano and Delclos 2006 | 1 specimen | ||||||||
recombined as Notocupes diazromerali | ||||||||||
MCCM LH 12001 (4 measurements) | ||||||||||
Zygadenia siniestri n. sp.
Soriano and Delclos 2006
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Soriano and Delclos 2006 | 3 specimens | ||||||||
recombined as Conexicoxa siniestri | ||||||||||
MCCM LH 23515, MCCM LH 21291, MCCM LH 15819 (4 measurements) | ||||||||||
Zygadenia martinclosas
Soriano and Delclos 2006
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Soriano and Delclos 2006 | 1 individual | ||||||||
recombined as Conexicoxa martinclosas | ||||||||||
MCCM LH 20151 | ||||||||||
Zygadenia longicoxa n. sp.
Soriano and Delclos 2006
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Soriano and Delclos 2006 | 2 specimens | ||||||||
recombined as Brachilatus longicoxa | ||||||||||
MCCM LH 928030, MCCM LH 23432 | ||||||||||
Zygadenia viridis n. sp.
Soriano and Delclos 2006
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Soriano and Delclos 2006 | 1 specimen | ||||||||
recombined as Brachilatus viridis | ||||||||||
MCCMLH 1805 (2 measurements) | ||||||||||
Insecta
- Coleoptera
- Cupedidae
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Anaglyphites zherikhini n. sp.
Soriano and Delclos 2006
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Soriano and Delclos 2006 | 1 specimen | ||||||||
recombined as Mesocupes zherikhini | ||||||||||
MCCM LH 15122 (2 measurements) | ||||||||||
Priacma sanzii n. sp.
Soriano and Delclos 2006
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Soriano and Delclos 2006 | 1 specimen | ||||||||
nomen dubium belonging to Cupedinae | ||||||||||
MCCM LH 24524 (2 measurements) | ||||||||||
Insecta
- Coleoptera
- Ommatidae
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Tetraphalerus ponomarenkoi n. sp.
Soriano and Delclos 2006
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Soriano and Delclos 2006 | 1 specimen | ||||||||
MCCM LH 78 (2 measurements) | ||||||||||
Insecta
- Coleoptera
- Parandrexidae
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Martynopsis laticollis n. gen., n. sp.
Soriano et al. 2006
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Soriano et al. 2006 | 2 specimens | ||||||||
MCCM LH-21091, 24513 (2 measurements) | ||||||||||
Insecta
- Coleoptera
- Nemonychidae
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Distenorrhinus (Parabrenthorrhinus) xavieri n. sp.
Zherikhin and Gratshev 2003
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Zherikhin and Gratshev 2003 | 1 specimen | ||||||||
MCCM LH 17330 (1 measurement) | ||||||||||
Insecta
- Odonata
- Aeschnidiidae
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Gigantoaeschnidium ibericus n. gen., n. sp.
Nel and Martínez-Delclòs 1993
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Fleck and Nel 2003 | 6 specimens | ||||||||
(6 measurements) | ||||||||||
Sinaeschnidia martinezdelclosi n. sp.
Fleck and Nel 2003
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Fleck and Nel 2003 | 2 specimens | ||||||||
LH 17332, ADR-317-I (2 measurements) | ||||||||||
Iberoaeschnidium conquensis n. gen., n. sp.
Nel and Martínez-Delclòs 1993
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Fleck and Nel 2003 | 1 specimen | ||||||||
LH-083-R (1 measurement) | ||||||||||
Insecta
- Ephemeroptera
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Ephemeroptera indet.
Hyatt and Arms 1890
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Insecta
- Diptera
- Ptychopteridae
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Eoptychoptera sp.
Handlirsch 1906
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Lukashevich and Arillo 2016 | 1 specimen | ||||||||
MCCM-LH 35084 | ||||||||||
Insecta
- Diptera
- Tipulidae
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Leptotarsus ibericus n. sp.
Ribeiro and Lukashevich 2014
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Ribeiro and Lukashevich 2014 | 3 specimens | ||||||||
MCCM LH 28206, 18122, 29995 (2 measurements) | ||||||||||
Leptotarsus buscalioniae n. sp.
Ribeiro and Lukashevich 2014
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Ribeiro and Lukashevich 2014 | 1 specimen | ||||||||
MCCM LH 35088 (2 measurements) | ||||||||||
Leptotarsus contractus n. sp.
Ribeiro and Lukashevich 2014
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Ribeiro and Lukashevich 2014 | 1 specimen | ||||||||
MCCM LH 15419 (2 measurements) | ||||||||||
Arachnida
- Opiliones
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Opiliones indet.
Sundevall 1833
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Palencia et al. 2019 | 1 specimen | ||||||||
MUPA LH-29969 | ||||||||||
Malacostraca
- Decapoda
- Atyidae
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Oplophorus roselli
Via 1971
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recombined as Delclosia roselli | ||||||||||
Delclosia martinelli
Rabada 1993
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Sanz et al. 2014 | |||||||||
Malacostraca
- Decapoda
- Astacidae
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Austropotamobius llopisi
(Via 1971)
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Sanz et al. 2014 | |||||||||
unclassified
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Undichna unisulca
de Gibert et al. 1999
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Moratalla et al. 2017 | |||||||||
Coelacanthimorpha
- Coelacanthiformes
- Latimeriidae
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? Holophagus leridae
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? "Holophagus" | ||||||||||
Amphibia
- Temnospondyli
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Anura indet.
(Fischer von Waldheim 1813)
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Sanz et al. 1988 | 1 individual | ||||||||
Iberobatrachus angelae n. gen., n. sp.
Baez 2013
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Baez 2013 | |||||||||
MCCM-LH 11393a-b - holotype (articulated specimen lacking the hindlimbs and left forelimb, two-dimensionally preserved as part and counterpart on two slabs); referred specimen - MCCM-LH 15999a-b (incomplete, articulated specimen preserved as part and counterpart on two slabs) | ||||||||||
Gracilibatrachus avallei n. gen., n. sp.
Baez 2013
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Baez 2013 | |||||||||
MCCM-LH 21171a, b - holotype (articulated skeleton of a postmetamorphic individual); referred specimens: MCCMLH-17422a, b, 11386, 11387, 26251, 26803a, b (in part), and, tentatively, 11394a, b | ||||||||||
Hylaeobatrachus sp.
Dollo 1884
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Sanz et al. 2014 | |||||||||
Amphibia
- Temnospondyli
- Urodela
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Caudata indet.
Scopoli 1777
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= Valdotriton gracilis n. gen., n. sp.
Evans and Milner 1996
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Evans and Milner 1996 | |||||||||
MCCM-LH 2850a-b, 6021, 928001a-b, ADR unnumbered | ||||||||||
Amphibia
- Temnospondyli
- Albanerpetontidae
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Celtedens ibericus n. sp.
McGowan and Evans 1995
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McGowan and Evans 1995 | |||||||||
LH 6020 - type (part and counterpart of a nearly complete skeleton, missing only part of the palate and tail); referred material (McGowan 2002): two complete specimens in part and counterpart (LH 030 R A & B; LH 15710 A & B) | ||||||||||
Mammalia
- Groeberida
- Nototheriidae
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Spinolestes xenarthrosus n. gen., n. sp.
Martin et al. 2015
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Martin et al. 2015 | |||||||||
MCCM-LH 30000 - holotype (complete skeleton); MCCMLH30000B (counter slab) | ||||||||||
Reptilia
- Testudines
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Toxochelyidae indet.
Baur 1895
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2 specimens | |||||||||
MCCM-LH 84, LH 254 (carapace fragment) | ||||||||||
= Hoyasemys jimenezi n. gen., n. sp.
Pérez-García et al. 2012
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Pérez-García et al. 2012 | |||||||||
Holotype: MCCM-LH 84, an almost complete skeleton, missing the dorsal portion of the skull. The specimen is preserved on two slabs. One shows the ventral view of the skull, the five proximal cervical vertebrae, some dorsal vertebrae, some fragments of the peripherals and of the lat− eral part of the costal plates, the plastron, sacral and caudal vertebrae and the appendicular skeleton. The other slab preserves the partial carapace and some bones of the hindlimb. | ||||||||||
Reptilia
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Jucaraseps grandipes n. gen., n. sp.
Bolet and Evans 2012
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Bolet and Evans 2012 | 1 individual | ||||||||
LH 18505 (holotype), almost complete skeleton in dorsal view | ||||||||||
Reptilia
- Teiidae
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Meyasaurus diazromerali n. sp.
Evans and Barbadillo 1997
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Evans and Barbadillo 1997 | 6 individuals | ||||||||
LH 370, LH 372, LH 33, LH 13510, LH 6026, LH 14317 | ||||||||||
Reptilia
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Lacertilia indet.
Owen 1842
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2 individuals | |||||||||
= Scandensia ciervensis n. gen., n. sp.
Evans and Barbadillo 1998
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Evans and Barbadillo 1998 | |||||||||
Hoyalacerta sanzi n. gen., n. sp.
Evans and Barbadillo 1999
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Evans and Barbadillo 1999 | 1 individual | ||||||||
LH 11000 (holotype skeleton) | ||||||||||
Crocodylomorpha indet.
(Hay 1930)
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Sanz et al. 2014 | |||||||||
Reptilia
- Gobiosuchidae
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Gobiosuchidae indet.
Osmolska 1972
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Sanz et al. 2014 | |||||||||
= Cassissuchus sanziuami n. gen., n. sp.
Buscalioni 2017
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Buscalioni 2017 | |||||||||
MCCM-LH 7991 - holotype; referred specimen: MCCMLH 6508 | ||||||||||
Reptilia
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Montsecosuchus sp.
Buscalioni and Sanz 1988
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Sanz et al. 2014 | |||||||||
Reptilia
- Atoposauridae
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Atoposauridae indet.
(Gervais 1871)
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Reptilia
- Bernissartiidae
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Bernissartidae indet.
Dollo 1883
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Sanz et al. 2014 | |||||||||
corrected as Bernissartiidae | ||||||||||
Reptilia
|
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Mantellisaurus atherfieldensis
(Hooley 1925)
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Llandres Serrano et al. 2013 | |||||||||
LH-30677, incomplete but fully-articulated right hindlimb | ||||||||||
= cf. Mantellisaurus sp.
Paul 2007
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Moratalla et al. 2017 | |||||||||
Reptilia
- Theropoda
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Megalosauridae indet.
(Fitzinger 1843)
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Canudo and Ruiz-Omeñaca 2003 | |||||||||
tooth | ||||||||||
= Theropoda indet.
Marsh 1881
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Canudo and Ruiz-Omeñaca 2003 | |||||||||
Reptilia
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? Pterosauria indet.
(Kaup 1834)
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Lockley et al. 1995 | 1 specimen | ||||||||
MCCM-LH-6500, isolated track | ||||||||||
= ? Theropoda indet.
Marsh 1881
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Vullo et al. 2009 | |||||||||
Theropoda indet.
(Marsh 1881)
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Moratalla et al. 2017 | 1 individual | ||||||||
MCCM-LH-Y-1-001, trackway | ||||||||||
Aves indet.
Linnaeus 1758
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Sanz et al. 2001 | 4 individuals | ||||||||
MCCM-LH 11386; 3 taxa in a pellet | ||||||||||
Eoalulavis hoyasi n. gen., n. sp.
Sanz et al. 1996
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Sanz et al. 1996 | 1 specimen | ||||||||
Concornis lacustris n. gen., n. sp.
Sanz and Buscalioni 1992
|
Sanz et al. 1995 | |||||||||
MCCM-LH-2814 | ||||||||||
cf. Concornis lacustris
Sanz and Buscalioni 1992
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Cambra-Moo et al. 2006 | 2 specimens | ||||||||
MCCM-LH 21006 a-b, tibiotarsus and tarsometatarsus | ||||||||||
cf. Iberomesornis sp.
Sanz et al. 1990
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Sanz and Buscalioni 1994 | |||||||||
Aves indet.
Linnaeus 1758
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new genus and species | ||||||||||
= Iberomesornis romerali n. gen., n. sp.
Sanz et al. 1990
|
Sanz et al. 1990 | |||||||||
MCCM-LH 22R - holotype (partial skeleton) | ||||||||||
Pelecanimimus polyodon n. gen., n. sp.
Pérez-Moreno et al. 1994
|
Pérez-Moreno et al. 1994 | 1 individual | ||||||||
MCCM-LH 7777 (1 measurement) | ||||||||||
Reptilia
- Carcharodontosauridae
|
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Concavenator corcovatus n. gen., n. sp.
Ortega et al. 2010
|
Ortega et al. 2010 | 1 individual | ||||||||
MCCM-LH 6666 (5 measurements) | ||||||||||
Reptilia
|
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Ornithocheiridae indet.
(Seeley 1870)
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Vullo et al. 2009 | 2 elements | ||||||||
MCCM-LH-17264, LH-21451, teeth | ||||||||||
Reptilia
- Istiodactylidae
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Istiodactylidae indet.
Howse et al. 2001
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Vullo et al. 2009 | 2 elements | ||||||||
MCCM-LH-28572, LH-15448, teeth | ||||||||||
Reptilia
|
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Europejara olcadesorum n. gen., n. sp.
Vullo et al. 2012
|
Vullo et al. 2012 | 1 specimen | ||||||||
MCCM-LH 9413, jaws and post-orbital elements of the skull preserved on part and counterpart slabs | ||||||||||
Actinopteri
- Pycnodontiformes
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Pycnodontiformes indet.
Berg 1937
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genus incertae sedis | ||||||||||
Actinopteri
- Pycnodontiformes
- Pycnodontidae
|
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Macromesodon sp.
Blake 1905
|
Wenz and Poyato-Ariza 1994 | |||||||||
Coelodus subdiscus
|
Wenz and Poyato-Ariza 1994 | |||||||||
Actinopteri
- Pycnodontiformes
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Eomesodon sp.
Woodward 1918
|
Wenz and Poyato-Ariza 1994 | |||||||||
Actinopteri
- Semionotiformes
- Macrosemiidae
|
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Notagogus sp.
Agassiz 1833
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Notagogus ferreri
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Actinopteri
- Lepisosteiformes
|
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Lepidotes sp.
Agassiz 1832
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Lepidotes microrbis
|
Sanz et al. 2014 | |||||||||
Lepidotes tanyrhis n. sp.
Wenz 2003
|
Wenz 2003 | |||||||||
MCCM-LH-7410 | ||||||||||
Lepidotes microrhis n. sp.
Wenz 2003
|
Wenz 2003 | |||||||||
MCCM-LH-8010a,b | ||||||||||
Actinopteri
- Amiiformes
- Caturidae
|
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Caturus sp.
Agassiz 1834
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Actinopteri
- Amiiformes
- Amiidae
|
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? Amiopsis sp.
Kner 1863
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Urocles woodwardi
|
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synonym of Amiopsis | ||||||||||
Vidalamia catalunica
(Sauvage 1903)
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Actinopteri
- Gonorhynchiformes
- Chanidae
|
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Rubiesichthys gregalis
Wenz 1984
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Actinopteri
|
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Gordichthys conquensis n. gen., n. sp.
Poyato-Ariza 1994
|
Poyato-Ariza 1994 | |||||||||
holotype: LH 1288 R | ||||||||||
Actinopteri
- Leptolepidiformes
- Leptolepididae
|
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? Ascalabos voithi
|
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synonym of Leptolepis | ||||||||||
Actinopteri
- Pholidophoriformes
- Pleuropholidae
|
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Pleuropholis sp.
Egerton 1858
|
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unclassified
|
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Planolites sp.
Nicholson 1873
|
Moratalla et al. 2017 | |||||||||
Treptichnus sp.
Miller 1889
|
Moratalla et al. 2017 | |||||||||
Lockeia sp.
James 1879
|
Moratalla et al. 2017 | |||||||||
Cruziana sp.
d'Orbigny 1842
|
Moratalla et al. 2017 | |||||||||
Palaeophycus sp.
Hall 1847
|
Moratalla et al. 2017 | |||||||||
Helminthoidichnites sp.
Fitch 1850
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Moratalla et al. 2017 | |||||||||
see common names |
Geography
Country: | Spain | State/province: | Castilla-La Mancha | County: | Cuenca |
Coordinates: | 40.1° North, 1.9° West (view map) | ||||
Paleocoordinates: | 30.5° North, 8.1° East | ||||
Basis of coordinate: | estimated from map | ||||
Altitude: | 428 meters | ||||
Geographic resolution: | outcrop |
Time
Period: | Cretaceous | Epoch: | Early/Lower Cretaceous |
Stage: | Barremian | 10 m.y. bin: | Cretaceous 2 |
*Period: | Early/Lower Cretaceous | *Epoch: | Neocomian - Gallic |
*International age/stage: | Late/Upper Hauterivian - Early/Lower Barremian | ||
Key time interval: | Late/Upper Barremian | ||
Age range of interval: | 130.00000 - 125.45000 m.y. ago | ||
* legacy (obsolete) database fields |
Stratigraphy
Formation: | La Huérguina | ||||
Local section: | Las Hoyas | Local bed: | 3 | ||
Local order: | bottom to top | ||||
Stratigraphic resolution: | group of beds | ||||
Stratigraphy comments: also called "Calizas de La Huérgina Formation" or "La Huérgina Limestones Formations" |
Lithology and environment
Primary lithology: | planar lamination "limestone" | ||
Secondary lithology: | packstone | ||
Includes fossils? | Y | ||
Lithology description: Limestone with laminates; three facies described: Massive limestone, packstones with peloids and sand-sized fragments of Charophyte stems. Occasionally, in the base of these beds, levels of gastropodal wackestones appear; Flaggy limestones, sometimes nodular. Ostracodal mudstones with occasional thick laminations, laterally grading to confolute laminations; Laminated limestones, mudstones with laminations containing abundant fossils. Laminae are very continuous laterally | |||
Environment: | lacustrine - large | Tectonic setting: | rift |
Geology comments: shallow to deep lacustrine facies; "produced in the context of a continental (freshwater) subtropical, seasonal summer wet, carbonate wetland that overlay a low-relief karstic terrain. The wetland was drained by carbonate-rich freshwater and comprised a typical environmental mosaic of swampy plains, ponds, lakes, ephemeral channels and sloughs." |
Taphonomy
Modes of preservation: | body,adpression,trace,soft parts |
Lagerst�tten type: | conservation |
Degree of concentration: | concentrated |
Size of fossils: | macrofossils,mesofossils |
Preservation of anatomical detail: | excellent |
Articulated whole bodies: | many |
Associated major elements: | some |
Disassociated major elements: | some |
Disassociated minor elements: | some |
Spatial resolution: | parautochthonous |
Collection methods and comments
Collection excludes: | some macrofossils,some microfossils |
Collection methods: | selective quarrying,surface (in situ),field collection |
Reason for describing collection: | general faunal/floral analysis |
Collection method comments: Repository: Museo de las Ciencias de Castilla-La Mancha, Cuenca (MCCM) |
Metadata
Database number: | 28006 | ||
Authorizer: | J. Alroy, M. Bell, M. Clapham, R. Benson, M. Carrano, P. Mannion, R. Butler | Enterer: | E. Leckey, M. Carrano, J. Alroy, J. Tennant, M. Oreska, M. Bell, R. Benson, M. Clapham, J. Karr, R. Butler, P. Mannion |
Modifier: | M. Carrano | Research group: | paleoentomology,vertebrate |
Created: | 2003-01-20 12:47:22 | Last modified: | 2021-01-22 10:43:48 |
Access level: | the public | Released: | 2003-01-20 12:47:22 |
Creative Commons license: | CC BY |
Reference information
Primary reference:
7671. | 5% 14640 | J. L. Sanz, S. Wenz, A. Yebenes, R. Estes, X. Martinez-Delclos, E. Jiminez-Fuentes, C. Dieguez, A. D. Buscalioni, L. J. Barbadillo and L. Via. 1988. An Early Cretaceous faunal and floral continental assemblage: Las Hoyas fossil site (Cuenca, Spain). Géobios 21(5):611-635 [J. Alroy/E. Leckey/J. Alroy] |
Secondary references:
44133 | A. M. Baez. 2013. Anurans from the Early Cretaceous Lagerstätte of Las Hoyas, Spain: New evidence on the Mesozoic diversification of crown-clade Anura. Cretaceous Research 41:90-106 [P. Mannion/P. Mannion/P. Mannion] | |
35206 | G. Bechly, A. Nel, X. Martínez-Declòs, E. A. Jarzembowski, R. Coram, D. Martill, G. Fleck, F. Escuillié, M. M. Wisshak and M. Maisch. 2001. A revision and phylogenetic study of Mesozoic Aeshnoptera, with description of numerous new taxa (Insecta: Odonata: Anisoptera). Neue Paläontologische Abhandlungen 4:1-219 [M. Clapham/M. Clapham/M. Clapham] | |
41482 | A. Bolet and S. E. Evans. 2012. A tiny lizard (Lepidosauria, Squamata) from the Lower Cretaceous of Spain. Palaeontology 55(3):491-500 [R. Benson/R. Benson] | |
62518 | A. D. Buscalioni. 2017. The Gobiosuchidae in the early evolution of Crocodyliformes. Journal of Vertebrate Paleontology 37(3):e1324459:1-21 [P. Mannion/P. Mannion/P. Mannion] | |
17880 | ETE | A. D. Buscalioni and M. Frenegal-Martinez. 2006. Archosaurian size bias in Jurassic and Cretaceous freshwater ecosystems. 9th International Symposium on Mesozoic Terrestrial Ecosystems and Biota, Abstracts and Proceedings Volume 9-12 [M. Carrano/M. Carrano/M. Carrano] |
34089 | ETE | O. Cambra-Moo, A. D. Buscalioni, J. Cubo, J. Castanet, M.-M. Loth, E. Margerie, and A. Ricqlès. 2006. Histological observations of enantiornithine bone (Saurischia, Aves) from the Lower Cretaceous of Las Hoyas (Spain). Comptes Rendus Palevol 5:685-691 [M. Carrano/M. Oreska/M. Carrano] |
17647 | ETE | J. I. Canudo and J. I. Ruiz-Omeñaca. 2003. Los restos directos de dinosaurios teropódos (excluyendo Aves) en España [Direct remains of theropod dinosaurs (excluding Aves) in Spain]. Ciencias de la Tierra 26:347-373 [M. Carrano/M. Carrano] |
55113 | E. Cuesta, I. Díaz-Martínez, F. Ortega and J. L. Sanz. 2015. Did all theropods have chicken-like feet? First evidence of a non-avian dinosaur podotheca. Cretaceous Research 56(1):53-59 [M. Carrano/M. Carrano] | |
41244 | X. Delclòs, A. Nel, D. Azar, G. Bechly, J. A. Dunlop, M. S. Engel, and S. W. Heads. 2008. The enigmatic Mesozoic insect taxon Chresmodidae (Polyneoptera): new palaeobiological and phylogenetic data, with the description of a new species from the Lower Cretaceous of Brazil. Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen 247:353-381 [M. Clapham/M. Clapham/M. Clapham] | |
37280 | S. E. Evans and L. J. Barbadillo. 1997. Early Cretaceous lizards from Las Hoyas, Spain. Zoological Journal of the Linnean Society 119:23-49 [R. Benson/R. Benson] | |
26529 | ETE | S. E. Evans and L. J. Barbadillo. 1998. An unusual lizard (Reptilia: Squamata) from the Early Cretaceous of Las Hoyas, Spain. Zoological Journal of the Linnean Society 124:235-265 [M. Carrano/M. Carrano] |
37784 | S. E. Evans and L. J. Barbadillo. 1999. A short-limbed lizard from the Lower Cretaceous of Spain. In D. M. Unwin (ed.), Cretaceous Fossil Vertebrates. Special Papers in Palaeontology 60:73-85 [R. Benson/R. Benson/M. Carrano] | |
25847 | ETE | S. E. Evans and A. R. Milner. 1996. A metamorphosed salamander from the Early Cretaceous of Las Hoyas, Spain. Philosophical Transactions of the Royal Society of London B 351:627-646 [M. Carrano/M. Carrano] |
35239 | G. Fleck, G. Bechly, X. Martinez-Delclos, E. Jarzembowski, R. Coram and A. Nel. 2003. Phylogeny and classification of the Stenophlebioptera (Odonata: Epiproctophora). Annales de la Société Entomologique de France 39(1):55-93 [M. Clapham/J. Karr] | |
35493 | G. Fleck and A. Nel. 2003. Revision of the Mesozoic family Aeschnidiidae (Odonata: Anisoptera). Zoologica 153:1-170 [M. Clapham/J. Karr] | |
28227 | ETE | N. S. Gupta, O. Cambra-Moo, D. E. G. Briggs, G. D. Love, M. A. Fregenal-Martinez and R. E. Summons. 2008. Molecular taphonomy of macrofossils from the Cretaceous Las Hoyas Formation, Spain. Cretaceous Research 29:1-8 [M. Carrano/M. Carrano] |
42228 | D. W. E. Hone, H. Tischlinger, E. Frey and M. Röper. 2012. A new non-pterodactyloid pterosaur from the Late Jurassic of southern Germany. PLoS ONE 7(7):e38900 [R. Benson/R. Benson] | |
45545 | M. Llandres Serrano, R. Vullo, J. Marugan-Lobon, F. Ortega, and A. D. Buscalioni. 2013. An articulated hindlimb of a basal iguanodont (Dinosauria, Ornithopoda) from the Early Cretaceous Las Hoyas Lagerstätte (Spain). Geological Magazine 150(3):527-576 [P. Mannion/J. Tennant/M. Carrano] | |
31125 | M. Lockley, T. J. Logue, J. J. Moratalla, A. P. Hunt, R. J. Schultz and J. W. Robinson. 1995. The fossil trackway Pteraichnus is pterosaurian, not crocodilian: implications for the global distribution of pterosaur tracks. Ichnos 4:7-20 [R. Butler/R. Butler] | |
56924 | E. D. Lukashevich and A. Arillo. 2016. New Eoptychoptera (Insecta: Diptera, Ptychopteridae) from the Lower Cretaceous of Spain. Cretaceous Research 58:254-264 [M. Clapham/M. Clapham] | |
56864 | T. Martin, J. Marugán-Lobón, R. Vullo, H. Martín-Abad, Z.-X. Luo and A. D. Buscalioni. 2015. A Cretaceous eutriconodont and integument evolution in early mammals. Nature 526:380-384 [P. Mannion/P. Mannion] | |
49227 | X. Martínez-Delclòs. 1993. Blátidos (Insecta, Blattodea) del Cretácico Inferior de España. Familias Mesoblattinidae, Blattulidae y Poliphagidae. Boletín Geológico y Minero 104:516-538 [M. Clapham/M. Clapham] | |
45506 | X. Martínez-Delclòs, A. Nel, and Y. A. Popov. 1995. Systematics and functional morphology of Iberonepa romerali n. gen. and sp., Belostomatidae from the Spanish Lower Cretaceous (Insecta, Heteroptera). Journal of Paleontology 69:496-508 [M. Clapham/M. Clapham] | |
44194 | G. McGowan and S. E. Evans. 1995. Albanerpetontid amphibians from the Cretaceous of Spain. Nature 373:143-145 [P. Mannion/P. Mannion] | |
42672 | G. J. McGowan. 2002. Albanerpetontid amphibians from the Lower Cretaceous of Spain and Italy: a description and reconsideration of their systematics. Zoological Journal of the Linnean Society 135(1):1-32 [M. Carrano/M. Carrano] | |
78477 | J. J. Moratalla, J. Marugán-Lobón, H. Martín-Abad, E. Cuesta, and A. D. Buscalioni. 2017. A new trackway possibly made by a trotting theropod at the Las Hoyas fossil site (Early Cretaceous, Cuenca Province, Spain): Identification, bio-dynamics and palaeoenvironmental implications. Palaeontologia electronica 20(3):59A:1-14 [M. Carrano/M. Carrano] | |
8943 | ETE | M. B. Mostovski and X. Martínez-Delclòs. 2000. New Nemestrinoidea (Diptera: Brachycera) from the Upper Jurassic - Lower Cretaceous of Eurasia, taxonomy, and palaeobiology. Entomological Problems 31(2):137-148 [C. Labandeira/A. Spencer-Lee/J. Karr] |
39071 | A. Nel, X. Delclos, and A. Hutin. 2005. Mesozoic chrysopid-like Planipennia: a phylogenetic approach (Insecta: Neuroptera). Annales de la Société Entomologique de France 41:29-69 [M. Clapham/M. Clapham/M. Clapham] | |
35441 | A. Nel and X. Martínez-Delclòs. 1993. Nuevos Zygoptera y Anisoptera (Insecta: Odonata) en el Cretacico inferior de Espana. Estudios Geológicos 49:351-359 [M. Clapham/J. Karr/M. Clapham] | |
33846 | ETE | F. Ortega, F. Escaco, and J. L. Sanz. 2010. A bizarre, humped Carcharodontosauria (Theropoda) from the Lower Cretaceous of Spain. Nature 467:203-206 [M. Carrano/M. Carrano/M. Carrano] |
67846 | L. Palencia, E. Peñalver, C. E. Prieto and F. J. Poyato-Ariza. 2019. First fossil harvestmen (Arachnida: Opiliones) from Spain and notes on the fossil record of Opiliones. Palaeontologia Electronica 22(1.5A):1-18 [M. Clapham/M. Clapham/M. Clapham] | |
42000 | A. Pérez-García, M. S. de la Fuente, and F. Ortega. 2012. A new freshwater basal eucryptodiran turtle from the Early Cretaceous of Spain. Acta Palaeontologica Polonica 57(2):285-298 [R. Benson/R. Benson/P. Mannion] | |
12449 | ETE | B. P. Pérez-Moreno, J. L. Sanz, A. D. Buscalioni, J. J. Moratalla, F. Ortega and D. Rasskin-Gutman. 1994. A unique multitoothed ornithomimosaur from the Lower Cretaceous of Spain. Nature 370:363-367 [M. Carrano/M. Carrano/M. Carrano] |
61983 | B. P. Pérez-Moreno and J. L. Sanz. 1997. Las Hoyas. In P. J. Currie & K. Padian (ed.), Encyclopedia of Dinosaurs 398-402 [M. Carrano/M. Carrano] | |
48667 | A. Ponomarenko and X. Martínez-Delclòs. 2000. New beetles (Insecta: Coleoptera) from the Lower Cretaceous of Spain. Acta Geologica Hispanica 35:47-52 [M. Clapham/M. Clapham] | |
47043 | F. J. Poyato-Ariza. 1994. A new Early Cretaceous Gonorynchiform fish (Teleostei: Ostariophysi) from Las Hoyas (Cuenca, Spain). Occassional papers of the Museum of Natural History, The University of Kansas, Lawrence, Kansas 164:1-37 [M. Bell/M. Bell] | |
39638 | A. P. Rasnitsyn and X. Martínez-Delclòs. 1999. New Cretaceous Scoliidae (Vespida=Hymenoptera) from the Lower Cretaceous of Spain and Brazil. Cretaceous Research 20:767-772 [M. Clapham/M. Clapham] | |
39604 | A. P. Rasnitsyn and X. Martínez-Delclòs. 2000. Wasps (Insecta: Vespida = Hymenoptera) from the Early Cretaceous of Spain. Acta Geologica Hispanica 35:65-95 [M. Clapham/M. Clapham/M. Clapham] | |
50631 | G. C. Ribeiro and E. D. Lukashevich. 2014. New Leptotarsus from the Early Cretaceous of Brazil and Spain: the oldest members of the family Tipulidae (Diptera). Zootaxa 3753:347-363 [M. Clapham/M. Clapham] | |
55500 | J. I. Ruiz-Omeñaca and J. I. Canudo. 2003. Dinosaurios (Saurischia, Ornithischia) en el Barremiense (Cretácico Inferior) de la península Ibérica [Dinosaurs (Saurischia, Ornithischia) in the Barremian (Lower Cretaceous) of the Iberian peninsula]. In F. Pérez Lorente (ed.), Dinosaurios y Otros Reptiles Mesozóicos de España 269-312 [M. Carrano/M. Carrano] | |
70303 | J. L. Sanz. 2000. Yacimientos ibéricos de dinosaurios [Iberian dinosaur localities]. Dinosaurios: Los Señores del Pasado [Dinosaurs: The Lords of the Land] 267-299 [M. Carrano/M. Carrano] | |
13500 | ETE | J. L. Sanz, J. F. Bonaparte, and A. Lacasa. 1988. Unusual Early Cretaceous birds from Spain. Nature 331:433-435 [M. Carrano/M. Carrano/M. Carrano] |
19339 | ETE | J. L. Sanz, A. D. Buscalioni, J. J. Moratalla, V. Francés, and M. Antón. 1990. Los reptiles Mesozoicos del registro español [The Mesozoic reptiles from the Spanish record]. Monografias del Museo Nacional de Ciencias Naturales 2:1-79 [M. Carrano/M. Carrano] |
13505 | ETE | J. L. Sanz and A. D. Buscalioni. 1994. Géobios, Mémoire Spéciale 16:213-217 [M. Carrano/M. Carrano/M. Carrano] |
24393 | ETE | J. L. Sanz, L. M. Chiappe, and A. D. Buscalioni. 1995. The osteology of Concornis lacustris (Aves: Enantiornithes) from the Lower Cretaceous of Spain and a reexamination of its phylogenetic relationships. American Museum Novitates 3133:1-23 [M. Carrano/M. Carrano] |
13506 | ETE | J. L. Sanz, L. M. Chiappe, B. P. Pérez-Moreno, A. D. Buscalioni, J. J. Moratalla, F. Ortega, and F. J. Poyato-Ariza. 1996. An Early Cretaceous bird from Spain and its implications for the evolution of avian flight. Nature 382:442-445 [M. Carrano/M. Carrano/M. Carrano] |
25164 | ETE | J. L. Sanz, L. M. Chiappe, Y. Fernández-Jalvo, F. Ortega, B. Sánchez-Chillón, F. J. Poyato-Ariza, and B. P. Pérez-Moreno. 2001. An Early Cretaceous pellet. Nature 409:998-999 [M. Carrano/M. Carrano] |
70301 | J. L. Sanz, C. Dieguéz, and F. J. Poyato-Ariza. 2000. Die Unter-Kreide von Las Hoyas, Cuenca, Spanien [The Lower Cretaceous of Las Hoyas, Cuenca, Spain]. In D. Meischner (ed.), Europäische Fossillagerstätten [European Fossil Lagerstätten] 155-160, 254-5 [M. Carrano/M. Carrano] | |
55576 | J. L. Sanz, F. Ortega, and M. Shibata. 2014. Dinosaurios Maravillosos de España [Marvelous Dinosaurs from Spain] 1-155 [M. Carrano/M. Carrano] | |
47397 | P. A. Selden and D. Penney. 2003. Lower Cretaceous spiders (Arthropoda: Arachnida: Araneae) from Spain. Neues Jahrbuch für Geologie und Paläontologie, Monatshefte 2003:175-192 [M. Clapham/M. Clapham] | |
35032 | C. Soriano and X. Delclos. 2006. New cupedid beetles from the Lower Cretaceous of Spain and the palaeogeography of the family. Acta Palaeontologica Polonica 51:185-200 [M. Clapham/J. Karr] | |
40087 | C. Soriano, A. G. Kirejtshuk, and X. Delclòs. 2006. The Mesozoic Laurasian family Parandrexidae (Insecta: Coleoptera:, new species from the Lower Cretaceous of Spain. Comptes Rendus Palevol 5:779-784 [M. Clapham/M. Clapham/M. Clapham] | |
5066 | ETE 778 | C. Soriano, A. G. Ponomarenko, and X. Delclòs. 2007. Coptoclavid beetles (Coleoptera: Adephaga) from the Lower Cretaceous of Spain: A new feeding strategy in beetles. Palaeontology 50(2):525-536 [C. Labandeira/C. Labandeira/J. Karr] |
31124 | R. Vullo, Á D. Buscalioni, J. Marugán-Lobón and J. J. Moratalla. 2009. First pterosaur remains from the Early Cretaceous Lagerstätte of Las Hoyas, Spain: palaeoecological significance. Geological Magazine 146:931-936 [R. Butler/R. Butler] | |
42229 | R. Vullo, J. Marugán-Lobón, A. W. A. Kellner, A. D. Buscaloni, B. Gomez, M. Fuente, and J. J. Moratalla. 2012. A new crested pterosaur from the Early Cetaceous of Spain: the first European tapejarid (Pterodactyloidea: Azhdarchoidea). PLoS ONE 7(7):e38900 [R. Benson/R. Benson] | |
45786 | S. Wenz. 2003. Les Lepidotes (Actinopterygii, Semionotiformes) de Cretace inferieur (Barremien) de Las Hoyas (Province de Cuenca, Espagne). Geodiversitas 25(3):481-499 [M. Bell/M. Bell/M. Bell] | |
47044 | S. Wenz and F. J. Poyato-Ariza. 1994. Les Actiopterygiens juveniles du Cretace Inferieur du Montsec et de Las Hoyas (Espagne). Geobios 16:203-212 [M. Bell/M. Bell] | |
39570 | V. V. Zherikhin and V. G. Gratshev. 2003. A new weevil-beetle (Insecta, Coleoptera, Nemonychidae) from the Lower Cretaceous of Spain. Paleontological Journal 37:407-408 [M. Clapham/M. Clapham] |